The Pteridophytes of Mexico

1064 Pages·2004·123.93 MB·English

by  John T. Mickel, Alan R. Smith

The
PTER IDOPHY TES
of
ME X ICO
Part I (Descriptions and Maps)
John T. Mickel
and Alan R. Smith
The
PTERIDOPHYTES
of
MEXICO
Memoirs of The New York Botanical Garden
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PatriciaK.Holmgren,DirectorEmeritaofthe
WilliamandLyndaSteereHerbarium
TheNewYorkBotanicalGarden
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TheNewYorkBotanicalGarden
ScottA.Mori,NathanielLordBrittonCuratorofBotany
TheNewYorkBotanicalGarden
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TheNewYorkBotanicalGarden
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email:[email protected]
ThomasF.Daniel
DepartmentofBotany
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The
PTER IDOPHY TES
of
MEX ICO
Part I (Descriptions and Maps)
John T. Mickel
and Alan R. Smith
Memoirs of
The New York Botanical Garden
Volume 88
The New York Botanical Garden
2004
(cid:2)Copyright2004byTheNewYorkBotanicalGarden
PartIandPartIIwereoriginallyissuedasasinglevolumeonJune30,2004.
ThisversionisprintedondemandbyTheNewYorkBotanicalGardenPress.
Allrightsreserved.Noportionofthisbookmaybereproducedinanyformor
byanymeans,includingelectronicstorageandretrievalsystems,
exceptbyexplicit,priorwrittenpermissionofthepublisherexceptfor
briefpassagesexcerptedforrevieworcriticalpurposes.
CompositionbyBinghamtonValleyComposition
InternationalStandardSerialNumber0077-8931
Thepaperusedinthispublicationmeets
therequirementsofANSI/NISOZ39.48–1992(R1997). TM
LibraryofCongressCataloging-in-PublicationData
Mickel,John.
ThepteridophytesofMexico/JohnT.MickelandAlanR.Smith.
p.cm. — (MemoirsoftheNewYorkBotanicalGarden;v.88)
Includesbibliographicalreferences(p.).
ISBN10(partI):0-89327-488-7(pbk.)
ISBN13(partI):978-0-89327-488-7(pbk.)
ISBN10(originalhardcover):0-89327-458-5
1. Pteridophyta—Mexico—Classification. 2.Pteridophyta—
Mexico—Identification. I. Smith,AlanReid,1943– II.
Title. III. Series.
QK1.N525no.88a
[QK525.85]
580s—dc22
[587/.09
TheNewYorkBotanicalGardenPress
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WhenRogersMcVaughinvitedtheseniorauthortoassisthimonathree-monthcollectingexpedition
in western Mexico in 1957, this new graduate student was introduced to the ferns of Mexico and
conceived the project of eventually treating the entire fern flora of the republic. Since this initial
visit, McVaugh has been a steadfast supporter of the project, especially in connection with the
treatmentforNuevaGalicia(Mickel,1992).Inadditiontobeingquiteknowledgeableaboutferns
himself,hehasbeentirelessinthecollectionandidentificationofMexicanpteridophytes.Onthat
same 1957 trip, the senior author and McVaugh became acquaintedwithJerzyRzedowski,whose
interest and assistance in our collecting and herbarium studies, as well as his abundant fern col-
lections, have been invaluable in the completion of this flora. It is with great appreciation and
warmaffectionthatwededicatethisbooktothesetwogiantsofMexicanbotanyandfloristics.
C O N T E N T S
art
P I
xi Contributors
xiii Acknowledgments
1 Abstract
1 Introduction
4 ConspectusofthePteridophyteGenerainMexico
6 KeytotheGeneraofMexicanPteridophytes
17 Acrostichum 242 Danaea 379 Macrothelypteris
18 Actinostachys 244 Dennstaedtia 380 Marattia
19 Adenoderris 249 Dicksonia 381 Marsilea
20 Adiantopsis 250 Dicranopteris 386 Megalastrum
21 Adiantum 251 Didymochlaena 389 Melpomene
45 Alsophila 251 Diplazium 395 Metaxya
47 Ananthacorus 268 Diplopterygium 396 Microgramma
48 Anemia 269 Doryopteris 400 Micropolypodium
60 Anetium 270 Dryopteris 402 Mildella
61 Anogramma
63 Arachniodes 282 Elaphoglossum 403 Nephrolepis
64 Argyrochosma 315 Enterosora 410 Neurodium
71 Aspidotis 317 Equisetum 410 Niphidium
72 Asplenium 320 Eriosorus 411 Notholaena
132 Astrolepis
135 Athyrium 321 Gleichenella 425 Odontosoria
139 Azolla 427 Oleandra
322 Hecistopteris 427 Olfersia
141 Blechnum 323 Hemidictyum 428 Onocleopsis
153 Bolbitis 324 Hemionanthes 429 Ophioglossum
158 Bommeria 325 Hemionitis 433 Osmunda
161 Botrychium 327 Histiopteris
328 Holodictyum 435 Pecluma
165 Campyloneurum 329 Huperzia 443 Pellaea
172 Ceradenia 337 Hymenophyllum 453 Pentagramma
174 Ceratopteris 348 Hypolepis 455 Phanerophlebia
176 Cheilanthes 460 Phlebodium
213 Cheiloplecton 354 Isoe¨tes 462 Pilularia
215 Cibotium 463 Pityrogramma
216 Cnemidaria 357 Jamesonia 466 Plagiogyria
217 Cochlidium 467 Pleopeltis
219 Ctenitis 358 Lastreopsis 475 Polybotrya
233 Culcita 360 Lellingeria 477 Polypodium
234 Cyathea 364 Lindsaea 511 Polystichum
239 Cyclopeltis 368 Llavea 525 Polytaenium
240 Cystopteris 368 Lomariopsis 528 Psilotum
370 Lonchitis 529 Pteridium
371 Lophosoria 533 Pteris
372 Loxogramme
373 Loxoscaphe 544 Radiovittaria
374 Lycopodiella
375 Lycopodium
377 Lygodium
CONTENTS vii
544 Saccoloma 609 Tectaria 683 Vittaria
546 Salvinia 615 Terpsichore
547 Schaffneria 621 Thelypteris 686 Woodsia
548 Schizaea 623 subg.Amauropelta 692 Woodwardia
549 Scoliosorus 638 subg.Cyclosorus
550 Selaginella 648 subg.Goniopteris 697 Literature
603 Sphaeropteris 663 subg.Meniscium IndextoScientificNames
603 Sphenomeris 667 subg.Stegnogramma
604 Sticherus 668 subg.Steiropteris
607 Stigmatopteris 668 subg.Thelypteris
669 Trichomanes
art
P II
703 Plates
IndextoScientificNames
C O N T R I B U T O R S
Thefollowingpersonsareattributedauthorshipforallorpartofselectedgenerictreatments.
R.JAMESHICKEY(Iso¨etes),BotanyDepartment,MiamiUniversity,Oxford,OH45056
W.CARLTAYLOR(Iso¨etes),MilwaukeePublicMuseum,Milwaukee,WI53233
IVA´NA.VALDESPINOQ.(Selaginella),Apartado10912,UniversidaddePanama´/ANAM,Panama´,Panama
MICHAELD.WINDHAM(Woodsia),GarrettHerbarium,UtahMuseumofNaturalHistory,UniversityofUtah,SaltLakeCity,
UT84112
A C K N O W L E D G M E N T S
Wearegratefultothecuratorsandadministratorswhomade Severalbotanistsgraciouslyfurnishedvaluabletaxonomicas-
their specimensavailable forstudy,bothon loanandon visita- sistance:JudyGarrisonHanks(Lygodium),RichardHauke(Eq-
tion. Herbaria consulted include ANSM, ARIZ, ASU, B, BM, uisetum), Hugo Navarette (Dennstaedtia), Benjamin Øllgaard
BR, C, CAS, CHAPA, DS, ENCB, F, FCME, GH, GUADA, (Lycopodiums.l.),W.H.Wagner,Jr.(Botrychium,Ophioglossum),
IBUG, IEB, ISC, IZTA, K, LAM, LL, MA, MEXU, MICH, and Michael Windham (Argyrochosma, Cystopteris, Notholaena,
MO, MSC, NY, P, PH, PR, PRC, S, SD, SERBO, TEX, Woodsia).GeorgeYatskievychbroughtseveralinterestingcollec-
UAMIZ,UC,UNL,US,XAL,andYU. tions to our attention. We also had valuable discussions with
Assistance with manuscript preparation and specimen proc- Rodolfo Aguirre-Clavera´n, Mar´ıa de la Luz Arregu´ın-Sa´nchez,
essingwasprovidedbyRuthRussellandDeeFerguson.Richard Ray Cranfill, Lucrecia Garc´ıa A., Francisco Lorea-Herna´ndez,
Rauh checked many details in the manuscript. Carol Mickel Robbin Moran, Leticia Pacheco, Blanca Pe´rez-Garc´ıa, Ramo´n
helpedwithdatarecordingatNYandmanyotherherbaria.We Riba,DanielTejero-D´ıez,andErnestoVela´zquezM.
aregreatlyindebtedtotheseniorauthor’sresearchassistantLee Financialsupportfortravelandillustrationscamefrommany
Brown, who processed as well as critically read the full manu- membersoftheNewYorkChapteroftheAmericanFernSociety
script.WethankSueBainbridgeforassistanceingeneratingthe and other fern friends. Especially generous gifts were received
base maps and Joan Smith for execution of the maps. Special fromIrvingKnobloch,EleanorandArdenMoyer,andGertrude
thanksgo to WilliamBuck of the New YorkBotanicalGarden andGeorgeYourke,andbequestsfromF.GordonFoster,Elinor
forhismeticulousreviewingandeditingofthemanuscript,and Merrell, and Elizabeth and Harold Tovell. Without their assis-
toDanielTejero-D´ıezandGeorgeYatskievychfortheirreviews tance,thisprojectcouldnothavebeencompleted.
and helpful comments. Brian Viger of the NewYorkBotanical
GardenPressprovidedvaluableassistanceincomputerizationof
themanuscript.