Table Of ContentTHE
SOUTHWESTERN
NATURALIST
ANTHONY A. ECHELLE, Managing Editor
Davip J. HAFNER, Associate Editor for Mammals
ELMER J. Finck, Associate Editor for Birds
STANLEY F. Fox, Associate Editor for Herps
Rosert J. Epwarps, Associate Editor for Fishes
KENNETH W. Stewart, Associate Editor for Invertebrates
FreD E. Smeins, Associate Editor for Botany (Ecology)
RAHMONA A. THOMPSON, Associate Editor for Botany (Taxonomy)
STANLEY F. Fox, Associate Editor for Spanish Manuscripts
VoLuME 39
PUBLISHED QUARTERLY BY
THE SOUTHWESTERN ASSOCIATION OF NATURALISTS
SOUTHWESTERN ASSOCIATION OF NATURALISTS
President President-Elect
MICHAEL L. KENNEDY WILLIAM J. MATTHEWS
DEPT. OF BIOLOGY BIOLOGICAL STATION AND DEPT. OF ZOOLOGY
MEMPHIS STATE UNIV. UNIV. OF OKLAHOMA
MEMPHIS, TN 38152 KINGSTON, OK 73439
Secretary Treasurer
LAURA L. JANECEK PAULA WILLIAMSON
SAVANNAH RIVER EcoLOGY LABORATORY DEPT. OF BIOLOGY
DRAWER E SOUTHWEST TEXAS STATE UNIV.
AIKEN, SC 29802 San Marcos, TX 78666
EDITORIAL STAFF
Managing Editor
ANTHONY A. ECHELLE
DEPT. OF ZOOLOGY
OKLAHOMA STATE UNIV.
STILLWATER, OK 74078
Associate Editor, Mammals Associate Editor, Invertebrates
Davip J. HAFNER KENNETH W. STEWART
NEw MEXICO MUSEUM OF NATURAL HISTORY DEPT. OF BIOL. Sci.
1801 MOUNTAIN Rp., NW UNIV. OF NORTH TEXAS
ALBUQUERQUE, NM 87104-1375 DENTON, TX 76203
Associate Editor, Birds Associate Editor, Botany
ELMER J. Finck FRED E. SMEINS (Plant Ecology)
DEPT. OF BIOLOGICAL SCIENCES DEPT. OF RANGELAND ECOLOGY AND
EMPORIA STATE UNIVERSITY MANAGEMENT
EMPORIA, KS 66801 Texas A&M UNIV.
COLLEGE STATION, TX 77843
Associate Editor, Herps Associate Editor, Botany
STANLEY F. Fox RAHMONA A. THOMPSON (Plant Taxonomy)
DEPT. OF ZOOLOGY DEPT. OF BIOLOGY
OKLAHOMA STATE UNIV. East CENTRAL STATE UNIV.
STILLWATER, OK 74078 ADA, OK 74078
Associate Editor, Fish Associate Editor, Spanish
ROBERT J. EDWARDS STANLEY F. Fox
DEPT. OF BIOLOGY DEPT. OF ZOOLOGY
Univ. OF TEXAS-PAN AMERICAN OKLAHOMA STATE UNIV.
EDINBURG, TX 78539 STILLWATER, OK 74078
BOARD OF GOVERNORS
Class of 1995
Troy L. BesT EDIE MARSH
AUBURN UNIVERSITY ANGELO STATE UNIVERSITY
ROBERT C. DOWLER LYNN W. ROBBINS
ANGELO STATE UNIVERSITY SOUTHWEST MISSOURI STATE UNIVERSITY
Class of 1996
Davip R. Epps KAREN MCBEE
EMPORIA STATE UNIVERSITY OKLAHOMA STATE UNIVERSITY
SARAH B. GEORGE ROBERT D. OWEN
UNIVERSITY OF UTAH TEXAS TECH UNIVERSITY
Class of 1997
WILLIAM H. BALTOSSER DwIGHT W. MOORE
UNIV. OF ARKANSAS AT LITTLE ROCK EMPORIA STATE UNIV.
ALICE F. ECHELLE CARYN C. VAUGHN
OKLAHOMA STATE UNIV. UNIVERSITY OF OKLAHOMA
The Southwestern Association of Naturalists was founded in May 1953 to promote the field study of plants
and animals (living and fossil) in the southwestern United States, Mexico, and Central America and to aid in
the scientific activities of its members. The Association holds an annual meeting with field trips and publishes
THE SOUTHWESTERN NATURALIST. Membership is open to all persons interested in natural history.
THE SOUTHWESTERN NATURALIST (ISSN 0038-4909) is published quarterly by The Southwestern
Association of Naturalists, Department of Biology, Southwest Texas State University, 601 University Drive,
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CONTENTS OF VOLUME 39
NUMBER 1
Mammalian species composition, diversity, and succession in conservation reserve program
grasslands. Dianne L. Hall and Michael R. Willig
Distribution of the threatened leopard darter, Percina pantherina (Osteichthyes: Percidae).
Alexander V. Zale, Stuart C. Leon, Matthew Lechner, O. Eugene Maughan, Mark T. Ferguson,
Steve O”Donnell, Brenda James, and Paul W. James
Aposematic coloration in Dactylotum variegatum (Orthoptera: Acrididae): support from verte-
brate feeding trials. Paul R. Neal, Mark R. Stromberg, and Karen A. Jepson-Innes
Are cicadas (Diceroprocta apache) both a “keystone” and a “critical-link” species in lower
Colorado River riparian communities? Douglas C. Andersen
Clonal diversity patterns in marginal populations of the geographically restricted shrub Neviusia
alabamensis (Rosaceae). Kenneth J. Freiley
The type locality of Sceloporus torquatus minor Cope, 1885 (Sauria: Iguanidae). Robert G.
Webb and Ralph W. Axtell
Microscopic eggshell characteristics in the collared lizard, Crotaphytus collaris (Sauria: Cro-
taphytidae). Stanley E. Trauth, Chris T. McAllister, and Wei Chen
Recolonization of wildlife on a coal strip-mine in northwestern New Mexico. Terry T. Ireland,
Gale L. Wolters, and Sanford D. Schemnitz
Utilization of coarse woody debris by Ephemeroptera in three Ozark streams of Arkansas.
Edward C. Phillips and Raj V. Kilambi
Biochemical variation in Alpheus (Decapoda, Caridea, Alpheidae) from the coast of Texas:
evidence for cryptic species. Matthew R. McClure and Ira F. Greenbaum
Trends in California mountain lion populations. K. Shawn Smallwood
Components of northern bobwhite brood habitat in southern Texas. /. Scott Taylor and Fred
S. Guthery
Factors affecting loss in mass of creosotebush leaf-litter on the soil surface in the northern
Chihuahuan Desert. W. P. MacKay, S. J. Loring, J. C. Zak, S. I. Silva, F. M. Fisher, and
W. G. Whitford
New psocids (Insecta: Psocoptera) from Socorro Island, Revillagigedo Archipelago, Mexico.
Alfonso Neri Garcia Aldrete
Notes
Characteristics of Ponderosa pines and Abert squirrel herbivory. W. Sylvester Allred and
Wiiliam S. Gaud
Food-item use by coyotes at the Naval Petroleum Reserves in California. Brian L. Cypher,
Kenneth A. Spencer, and Jerry H. Scrivner
Movements and nesting habitat of lesser prairie-chicken hens in Colorado. Kenneth M.
Giesen
Current status of the rice rat, Oryzomys couesi peninsularis. Sergio Ticul Alvarez-Castañeda ...
Announcements
NUMBER 2
Distribution and abundance of the Mexican lampreys (Petromyzontidae: Lampetra: subgenus
Tetrapleurodon). John Lyons, Philip A. Cochran, Oscar J. Polaco, and Evodio Merino-
Nambo
Changes in competition between a C, grass and a woody legume with differential herbivory.
O. W. Van Auken
Ecophysiology of three C, perennial grasses in the northern Chihuahuan Desert. R. S. Senock,
D. L. Devine, W. B. Sisson, and G. B. Donart
iii
Species diversity, selectivity, and habitat associations of small mammals from coastal California.
Gary M. Fellers
An assessment of riparian environmental quality by using butterflies and disturbance suscep-
tibility scores. S. Mark Nelson and Douglas C. Andersen
Effects of diet composition and ambient temperature on food choice of captive mallards. C.
Brad Dabbert and Thomas E. Martin
Geographic variation in morphology of spikedace, Meda fulgida, in Arizona and New Mexico.
Allison A. Anderson and Dean A. Hendrickson
Reproductive characteristics of Merriam’s pocket gopher (Pappogeomys merriami merriami)
from Huitzilac, Morelos, Mexico (Rodentia: Geomyidae). Beatriz Villa-C. and Richard
M. Engeman
Ecological conditions and the notonectid-fairy shrimp interaction. B. D. Woodward and J.
Kiesecker
Correlations between fecal nitrogen and diet composition in free-ranging black-tailed deer.
Brian N. Massey, Floyd W. Weckerly, Charles E. Vaughn, and Dale R. McCullough
Notes
List of reptiles of Morelos, Mexico, and their distribution in relation to vegetation types. Rubén
Castro-Franco and Maria Guadalupe Bustos Zagal
Observations on long-nosed bats (Leptonycteris) in New Mexico. Reginald A. Hoyt, J. Scott
Altenbach, and David J. Hafner
Natural regeneration in encinal woodlands of southeastern Arizona. Simone Borelli, Peter F.
Ffolliott, and Gerald J. Gottfried
Current status of terrestrial birds on the islands of Mono Lake, California. Linnea S. Hall,
Michael L. Morrison, and William M. Block
Latitudinal differences in sex ratios among nonbreeding western sandpipers in Puerto Rico
and Ecuador. Brian A. Harrington and Ben Haase
Timing of remigial molt in fulvous whistling ducks nesting in Louisiana. William L. Hohman
and David M. Richard
Growth and survivorship of Moapa dace (Osteichthyes: Cyprinidae: Moapa coriacea) in an
isolated stream reach on Moapa National Wildlife Refuge. G. Gary Scoppettone and
Howard L. Burge
Investigator visitation and predation rates on bird nests in burned and unburned tallgrass
prairie in Oklahoma: an experimental study. Paul Hendricks and Dan L. Reinking
Suppression of saguaro cactus flower-bud formation by roosting vultures in Arizona. Peter S.
Bennett and Michael R. Kunzmann
Helminths from mallard ducks, Anas platyrhynchos, wintering in the post-oak savannah of
southcentral Texas. Norman O. Dronen, James R. Lindsey, Lisa M. Ross, and George M.
Predation by dipteran larvae on fairy shrimp (Crustacea: Anostraca) in Utah rock-pools. Tim
B. Graham
Effects of predator exclusion on rodent abundance in an Arizona semidesert grassland. Carl
E. Bock and Jane H. Bock
Noteworthy record of the western small-footed Myotis (Mammalia: Chiroptera: Myotis cilto-
labrum). Ricardo López-Wilchis, Gerardo López-Ortega, and Robert D. Owen
Longevity of Illinois bundle flower (Desmanthus illinoensis) seeds. Ralph W. Kelting
Announcements
NUMBER 3
Seasonal variation in habitat use by the Neosho Madtom (Teleostei: Ictaluridae: Noturus
placidus). Linda Fuselier and David Edds
Distribution and status of the Chihuahua chub (Teleostei: Cyprinidae: Gila nigrescens), with
notes on its ecology and associated species. David L. Propst and Jerome A. Stefferud ...
Hybrid Cnemidophorus (Sauria: Teiidae) in Ninemile Valley of the Purgatoire River, Colorado.
James M. Walker, Harry L. Taylor, and James E. Cordes
Molecular systematics of the genus Pimephales (Teleostei: Cyprinidae). Timothy R. Schanid,
Thomas E. Dowling, and John R. Gold
Summer emergence of mayflies, stoneflies, and caddisflies from a Colorado mountain stream.
R. Edward DeWalt, Kenneth W. Stewart, Stephen R. Moulton II, and James H. Kennedy
Demography and status of the island fox and the island spotted skunk on Santa Cruz Island,
California. Kevin Crooks
Spatial, seasonal, and diel variation in abundance of the bay anchovy, Anchoa mitchilli (Teleostei:
Engraulidae), in a tropical coastal lagoon of Mexico. Manuel Castillo-Rivera, Griselda
Moreno, and Rodolfo Iniestra
Fawning habitat of desert mule deer. Kevin B. Fox and Paul R. Krausman
Systematics and zoogeography of the Mer Rouge pocket gopher (Geomys breviceps breviceps)
based on cytochrome-b sequences. James W. Demastes
Notes
Two small mammals new to the fauna of El Salvador. Mark D. Engstrom, Burton K. Lim,
and Fiona A. Reid
Chromosomal variation in Mexican populations of the Genus Dendroctonus (Coleoptera: Sco-
lytidae). Yolanda Salinas-Moreno, Gerardo Zuniga-Bermudez, and Ramon Cisneros-
Barrios
Great horned owl food habits at Mono Lake, California. Paul A. Aigner, Michael L. Morrison,
Linnea S. Hall, and William M. Block
A problematic specimen of Peromyscus with an unusual karyotype. Steven A. Smith, Ira F.
Greenbaum, and David J. Schmidly
The effect of box-trap design on rodent captures. Eric E. Jorgensen, Stephen Demarais, and
William R. Whitworth
Color preference of the island endemic lizard Uta palmeri in relation to rat eradication cam-
paigns. Bernie R. Tershy and Dawn Breese
Diurnal aggregation of banded geckos under field conditions. Russell L. Burke
Observations on mountain plover (Charadrius montanus) breeding in Utah. Keith S. Day ...
Morphometric comparison of pupfish populations, Cyprinodon nevadensis, at Shoshone and
Tecopa, California. Frances R. Taylor and John W. Pedretti
Natural cactus spine injury in the tree lizard, Urosaurus ornatus (Phrynosomatidae). Stephen
R. Goldberg, H. J. Holshuh, George A. Middendorf III, and Naida Zucker
Mountain lion predation on pronghorn in central Arizona. Richard A. Ockenfels
Announcements
NUMBER 4
Development of the tail autotomy adaptation in lizards under disparate levels of predation at
high and low elevations in Mexico. Stanley F. Fox, Susana Perea-Fox, and Rubén Castro
Population declines of the native frog, Rana muscosa, in Sequoia and Kings Canyon National
Parks, California. David F. Bradford, David M. Graber, and Farinaz Tabatabai
Geographic variation in sexual dimorphism in the collared lizard, Crotaphytus collaris (Sauria:
Crotaphytidae). /. Kelly McCoy, Stanley F. Fox, and Troy A. Baird
Pollination biology of Abronia macrocarpa (Nyctaginaceae), an endangered Texas species. Paula
S. Williamson, Lisa Muliani, and Gena K. Janssen
Long-term grazing influences on Chihuahuan Desert rangeland. Jerry L. Holechek, Ackim
Tembo, Alipayou Daniel, Michael J. Fusco, and Manuel Cardenas
Vv
Effects of Abert Squirrel herbivory on foliage and nitrogen losses in Ponderosa pine.
Sylvester Allred and William S. Gaud
Den-site selection in the island spotted skunk of Santa Cruz Island, California. Kevin R.
Double captures of small mammals in single-capture traps: random encounters or social trav-
eling? James F. Taulman, Ronald E. Thill, and James H. Williamson
Pocket gopher herbivory and mortality of ocotillo on stream terrace, bajada, and hillside sites
in the Colorado Desert, southern California. George W. Cox and Jodee Hunt
Predation by odonate nymphs on larval razorback suckers (Xyrauchen texanus) under laboratory
conditions. Michael J. Horn, Pau! C. Marsh, Gordon Mueller, and Tom Burke
Notes
Undescribed subspecies of red-tailed hawk from Baja California. Robert W. Dickerman
Observations of nest construction and bathing behaviors in the nine-banded armadillo, Dasypus
novemcinctus. James F. Taulman
Faunal use of nest boxes in the Ouachita Mountains of central Arkansas. Paul T. Caster, Gary
A. Heidt, and Karen D. Stone
The effect of bot fly (Cuterebra) infestation on cold-night trap mortality in cactus mice (Pero-
myscus eremicus). Lyle B. Nichols
Evidence of a symbiotic relationship between cellulolytic bacteria and a freshwater herbivorous
turtle. R. Brent Thomas, Don Moll, and Jack Steiert
Collection of Age-0 razorback suckers (Xyrauchen texanus) in the lower Green River, Utah.
F. Brandt Gutermuth, Leo D. Lentsch, and Kevin R. Bestgen
Effects of temperature on escape behavior in the cribellate spider, Oecobius annulipes (Araneae,
Oecobiidae). Vincent A. Cobb
Winter specimen of the common loon (Gavia immer) from the state of Hidalgo, Mexico. Patricia
Ramírez B., Atahualpa E. Desucre-Medrano, Adolfo G. Navarro S., Pedro J. Romo O., and
Hugo J. Castro
Announcements
Index to Volume 39
E
