Table Of ContentTRITICALE: TODAY AND TOMORROW
Developments in Plant Breeding
VOLUME 5
Triticale:
Today and Tomorrow
Edited by
HENRIQUE GUEDES-PINTO
Departamento de Genetica e Biotecn%gia,
Universidade de Tras-os-Montes e Alto Douro,
Vila Real, Portugal
NORMAN DARVEY
Plant Breeding Institute,
University of Sydney,
Cobbity, Australia
and
VALDEMAR P. CARNIDE
Departamento de Genetica e Biotecnologia,
Universidade de Tras-os-Montes e Alto Douro,
Vila Real, Portugal
KLUWER ACADEMIC PUBLISHERS
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ISBN-13:978-94-010-6634-1' e-ISBN-13:978-94-o09-0329-6
DOl: 10.1007/978-94-009-0329-6
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Contents
List of Referees xiii
List of Sponsors XIV
Welcome and Network Information -Norman Darvey
I - Triticale: Today and Tomorrow
Triticale: present status and challenges ahead - George Varughese 13
Triticale - today and tomorrow - Rolf Schlege 21
Marketing - Dilemma or Dream - Cristopher Green 33
How Germany became the number 3 in the triticale world and why I am
confident about the future development - Peter Kruse 41
The verdict in triticale - The case for - Kath Cooper 45
The verdict of triticale - A critical view - Tamas Lelley 49
Tritordeum: triticale's new brother cereal - Antonio Martin, C. Martinez-Araque,
D. Rubiales, J Ballesteros 57
A retrospection on triticale - Enrique Sanchez-Monge 73
Less known studies on triticale in central and Eastern Europe -Anton Tanjsek
and Ivan Kreft 83
II - Cytogenetics
Triticale genomic and chromosome's history -Nicolas Jouve and Consuelo Soler 91
Control of nucleolar expression in triticale -J Perry Gustafson and
R. B. Flavell 119
Expression of lR rDNA loci in triticale: genetic and developmental controls -
Wanda Viegas, M Silva, N. Neves, A. Castilho, A. Bariio, A. Queiroz,
Heslop-Harrison, L. Morais-Cecilio, L. Amado, M Delgado and
Vitor Carvalho 127
The expression of nucleolus organizer regions (Nors) in bread wheat x rye
hybrid lines carrying a lRS/lBL chromosome translocation -JP. Silva,
R. Vieira, N. Jouve and T. Mello-Sampayo 135
Can bread-making quality be introduced into hexaploid triticale by whole-
chromosome manipulation? - E. Kazman and T. Lelley 141
Intergenomic interaction in wheat x rye hybrids non-additive expression of
gliadins and isozymes - Victor Carvalho,and W. Viegas 149
Comparative analysis of telomeric heterochromatin of rye chromosomes in rye
and triticale by fish -Angeles Cuadrado and Nicolas Jouve 155
Production and fertility of hexaploid primary triticales - G. Kleijer and
A. Fossati 165
VI
Study of hybrids between 2n=6x=42 and 2n=6x=42 triticale forms.
I. investigations in F2 - Stoyan M Tsvetkov 173
Tetraploid triticale from Aegilops squarrosa L. x Secale L. spp. -A. Cabrera,
D. Rubiales, J. Ballesteros and A. Martin 179
Molecular cytogenetics analysis of triticale x tritordeum F 1 hybrids - Jose
Lima-Brito, Pat Heslop-Harrison and Henrique Guedes-Pinto 183
Different chromosome combinations on tetra-and hexaploid level from hybrids
of tetraploid rye x tetraploid triticale - Barbara Apolinarska 189
Cytological analysis of F 1 Aegilops hybrids with Triticosecale - D. Gruszecka,
C. Tarkowski, G. Stefanowska and K Marciniak 195
C-banding characteristics of rye chromosomes in winter hexaploid triticale
crossed with maize (Zea mays) - Maria Rogalska Stanislawa 203
Cytological disorders in hexaploid triticale (Triticosecale Wittmack). Association
and stability between two indexes -A. Juarez, A. Ordonez, R. Maich and D.
Manero De Zumelzu 207
Gelelectrophoretic gliadin patterns of euplasrnic and alloplasmic primary triticale
and the corresponding wheat parents - Thomas Gunther, Claus-Ulrich Hesemann
and Gitta Oettler 211
Development of a dominant male-sterile triticale (Ms3) for recurrent selection
and hybrid seed production - S. Venkatanagappa and N. L. Darvey 217
Cms system in hexaploid triticale - Roman Warzecha, Krystyna Salak-Warzecha
and Zygmunt Staszewski 225
Genetic control of non-starch polysaccharides in wheat rye addition
lines-Malgorzata Cyran 233
In - Germplasm
Triticale germplasm: a basic tool for plant breeding - B. Skovmand, W Pfeiffer
and G. Varughese 243
The contribution of rye germplasm towards cereal improvement -A. Mujeeb-
Kazi, Mdhm William, RL. Villareal, RJ. Pena, S. Rajaram, MN. Islam-Faridi,
L. Gilchrist and G. Varughese 253
Assembly and analysis of a North American triticale genetic resource collection -
Calvin 0. Qualset, B.J. Furman, J. Heaton, B. Skovmand and D. Wesenberg 261
Establishment of a network for genetic resources - Norman L. Darvey, G.
Oettler and D. WH Pfeiffer 269
Dwarfmg genes of wheat and rye and its expression in triticale -Andreas
Borner and Jens Plaschke 275
Transfer of genes Rht 1, Rht 2, and Rht 3 from wheat to triticale - C. Tarkowski,
D. Gruszecka, J. Bichta and K Kowalczyk 281
Behaviour of triticale germplasm selected at ENMP by introgression of winter
genes on spring genotypes - B. Mar;iis, J. Coutinho, F. Bagulho, C. Gomes and
N Pinheiro 285
Vll
Variation of some physiological and spike characters affecting the reproductive
behaviour of introgressive triticale lines with T. monococcum genetic
information - W Sodkiewicz and M Tomczak 291
Biological potential of triticale germplasm (X Triticale M.) and its utilization in
breeding - RychtfIrik Julius 299
The analysis of heretability and combining ability of the protein content in
triticale - Sun Yuanshu and Zhang Yuqing 309
Evaluation of the triticale breeding materials with respect to their reaction to
fungal diseases - Stanislaw Wegrzyn, Anna Strzembicka, Helena Grzesik and
Zojia Gajda 315
Facultative triticale: hydroponic evaluation of doubled haploids with superior
mid-generation progenies -A. Arzani and NL. Darvey 321
IV - Biotechnology
Biotechnology for basic studies and breeding of triticale -J Zimny and H Lorz 327
RFLP markers and their applications in cereal breeding - Katrien M Devos and
Mike D. Gale 339
Somaclonal variation detected on the molecular level in the mitochondrial
genomes of regenerated primary CMS Triticale plants - R. Weigel, TH Stoesser,
M Wolf, M Stierle and C-U Hesemann 349
Investigations on morphogenic suspension cultures initiated from mature
embryos in triticale - F. J Maier, G. Gettler and C-U Hesemann 357
Response of mature triticale embryos to aluminium-toxic callus induction media -
Cs. Gaus, G. Gettler and C-U Hesemann 365
Plant regeneration an in vitro selection of triticale - G. Kleijer, NN Diop and
A. Fossati 373
Induction of haploids in triticale (X Triticosecale Wittmack) by crossing it with
corn (Zea mays) - Stanislawa, Maria Rogalska, J Cybulska-Augustyniak and
Wojciech Mikulski 379
Endosperm protein of androgenetic double-haploid lines of 6x-triticale - Juan
Manuel Gonzalez, Ana Fuertes, Nicolas Jouve and Sylvie Bernard 383
Positive effect of Timopheevi cytoplasm on anther culture response of triticale -
A. Arzani and NL. Darvey 391
Genetic control of anther culture induction in triticale - L.N Kaminskaya, L. V
Kotyljova and S.N Matveenko 395
Callus formation and the culturability of inflorescence primordia of the mutant
triticales - Gallia Butnaru 401
Isozyme and endosperm protein markers in the determination of chromosomal
constitution in X Triticosecale Wittmack - Pilar Rubio Nicolas Jouve
Consuelo Soler and Angeles Bernardo ' '409
viii
v -Abiotic Stresses
Stress tolerance in newer triticales compared to other cereals - Robin S. Jessop 419
High levels of salt tolerance revealed in triticale - Robert MD. Koebner and
P.K. Martin 429
Aluminum tolerance in triticale, wheat and rye -Augusto C Baier, D.J Somers
and lP. Gustafson 437
Screening cultivars for aluminium tolerance -Ana Maria Antunes, Jose Pereira
and M Antonieta Nunes 445
Screening methods for aluminium tolerance in seedlings of triticale - D. Morath,
G. Gettler and A.E. Melchinger 453
The variability of aluminium tolerance among triticale strains and cultivars bred
in Poland- Andrzej Aniol 461
Seedling tolerance to aluminium in spring triticale from Poland and CIMMTY -
Budzianowski G., W Mackowiak, K. Paizert and B. Apolinarska 467
Changes in pH of the root medium produced by cultivars of triticale -Ana
Maria Antunes, Jose Pereira and M Antonieta Nunes 475
Aluminum toxicity and protein synthesis in near isogenic lines of Triticum
aestivum L. differing in aluminum tolerance - Daryl J Somers, Keith G. Briggs
and 1 Perry Gustafson 481
Responses of triticale anJ wheat to hypoxia -1 W Johnson and B. Huang 489
VI - Diseases
Triticale diseases - a review - Edward Arseniuk 499
Screening for fusarium scab resistance in triticale - Mariana Ittu, GH Ittu and
NN Saulescu 527
Inheritance of head resistance of winter triticale to Phaeosphaeria nodorum -
Henryk Wos 535
Evaluation of new winter octoploid triticales for their reaction to single and
combined infections with bydv and Typhula ishikariensis Imai - B. Bizimungu,
J Collin, CA. ST-Pierre and A. Comeau 541
Evaluation of selection criteria for tolerance to Fusarium graminearum in
triticale - Rebeca M Gonzalez, Castrejon CA. and Eulalio Venegas Gonzalez 549
Resistance of triticale to root lesion nematode in south Australia - V A.
Vanstone, M Farsi, A. 1 Rathjen and K. V Cooper 557
VII - Triticale Breeding
Breeding strategies for triticale -Augusto C Baier, J Perry Gustafson 563
Triticale: potential and research status of man-made cereal crop - W H Pfeiffer 571
Semidwarf winter triticale - Tadeusz Wolski and Jaroslaw Gryka 581
Developing Spring and winter triticale with reduced-awn expression - Don F.
Salmon, lH Helm and P.E. ledel 589
ix
Outcrossing in winter triticale, measured by occurence of tall plants - W Sowa
and HKrysiak 593
Estimates of combining ability for some traits in triticale - Stanislaw Wegrzyn,
Helena Grzesik 597
Effect of low nitrogen input on agronomic traits in triticale - Gitta Gettler 603
Genetic variation, G*E interactions and selection response for hagberg falling
number in triticale -L. Jestin and H Bonhomme 609
Preharvest sprouting resistance in triticale: preliminary results - G. Haesaert
and A. De Baets 615
Facultative triticale: comparison of local and CIMMYT germplasm in
hidroponics and field conditions -A. Arzani and NL. Darvey 623
An attempt at tetraploid triticale improvement - Boguslaw Lapinski, Barbara
Apolinarska, Grzegorz Budzianowski, Malgorzata Cyran and Maria Rakowska 627
Some remarks on triticale breeding evolution in Portugal. a) Role played by the
National Plant Breeding Station - Francisco Bagulho, M T. Barradas, B.Mar;as
and J Coutinho 635
Triticale research and breeding programmes in France: recent developments -
Bernard M, Bernard S, Bonhomme H, Faurie c., Gay G. and L. Jestin 643
Triticale breeding in Switzerland - A. Fossati, D. Fossati and G. Kleijer 649
Crop management of triticale in Belgium - G. Haesaert, J Latre and A. de
Baets 655
Triticale breeding in Fundulea - Romania - GH Ittu and NN Saulescu 663
Breeding triticale for the south western cape - HS Roux and G.F. Marais 669
Registration of three new triticale varieties: Crato, Arruda and Alter -J
Coutinho, B. Mar;as"F. Bagulho, M Conceir;iio Gomes and JCoco 675
Characterization of lamb-2 a new triticale variety for the highly variable
moisture situation of central Mexico - Rebeca M Gonzalez Iniguez and
W H Pfeiffer 679
VIII - Regional Trials and Crop Management
Adaptation of triticale to Southern Brazil -Augusto Carlos Baier 687
Triticale at high latitudes in Alberta Canada - D.F. Salmon, VS Baron, JG.
Mcleod and JH Helm 693
Regional trials on triticale - National list -Ana Paula A. Cruz de Carvalho 701
Quantitative and qualitive components of eleven greek triticale cultivars (X
Triticosecale Wittmack) compared to cv. "Vergina" (T. aestivum L. em. Thell.) -
D.M Gogas, N Triantaphilakos, S Stratilakis, S GeorgiadiS, T. Avramidis, G.
Skipitaris, H Kartitsi, H Tsivopulu and I Eystathiu 709
Triticale in Morocco: a promising crop - M Mergoum 719
Performance of drira out-cross of triticale - SK Yau 725
Triticale cultivation for diversifying and stabilizing a wheat dominated
agro-ecosystem - G.S Dhindsa and Jaswinder Singh 731
x
Some physiological features of triticale and their consequences on crop
managment in France - Georges Laroche and Gate Philippe 737
Succesive effect of herbicides on triticale seeds germination and plant growth -
Slawomir Stankowski and Robert Maciorowski 743
Comparative weed supression by triticale, cereal rye and wheat - D. Lemerle
and K Cooper 749
IX - Triticale Uses
Factors affecting triticale as a food crop - Roberto J. Pena 753
Triticale malting - an evaluation of characteristics and production - M L. Bairiio
da Costa and M J. Cabo Verde 763
Relationship between the pentosans of triticale flour and bread loaf volume - M
Cyran and M Rakowska 771
Rheological and biochemical properties of meal from winter triticale varieties in
connection with the resistance to preharvest germination - Stanislav Grib,
Mikhail Shishlov and Natalia Shishlova 779
Environmental and genotypical influences on triticale grain quality in Northeast
of Portugal - J. S. Oliveira, Benilde Mendes, Ana Fernando, Rui Fernandes, H
Guedes-Plnto, 0. Pinto-Carnide, V. Carnide, Bruno de Sousa, F. Cabral and
J. Baieta 785
Concentrations of minerals in the grains of a high-and a low-protein winter
triticale - D. Fossati, A. Fossati and B.Feil 793
Breadmaking quality in tritordeum: the use-possibilities of a new cereal -Juan
B. Alvarez and Luis M Martin 799
Influence of feeding growing-finishing pigs triticale, wheat or maize based diets
on reSUlting carcass composition, and on taste and quality characteristics of
pork- JH Brendemuhl, R.o. Myer and D.D. Johnson 807
Synthetic amino acid supplementation of triticale and wheat based diets for
growing-finishing pigs - R.o. Myer, JH Brendemuhl and R.D. Barnett 813
Nutritive value and feed efficiency of broiler diets containing different levels of
triticale - El-Yassin Fayez, Haj-Omar Nedal and Abboud Mousa 819
Evaluation of triticale as a forage plant through the analysis of the cinetic of
some qualitative parameters from stem elongation to maturity - S. Bocchi, G.
Lazzaroni and T Maggiore 827
Effect of the seeding rate on forage yield and quality of triticale - V. Carnide,
H Guedes-Pinto, A. Mascarenhas-Ferreira and C. Sequeira 835
Aptitute of spring and winter triticales for dual-purpose (forage and grain) in
mediterranean conditions - Conception Royo, E. Penella, F. TriM and J.L.
Molina-Cano 843
The effects of sowing date on the growth and yield of dual-purpose and grazing
triticales under cool Australian conditions -J. A. Agyare, R. L. Wright and
R.S. Jessop 851
