THE TYPE SPECIMENS OF BROMELIACEAE IN THE HERBARIUM OF THE MUSEUM OF NATURAL HISTORY IN VIENNA, AUSTRIA

18 Pages·1994·1.3 MB·

by  Till, Walter

Selbyana 15: 94-111
THE TYPE SPECIMENS OF BROMELIACEAE IN THE
HERBARIUM OF THE MUSEUM OF NATURAL HISTORY IN
VIENNA, AUSTRIA
WALTER TILL
Botanical Institute, University of Vienna, Rennweg 14, A-1030 Vienna, Austria
ABsTRACT. The fate of the herbarium of the Museum of Natural History in Vienna, Austria, during
World War II is briefly reported. A complete and annotated account of the type specimens ofBromeliaceae
still kept there is presented including type photographs from other herbaria. The following basionyms have
been lectotypified in this paper: Aechmea organensis, Aechmea petropolitana, Tillandsia globosa, Tillandsia
ventricosa, Vriesea carinata, Vriesea carinata var. inflata, and Vriesea paraibica.
INTRODUCTION been reported (Till, 1987). The following account
comprises all type specimens of Bromeliaceae
The Museum of Natural History in Vienna,
which have survived World War II and which
Austria, houses one of the most important bo
have been acquired since then to December 1993.
tanical collections in the world. The nearly four
Included are type photographs from other her
million specimens include many of the classical
baria and a few specimens which cannot be iden
collections (Holmgren et ai. 1990) which contain
tified with full certainty as types but which are
an extraordinarily high number of type speci
suspicious. Publication dates and abbreviations
mens.
of protologue literature for books follow Stafleu
With regard to Bromeliaceae, before World
& Cowan (1976,1979,1981,1983,1985, 1986,
War II the Museum's phanerogamic collections
1988), those for journals Lawrence et aI. (1968)
had been especially important for the specimens
and Bridson & Smith (1991).
ofF. Antoine, J.S. Blanchet, T.P.X. Haenke, N.J.
All entries first provide the name and the pro
Jacquin, C.F.P. von Martius, E.F. Poeppig, J.E.
tologue citation, then, in quotation marks, the
Pohl, H. Wawra von Fernsee, H.R. Wullschliigel,
original label information, followed by deter
and of many others. Unfortunately this situation
mination(s) in chronological order. Ifexistant or
changed in 1945.
known, photographs of the specimens in other
Alarmed by the tragedy of the Herbarium and
herbaria are cited. The herbarium abbreviations
the Library of the Botanical Museum Berlin
follow Holmgren et. aI. (1990). Additional re
Dahlem in March 1943, approximately 12,000
marks from the author are found under the para
fascicles and the library of the Botanical De
graph notes. Article 9.1 of the ICBN (Greuter et
partment of the Vienna Museum were trans
aI., 1988) has been strictly applied. Photographs
ferred during the summer of 1943 to seven dif
of the types of several hitherto not (or insuffi
ferent localities: the library within Vienna, the
ciently) figured taxa are provided.
Herbarium to Gaming, Kirchstetten near Staatz,
Klosterneuburg, Lunz, Ober-Hofiein near Retz,
and Purgstall, all villages in Lower Austria. The RESULTS
collections were returned in November 1945 and Bromelioideae
the spring of 1946 (Petrak, 1948).
After the end of war activities, the Botanical Aechmea alopecurus Mez in Martius, Fl. bras.
Department was informed in July 1945 that part 3(3): 367. 1892. "PohI5230, Brasilia", HO
of the material stored in Ober-Hofiein had been LOTYPE: W.-Macrochordium ? Vriese,
damaged by fire. Petrak (1948) listed all the fam det. Beer; Aechmea alopecurus Mez n.spec.,
ilies which had been destroyed completely or for det. C. Mez.
the major part, among them Bromeliaceae. This NOTE: One inflorescence only. For more
list may be the reason why the Bromeliaceae of details see Till (1987).
the Museums Herbarium (acronym: W) generc Aechmea ampla L.B.Smith, Phytologia 24: 433,
ally have been considered as to be lost. pI. 2, Figs. 5-7. 1972.
"Bahia, Canavieiras, Na restinga,
29.1.1965, leg. Lanna 742 et Castellanos
25491", HOLOTYPEand ISOTYPE: Herb.
MATERIAL AND METHODS
Centro Pesquisas Florestais e Conservacao
In a prior paper the survival of the type of da Natureza-Tijuca-Rio de Janeiro, pho
Aechmea a!opecurus Mez in Martius has already tos W.
94
1994] TILL: BROMELIADS-TYPE SPECIMENS IN VIENNA, AUSTRIA 95
FIGURE 1. Aechmea angustijolia.
Aechmea angustifolia Poeppig & Endlicher, Nov. rosette with a poor inflorescence on the right
gen. sp. pI. 2: 43, pI. 159. 1838. FIGURE 1. side. At the bottom of the sheet are two
"Peruvia, Provo Maynas alta. Ad arbores labels, that ofPoeppig s.n. (left) and ofPoep
circum Missionem Tocache, luI. 1830, pig 1823 (right). It is uncertain which label
Poeppig 1823", LECTOTYPE: W; "May belongs to which individual plant specimen.
nas. In arbor. ad Yurimaguas, 1831, Poeppig From the arrangement of the specimens and
s.n.", SYNTYPES: W.-Hop!ophytum Br., the labels, however, it seems logical to con
det. Beer; Aechmea angustifolia, det. C. Mez. sider the rosette with the poor inflorescence
NOTE: The sheet in W bears two inflo as the prior collection (= Poeppig 1823) and
rescences on the left half and a complete the two inflorescences as a later gathering (==
96 SELBYANA [Volume 15
Poeppig s.n.) to add the missing inflores W. - Portea petropolitana Mez, det. Carl Mez
cence characteristics. If these arguments are III/93; rev. W. Till 1980.
acceptable, L.B. Smith's choice of a lecto NOTE: Six inflorescence fragments in
type ("Poeppig 1823, holotype, W", Smith postflowering state are mounted on two
& Downs, 1979) must include the rosette sheets. Sheet # 651 (Museum's numbering
specimen. It should further be noted that C. at right top corner) is chosen here as the
Mez had seen Poeppig 1823 also from her lectotype. Currently named Portea petro
barium G-BOIS. That specimen would rep politana (Wawra) Mez in Martius.
resent an isolectotype. Bromelia melanantha Ker, Bot. Reg. 9: pI. 766
Aechmea friedrichsthalii Mez & Donnell Smith (1824)."de Lambert", ISOTYPE: W.-Bro
in Donnell Smith, Bot. Gaz. (Crawfords melia melanantha Bot. Reg.; [Bromelia] bi
ville) 19: 263. 1894. color.
"Guatemala. Ins. Catina in fl. So. Juan. NOTE: The specimen consists of two
Parasita. 1841 Friedrichsthal 609.", LEC leaves and one inflorescence which are glued
TOTYPE: W. Ae. micrantha Mez n.sp., det. on a sheet of 39,4 x 27 cm. The original
C. Mez; Aechmea Friedrichsthalii Mez et J.D. sheet in the typical British format had been
Smith n.sp., det. C. Mez 1894. glued in 1889 on a sheet of the Vienna mu
NOTE: One inflorescence and two leaves. seum's format when the collection of Rei
Currently treated as synonym of Aechmea chenbach filius had been acquired and this
penduliflora Andre. individual sheet had been numbered #
Aechmea noettigii Wawra, Oesterr. Bot. Z. 30: 262527. Due to this remounting the hand
117.1880. writing at the backside top of the original
"Reise d. Prinz. Aug. u. Ferdin. v. S. sheet had been obscured: "Hort. Boyt on
Coburg nach Brasilien., 1879., Nro. 108., A.B.L. 1824". The comparison with the
Aechmea Nottigii Wwr., Entre Rios, Dr. H. handwriting illustrated in Miller (1970: 492)
Wawra.", HOLOTYPE: W.-Portea Noet leaves no doubt that it originates from Ayl
tigii Mez, det. Carl Mez 111193; Portea pe mer Bourke Lambert himselfw ho cultivated
tropolitana var. noettigii (Wawra) L.B. Smith, the Bromelia melanantha at Boyt on House
Typus, det. W. Till 1980. near Heytesbury, Wiltshire. Lambert's fa
NOTE: One inflorescence and three leaves. mous herbarium was sold by auction and
Currently named Portea petropolitana (Wa one lot was obviously bought by Reichen
wra) Mez in Martius var. noettingii (Wawra) bach pater. Ker (1824) stated: "We were
L.B. Smith. supplied with the plant for our drawing by
Aechmea organensis Wawra, Oesterr. Bot. Z. 30: Mr. Lambert ..." . As it remains unsolved
116. 1880. whether Ker returned the plant to Lambert
"Reise d. Prinz. Aug. u. Ferdin. v. S. or prepared a herbarium specimen to be kept
Coburg nach Brasilien., 1879, Nro. 317., with him, the holotype hence is considered
Aechmea organensis Wwr., Serra d. Orgaos, as to be lost. The specimen in W represents
Dr. H. Wawra", LECTOTYPE (designated an unremembered isotype. The species is
here) and ISOLECTOTYPES: W.-Aech currently named Maeroehordion bromeli
ifolium (Rudge) Beer.
mea organensis Wawra, det. C. Mez 1894).
Bromelia paniculigera Reichenbach, Iconogr. bot.
NOTE: The sheet bears one flowering ro
exot. 3: 14, tab. 239, 240.1830; non Swartz,
sette, three additional inflorescences and one
1788.
additional leaf. In agreement with Art. 9.1
"Bromelia paniculigera Sw., Rchb. Bot.
oflCBN (Greuter et aI., 1988) the flowering ex. 239 Orig." "Bromelia paniculigera Sw,
rosette is chosen as the lectotype as it is the Reich. Bot exot t 240 (Orig)".
basis for pI. 23 of Hin. princ. S. Coburgi NOTE: There are two museum sheets each
(1883). A sequence of these illustrations is bearing (glued) two drawings and paintings
present in the museum's archive: one pencil respectively. Although Reichenbach's name
drawing, one hand-colored engraving, and is a younger homonym of Bromelia pani
one preprint of plate 23. culigera Swartz, the original icons demon
Aechmea petropolitana Wawra, Oesterr. Bot. Z. strate Reichenbachs interpretation of the
30: 116. 1880. name concerned. The species is currently
"Reise d. Prinz. Aug. u. Ferdin. v. S. named Aechmea aquilega (Salisbury) Gri
Coburg nach Brasilien., 1879., Nro. 63. sebach var. aquilega.
Aeehmea petropolitana Wwr., Petropolis Bromelia strobilina Beurling, Kongl. Svenska
(Alto Imperador), Dr. H. Wawra.", LEC Vetensk. Acad. Hand!. 15: 110. 1854.
TOTYPE (designated here) and ISOTYPES: "Bromelia strobilina Beurl., Porto-Bello,
1994] TILL: BROMELIADS-TYPE SPECIMENS IN VIENNA, AUSTRIA 97
in rupibus marinis; April 1826: Billberg.", "The New York Botanical Garden Ven
ISOTYPES: S, photo W. ezuelan Expedition 1957-58, Cerro de La
NOTE: The sheet in S bears two inflores Neblina, Rio Yatua, Territorio de Amazo
cences and one leaf, all representing isotypes nas, Bassett Maguire, John J. Wurdack, and
according to Art. 9.1 ofICBN. No lectotype Celia K. Maguire 42207, November 24,
has been hitherto chosen. Currently named 1957", ISOTYPES: W.
Aechmea strobilina (Beuding) L.B. Smith & NOTE: The sheet bears two flowering ro
Read. 1 settes. No herbarium is mentioned in the
J?.
Prantleia glabra Mez in Martius, bras. 3(3): protologue to keep the "type", so the Smith
258, pI. 58, Fig. 1. 1891. & Downs (1974) citation ofa "holotype" at
"3436. Inhumaes. Pohl.", SOLECTO US actually represents the lectotype.
TYPE: W. - Orthophytum glab m Mez, det. Navia affinis L.B. Smith in Maguire and Wur
C. Mez. ~' dack, Mem. New York Bot. Gard. 10(2): 21,
NOTE: Mez had seen Pohl 436 also in Fig. 9 a,b. 1960.
Brussels (BR). Smith & Downs (1979) have "The New York Botanical Garden Ven
chosen that specimen as the Ie otype ("ho ezuelan Expedition 1957-58, Cerro de La
lotype"). Currently named rthophytum Neblina, Rio Yatua, Territorio de Amazo
glabrum (Mez in Martius) Me in A. et C. nas, Bassett Maguire, John J. Wurdack, and
DC. Celia K. Maguire 42470, December 24,
Prantleia leprosa Mez in Martius, . bras. 3(3): 1957", ISOTYPES: W.
259, tab. 58, Fig. 2. 1891. NOTE: The sheet bears two complete ro
"5229. Cacho d'Infemo. P hl.", LEC settes. As in Brocchinia delicatula, the spec
TOTYPE (Smith, 1955): W.-lrthoPhytum imen in US is the lectotype.
leprosum Mez, det. C. Mez. Navia ftuviatilis L.B. Smith in Maguire and Wur
NOTE: Currently named rthophytum dack, Mem. New York Bot. Gard. 9(3): 298,
leprosum (Mez in Martius) Met in A. et C. Fig. 27 a-g. 1957.
DC. I "Plants of the Pakaraima Mountains,
PseudaechmeaambiguaL.B. Smith&R.W. Read, British Guiana-1951, Membaru-Kuru
Phytologia 52: 53, pIs. 3, 4. 1982. pung Trail, Bassett Maguire and D.B. Fan
"Flora Colombiana. Herbario Departa shawe 32360, October 29-November 4,
mento de Biologia. Universidad de Antio 1951", ISOTYPES: W. - Navia rupestris
quia. Bromeliaceae. Epiphyte. Inflores (Gleason) Sandwith, rev. W. Till 1980.
cences showy, fl bracts bright red, rhachis of NOTE: The sheet bears two just-post
inflorescences yellowish orange. Pedocens flowering rosettes. Currently considered as
noght pink, base offl buds green, unopened a synonym of Navia rupestris (Gleason)
fls bright violet-blue. Periphery of primary Sandwith.
forest. Departamento de Antioquia, Muni Navia nubicola L.B. Smith in Maguire and Wur
cipio de Zaragoza, Corregimientode Provi dack, Mem. New York Bot. Gard. 9(3): 303,
dencia, trail in advanced humid secondary Fig. 38 a-d. 1957.
tropical rain forest, vicinity of the hydro "The New York Botanical Garden Ven
electric plant. alt. 500-650 m. Colector: D.D. ezuelan Expedition 1953-54, Cerro de la
Soejarto y J.D. Villa No: 2703. Fecha: Feb Neblina, Rio Yatua, Territorio de Amazo
ruary 9, 71", HOLOTYPE: ECON, photo nas, Bassett Maguire, John J. Wurdack, and
W; ISOTYPE: ECON, photo W. George S. Bunting 37382, Jan. 24, 1954",
Streptocalyx biftoms L.B.Smith, Phytologia 24: PARATYPE: W.
448, pI. 5, Figs. 15, 16. 1972. Navia ocellata L.B. Smith in Maguire and Wur
"Plants of Ecuador. Provo Pastaza. Strep dack, Mem. New York Bot. Gard. 9(3): 310,
tocalyx bif/orus L.B. Smith. Type. Topo Fig. 51 a-c. 1957.
along Rio Topo; tropical rain forest; elev. "The New York Botanical Garden, Plants
1300 m. Bracts yellow; flowers pink; leaves of the Kunhardt Venezuelan Expedition,
deep red; epiphytic on tree trunk. C.H. Dod 1948-49, Cerro Sipapo (Panique), Territorio
sonandL.B. Thien No. 2070. Jan. 11, 1962", Amazonas, Bassett Maguire and Louis Politi
HOLOTYPE: WIS, photo W. 27866, December 23, 1948", PARA
TYPES:W.
NOTE: The sheet bears two complete ro
Pitcairnioideae
settes.
Brocchinia delicatula L.B. Smith in Maguire and Navia trichodonta L.B. Smith in Maguire and
Wurdack, Mem. New York Bot. Gard. 10(2): Wurdack, Mem. New York Bot. Gard. 9(3):
19, Fig. 6 a-e. 1960. 301, Fig. 32 a-d. 1957.
98 SELBYANA [Volume 15
"Plants of the New York Botanical Gar ly unlikely that such a striking plant which
den Venezuelan Expedition, 1950-51, Cerro had been collected only two times in the past
Huachamacari, Rio Cunucunuma, Amazo century would have received the same num
nas, Bassett Maguire, R.S. Cowan, and John ber by two different collectors. The sheet in
J. Wurdack 30228, 1950", ISOTYPES: W. W is therefore considered as an isotype. The
NOTE: The sheet bears two complete ro plant is currently named Guzmania pearcei
settes. (Baker) L.B. Smith.
Pitcairnia reftexiftora Andre var. minor L.B. Tillandsia acbyrostacbys E. Morren ex Baker var.
Smith & R.W. Read, Phytologia 30: 290. stenolepis L.B. Smith, Contr. U.S. Natl.
1975. Herb. 29: 505, Fig. 68e. 1951.
"Plants of Ecuador. Pitcairnia reflexiflora "e.G. Pringle, Plantae Mexicanae. 1892.
Andre var. minor L.B. Smith. Type. Km. State of Jalisco. 4111 Tillandsia variabilis,
110 from Duran; t[a]ropical cloud forest; Schi. ex char. On Burseras, hills near Gua
elev. 1300 m.; Flowers red; side of embank dalajara. 25, June.", ISOTYPE: W.-Til
ment. e.H. Dodson and L.B. Thien 2097, landsia achyrostachys E. Morren var. sten
Jan. 15, 1962", HOLOTYPE: WIS, pho olepis L.B. Smith, det. W. Till, 1979.
toW.
Tillandsia araujei Mez in Martius, Fl. bras. 3(3):
Pitcairnia ruderalis L.B.Smith, Phytologia 18:
600, pI. 112, Fig. 2. 1894.
139, Figs. 10, 11.1969. "5516. Tillandsia apicrioides Schott. Bra
"Pitcairnia ruderalis L.B. Smith. Type.
silia. Schott.", SYNTYPES: W.-Nomen
[Peru, Loreto.], Obenteni on + pajonal edge
publici juris non factum, ob Catopsidis spe
or old chacra, ca. 1150 m. Also seen else
ciem eodem signatam neglectum, Mez; Til
where in Gran Pajonal in old chacras and landsia araujei Mez n.sp., det. Carl Mez.
moist sites (where probably most common).
NOTE: The sheet bears one leafy stem
M.S. Chrostowski No.: 66-5b. Date: Aug. 13,
with the inflorescence lost at the left side
1966. Herbarium University of Wisconsin
and one branched leafy stem of which each
Madison", HOLOTYPE: WIS, photo W.
branch terminates with an inflorescence at
the right side.
Tillandsioideae Tillandsia azurea Presl, Reliq. haenk. 1: 124, pI.
24. 1827.
Guzmania cuzcoensis L.B. Smith, Phytologia 24:
"Tillandsia azurea. Presi. Haenke 96.",
445, pI. 5, Figs. 7-8. 1972.
ISOTYPE: W.-Tillandsia II, Beer; Till.
"Peru, Provo La Convencion, Dept. Cuz
straminea HBK., det. C. Mez.; Tillandsia
co, 73°36'W; 12°38'S, T.R. Dudley 11317,
purpurea Ruiz & Pavon, rev. W. Till 1980.
23 July 1968", HOLOTYPE: NA, photo W.
NOTE: The specimen had been acquired
Guzmania seemannii Baker, Handb. BromeI. 153.
by the museum in II. 1829. "Haenke 96."
1889.
is written in the same red ink as had been
"No. Tillandsia. An Baumen zwischen
used for the acquisition date. Treated as a
Ona & Saraguro Provo Loja in Ecuador
synonym of Tillandsia purpurea Ruiz-Lo
10000'. Bliithenstand hellroth Blumen gelb.
pez & Pavon y Jimenez (Smith & Downs,
August 1878. leg. Lehmann Ecuador".,
1977) but more recently sometimes sepa
SYNTYPES: W.-Tillandsia seemannii
rated from the former under T. straminea
(Baker) Mez, det. L.B. Smith III, 1959.
H.B.K.
NOTE: The sheet bears two rosettes with
inflorescence. Currently named Racinaea Tillandsia band ens is Baker var. inter media
seemannii (Baker) Spencer & Smith. Hassler, Annuaire Conserv. Jard. Bot. Ge
Sodiroa pearcei Baker, J. Bot. 25: 53. 1887. neve 20: 333. 1919.
FIGURE 2. "Dr. E. Hassler, Plantae Paraguarien
"416, Tillandsia 2, East. And.", ISO ses.-1903. Plantae a custode herbarii mei
TYPE: W. - Guzmania pearcei (Baker) L.B. T. Rojas in Chaco septentrionali lectae. No.
Smith, det. L.B. Smith III, 1959. 2778 Tillandsia bandensis var. intermedia
NOTE: The sheet bears an additional la HassI. (ex Hassler). Gran Chaco: Santa Elisa
bel from the Vienna Museum which attri lat. S. 23°10'. Lect. mens. Februarius", ISO
butes the plant to [W.] Lobb with Peruvian TYPES: W.
provenance. As both R.W. Pearce and W. NOTE: The sheet bears three flowering
Lobb collected plants for J.G. Veitch, it is plants. Currently not separated from T. ban
very persuasive to assume that the collection densis.
has erroneously been attributed to Lobb by Tillandsia bangii Baker in Rusby, Mem. Torrey
the museum's staff. It is, moreover, extreme- Bot. Club 6: 124. 1896.
1994] TILL: BROMEUADS-TYPE SPECIMENS IN VIENNA, AUSTRIA 99
FIGURE 2. Sodiroa pearcei.
"Plantae Bolivianae. A Miguel Bang Lec Tillandsia bicolor Brongniart in Duperrey, Voy.
tae. Ex. Herbario Collegii Columbiae, a N.L. monde 185. 1834, Atlante pI. 36. 1826.
Britton et H.H. Rusby distributae. Tilland "Tillandsia. parasita. He Sa. Catherine,
sia. Vic. Cochabamba. No. 1283. 1891.", Bresil. d Urville", ISOTYPE: W.-Tilland
ISOTYPE: W. - Tillandsia deppeana Steud. sia aeranthos (Loisel.) L.B. Smith, det. L.B.
var. lati/olia (Griseb.) L.B. Smith, det. L.B. Smith (III 1959); Tillandsia tenuifolia L. var.
Smith (III 1959); Tillandsia fendleri Grise tenuifolia, rev. W. Till 1980.
bach var. fendleri, rev. W. Till 1980. NOTE: Currently treated as a synonym of
NOTE: Currently treated as a synonym of Tillandsia aeranthos (Loiseleur Deslongch
Tillandsia fendleri Griseb. var. fendleri. amps in Mordant de Launay) L.B. Smith.
100 SELBYANA [Volume 15
Tillandsia boliviana Mez, Bull. Herb. Boissier sia globosa Wwr. Entre rios. Dr. H. Wa
ser.2. 4: 1130. 1904. wra.", LECTOTYPE (designated here: right
"Plantae Bolivianae. A Miguel Bang lec plant): W, ISOLECTOTYPE (left plant):
tae. Ex. Herbario Collegii Columbiae, a N.L. W.-Till. globosa Wawra, det. Carl Mez.;
Britton et H.H. Rusby distribu Specim. exam. Wilhelm Weber, July 1981.
tae.Tillandsia, Probably n. sp. No. 2202", Tillandsia globosa Wawra var. crinifolia Wawra,
ISOTYPE: W.-Tillandsia boliviana Mez, Oesterr. Bot. Z. 30: 223. 1880.
det. L.B. Smith III. 1959. "Reise d. Prinz. Aug. u. Ferdin. v. S.
Tillandsia bryoides Griseb ach ex Baker, J. Bot. Coburg nach Brasilien. Nro. 142c. Tilland
16: 236. 1878. sia globosa var. crinifolia Wwr. Dr. H. Wa
"Flora Argentina No. 128. Tillandsia. Es wra.", HOLOTYPE: W.-Till. globosa Wa
tancia Germania prope Cordoba. Junio-De wra, det. Carl Mez.; Specim. exam. Wilhelm
cembre 1874. Dr. P.G. Lorentz", SYN Weber, July 1981.
TYPES: W.-T. tricholepis Baker, rev. W. NOTE: Currently not separated from T.
Till 1979. globosa Wawra var. globosa.
NOTE: T. bryoides as based on Lorentz Tillandsia. herzogii Wittmack, Meded. Rijks
128 is treated as a synonym of Tillandsia Herb. 29: 89. 1916.
tricholepis Baker (Till, 1984, 1989). The sheet "Plantae in itinere secundo per Boliviam
in W bears four clusters of the Lorentz col lectae. leg. Th. Herzog No. 1151", ISO
lection. TYPE: W.-T. reichenbachii Baker 1889,
Tillandsia flagellata L.B. Smith, Phytologia 6: rev. W. Till 1979.
258, pI. 1, Figs. 9, 10. 1958. NOTE: Currently treated as a synonym of
"United States Department of Agricul Tillandsia reichenbachii Baker.
ture, Gray Herbarium of Harvard Univer Tillandsia hieronymi Mez in A. et C.DC., Mon
sity, New York Botanical Garden. Explo ogr. phan. 9: 876. 1896.
rations in South America. Tillandsia cf "Flora Argentina No. 124. Tillandsia. Es
platyrhachis. On tree. Dry hill. Ecuador: tancia Germania prope Cordoba. Junio-De
cembre 1874. Dr. P.G. Lorentz", ISOLEC
Province Oro. Portovelo (Gold mine near
TOTYPES: W. - Tillandsia capillaris Ruiz
Zaruma). Alt. 600-1000 M, No. 21245, A.S.
& Pavon forma hieronymii (Mez) L.B.Smith,
Hitchcock, Collector, Aug. 30-Sept. 1, 1923."
Syntypus, rev. W. Till 1979.
ISOTYPES: US, photo W.
NOTE: The lectotype has been chosen by
NOTE: The sheet in US bears one fruiting
Till (1984) and is kept in B. The sheet in W
rosette and one infrutescence. No lectotype
bears two clusters of the Lorentz collection.
has hithero been chosen.
Currently treated as a synonym of Tillandsia
Tillandsia funckiana Baker, Handb. Bromel. 196. capillaris Ruiz Lopez & Pav6n y Jimenez.
1889. Tillandsia incana Wawra, Oesterr. Bot. Z. 30:
"No. 1258 Funcke & SchUm, Venezuela, 223. 1880. FIGURE 3.
&c. CoIl. 1846.", ISOTYPE: W. - Tillandsia "Reise d. Prinz. Aug. u. Ferdin. v. S.
andreana E. Morren ex Andre, det. L.B. Coburg nach Brasilien. Nro. 508. Tillandsia
Smith (III 1959). incana Wwr. Itatiaia (Catingaa). Dr. H. Wa
NOTE: Currently accepted as a species wra.", HOLOTYPE: W. - Tillandsia gemi
distinct from T. andreana. niflora Brongn. var. incana (Wawra spec.),
Carl Mez 111/93.
Tillandsia gilliesii Baker, J. Bot. 16: 240. 1878. NOTE: One preanthetic rosette. The sheet
"Tillandsia. Mendoza. Hook. 837", prob bears an additional leaf from a plant culti
ably ISOTYPES: W.-Till. myosura Gris., vated in the "Augarten" in Vienna under
det.CarlMez 1894.; TillandsiagilliesiiBak., the name Tillandsia Rolissonii. Currently
det. A. Castellanos, XI 1935. named Tillandsia geminijlora Brongniart in
NOTE: Hooker himself never had col Duperrey var. incana (Wawra) Mez in Mar
lected in Argentina but had received plants tius.
from Gillies. Hence the four plants on the Tillandsi* lancifolia Baker, Handb. Bromel. 202.
sheet are most probably isotypes as Hook 1889.
er's herbarium had been seen and studied "3458. fl.lutei, ad rupes. Bras. Blanchet.",
by Baker in Kew. ISOTYPE: W.-Vriesea Platzmanni E.
Tillandsia globosa Wawra, Oesterr. Bot. Z. 30: Morr., det. Carl Mez 111/93.
222. 1880. NOTE: Currently named Vriesea lanci
"Reise d. Prinz. Aug. u. Ferdin. v. S. folia (Baker) L.B. Smith emend. Martinelli
Coburg nach Brasilien. Nro. 142b. Tilland- & Leme.
1994] TILL: BROMELIADS-TYPE SPECIMENS IN VIENNA, AUSTRIA 101
FIGURE 3. Tillandsia incana.
Tillandsia laxissima Mez, Bull. Herb. Boissier M.[onte] V.[erde] Dec 30", ISOTYPE: W.
ser.2. 5: 108. 1905. Tillandsia lescaillei C. Wright ex Sauv., Iso
"Plantae Bolivianae. A Miguel Bang Lec type, det. L.B. Smith (III 1959).
tae. Ex. Herbario Collegii Columbiae, a N.L. NOTE: The specimen is recognized as a
Britton et H.H. Rusby distributae. N. Gen. type by C. Wright's characteristic hand
in Bromeliaceae. No. 2301", ISOTYPE: writing. Currently named Racinaea lescail
W. - Tillandsia laxissima Mez, det. L.B. lei (C. Wright ex Sauvalle) Spencer & Smith.
Smith (III 1959). Tillandsia mandonii E. Morren ex Mez in A. et
Tillandsia lescaillei C. Wright ex Sauvalle, An C. DC., Monogr. phan. 9: 871. 1896.
ales Acad. Ci. Med. Habana 8: 52. 187l. "G. Mandon.-P1antae Andium Boli
"Tillandsia. On trees in thick woods. viensium. No. 1180.-Tillandsia. Hab. Pro-
lO2 SELBYANA [Volume 15
FIGURE 4. Tillandsia martinicensis.
va. Larecaja. - Viciniis Sorata; inter San Pe W. - Guzmania p/umieri (Griseb.) Mez, det.
dro et Coaconi, in scopulosis super arbores. L.B. Smith III 1959.
Alt. Reg. tempa. 2650 m. Mart. 186l.", NOTE: The single inflorescence originates
ISOLECTOTYPES: W.-Till. mandonii from the collection of Reichenbach fil. and
Ed.Morr., det. Carl Mez 1894. was acquired by the museum in 1889 under
NOTE: The lectotype has been chosen by the number 132030. It probably represents
Till (1984) and is kept in BM. a syntype. The plant is currently named
Tillandsia martinicensis Baker, J. Bot. 26: 45. Guzmania p/umieri (Griseb.) Mez in A. et
1888. FIGURE 4. C. DC.
"Martinique leg. Hahn.", SYNTYPE ?: Tillandsiamathewsii Baker, J. Bot. 25: 236.1887.
1994] TILL: BROMELIADS-TYPE SPECIMENS IN VIENNA, AUSTRIA 103
"651. Tillandsia. Obragilla, Mathews. as a syno.nym o.f Tillandsia tenuifolia L. var.
Ho.o.k 837", ISOTYPE: W, F pho.to. 29981. tenuifolia.
Tillandsia II, Beer; Till. humilis Pres1, det. Tillandsia pulchra Ho.o.ker var. vaginata Wawra,
Carl Mez. Oesterr. Bo.t. Z. 30: 224. 1880.
NOTE: Currently treated as a syno.nym o.f "Reise d. Prinz. Aug. u. Ferdin. v. S.
Tillandsia humilis PresL CQburg nach Brasilien. Nro. 212. Tillandsia
Tillandsia mollis H. Hro.madnik & W. Till, PI. pulchra Ho.o.k. var. vaginata Wwr. Juiz de
Syst. Evo.L 142: 123, Abb. 1-6. 1983. FQra (Catinga's). Dr. H. Wawra.", ISO
"Tillandsia mollis H. Hro.madnik & W. TYPES: W. - Tillandsia pulchella HQo.k. var.
Till spec. no.v. Bro.meliaceae. Bo.livia, De pitiphylla Mart (spec.). Carl Mez 111193.; Til
partamento. Tarija, near the pass "Cumbre landsia tenuifolia L. var. vaginata (Wawra)
del co.ndo.r", between Tarija and Narvaez, L.B. Smith, rev. W. Till 1979.
ca. 25 km west o.f Narvaez, 2700 m.s.m., NOTE: Two. sheets bear three and fo.ur
leg. H. Hromadnik (HR 9088), On perpen plants respectively, nearly all in full blQQm.
dicular ro.cks in so.uthern expo.sitio.n with No. lecto.type has been chQsen. Currently
T.capillaris R. & P. fo.rma virescens (R. & named Tillandsia tenuifolia L. var. vaginata
P.) L.B. Smith, T.usneoides (L.) L., Abro (Wawra) L.B. Smith.
meitiella sp. cf nov., Cleistocactus straussii Tillandsia rectangula Baker, J. Bo.t. 16:238. 1878.
(Heese) Backbg. and Aylostera spinosissima "Flo.ra Argentina No. 127. Tillandsia fec
(Backbg.)Backbg.", ISOTYPES: W. tangula Bak. Estancia Germania propeCo.r
No.te: The sheet bears three post-flo.wering dQba. Junio-Decembre 1874. Dr. p.G. Lor
plants. entz", ISOLECTOTYPES: W.
TiIlandsia myosura Grisebach ex Baker, J. Bo.t. NOTE: The lectQtype has been cho.sen by
16: 240. 1878. Till (1984) and is kept in K. The sheet in W
"Flo.ra Argentina No. 122. Tillandsia bears three clusters o.fthe Lo.rentz collection.
myosura Gris. Estancia Germania prope Tillandsia recurvata (L.) L. var. ciliata E. Mo.rren
Co.rdo.ba. Junio-Decembre 1874. Dr. P.G. ex Mez in Martius, Fl. bras. 3(3):610. 1.894.
Lorentz", ISOLECTOTYPES: W. "Tillandsia ciliata Link, [3/4.], Valle de
NOTE: The lecto.type has been cho.sen by Mexico., leg. Schmitz. Mexico.", SYN
Till (1984) and is kept in BM. The sheet in TYPES: W; "367. Tillandsia unijlora.[env:
W bears two. clusters o.f the Lo.rentz .co.llec de] Mexico., [Juni], [lJerlandier)", SYN
tio.n. TYPES: W.
Tillandsia pohliana Mez in Martius, Fl. bras. NOTE: The mere citatiQn alo.ne Qf Ber
3(3): 597, pI. Ill. 1894. landier 367 as the type (Smith & DQwns,
"3658. San- Miguel. Pohl.", LECTO 1977) canno.t be considered as a lectQtypi.
TYPE (Smith & Do.wns, 1977): W.-Til ficatiQn as nQne o.fthe at least two. speciJ;nens
landsia Pohliana Mez n.sp.,det. Carl Mez o.f the herbaria P and US has been desig
III/93; Till. meridionalis Bak., Mez. nated as the lecto.type. OfbQth ·o.fthe syntype
NOTE: The inflo.rescence is damaged and co.llectiQns three sheets are kept in W. Cur
no.w co.nsists o.nly o.f two. basal flo.ral bracts rently no.t separated frQm T. recurvata.
and o.ne calyx. Tillandsia retorta Grisebach ex Baker, J. Bo.t. 16:
Tillandsia pulchella Ho.o.ker var. surinamensis 238. 1878.
Mique1 ex Mez in Martius, Fl. bras. 3(3): "FlQra Argentina No. 125. Tillandsia re
603. 1894. FIGURE 5. torta Gris. Estancia Germania prope Co.r
"Nro. 592.c. Tillantisia surinamensis Miq. dQba. Junio-Decembre 1874. Dr. P.G. Lor-
tpse. In Surinam cQllegit Dr. Hostmann. Ed. entz", ISOLECTOTYPES: W. .
~.F. HQhenacker 1846.", HOLOTYPE: NOTE: The lecto.type has been chQsen by
. - Tillandsia II, Beer; Tillandsia pulchel Till (1984) and is kept in HM. The sheet in
a HQQk. var. Surinamensis Miq. (spec.), Carl W bears two. clusters Qfthe Lorentzco.llec
ez 111193.; T. tenuifolia L. var. surina tiQn.
ensis (Mez) L.B. Smith, Iso.typus!, rev. W. TillandsiasoratensisBaker, J. BQt. 25: 235.1887.
ill 1979. "G. Mandon.-Plantae Andium BQli
NOTE: Smith and Do.wns (1977) cite the viensium. No. 1185. - Tillandsia. Hab. PrQ
erbarium U fo.r the type. In the pro.to.lQgue, va. Larecaja.-ViciniisSo.rata; colle Catar
owever, U is not mentioned. Acco.rding to. guata; super arbo.res. Alt. Reg. tempa. 2600
o.uda (1987), no. specimen is present in U m. Decembr. 1858.", ISOTYPE: W.
ut o.ne in P. The specimen in W is hence NOTE: Currently treated as a synQnym Qf
Qnsidered as the hQIQtype as it was seen, Tillandsia streptocarpa Baker var. strepto
evised, and cited by Mez. Currently treated carpa.