Table Of ContentThe evolution ofsheep and goathus bandry
in centralAnatol ia
BenjaminS.ARBUCKLE
BaylorUniversity
Departmentof Anthropology,ForensicScience,andArchaeology
OneBearPlace#97173,Waco,TX76798-7173 (U.S.A.)
[email protected]
AliyeÖZTAN
AnkaraUniversity,DilveTarih-Co˘grafyaFacültesi
ArkeolojiBölümü,Ankara(Turkey)
SevilGÜLÇUR
IstanbulUniversity,EdebiyatFacültesi
PrehistoryaAnabilimDalı,Istanbul(Turkey)
ArbuckleB.S.,ÖztanA.&G ülçurS.2009. —The evolutionofsheepandgoathusbandry in
centralAnatolia. Anthropozoologica44(1):129-157.
ABSTRACT
Thispaperexplores t he evolution ofsheepandgoathusbandry in central
Anatoliathrough the analysisofnewandpublished faunaldata f rom
Neolithic, Chalcolithic, andBronzeAgesitesin theregion. Themajor
KEYWORDS patternsofchangeovertimein thepastoralsystemared iscussedincludingthe
Pastoralism,
beginningsofherding,thea ppearanceofthepractice ofyoung malekill-off,
herdmanagement,
sheep, andthetiming oftheonsetoftheintensiveuseofsecondary products.Results,
goat, including the discovery oflongstanding differencesin themanagementof
domestication,
Neolithic, sheepandgoats as well as synchronic differencesin herding strategies,have
Chalcolithic, importantimplicationsfor u nderstanding theroleof a nimal husbandry
BronzeAge,
in centralAnatoliancommunitiesandindicatethat t hehistory ofpastoral
centralAnatolia,
Turkey. productionintheregion iscomplexandmulti-faceted.
RÉSUMÉ
L’évolutiondel’élevage desmoutonsetdeschèvresenAnatolie centrale
Cetart icle aborde l’évolution de l’élevage desmoutonsetdeschèvr esen
l’Anatolie centrale àtravers l’analyse de donnéesnouv ellesetpubliées
ANTHROPOZOOLOGICA •2009•44 (1)©PublicationsScientifiquesduMuséumn ationald’Histoirenaturelle,Paris. 129
S.ArbuckleB.,ÖztanA.&Gülçur S.
provenantde sitesde larégion datantduNéolithique, duChalcolithique e t
de l’âge deBronze.Lesprincipaux modesde changementdansletempsdu
MOTSCLÉS
Pastoralisme, systèmepastoralsontexaminés, d epuislesdébuts de l’élevage, l’apparition
gestiondutroupeau, de lapratique d el’abattage d esjeunesmâle e tlemomentde l’apparition
mouton,
de l’utilisation intensive desproduits secondaires.Les r ésultats,ycompris
chèvre,
domestication, la découverte de différencesdurablesdanslegestion desmoutonsetdes
Néolithique, chèvresainsique d e différences synchroniquesdansles s tratégiesd’élevage,
Chalcolithique, ontd’import antesimplicationspour la compréhension du rôle del’élevage
âgeduBronze,
danslescommunautésd’Anatolie c entrale.Ilsmontr ent quel’histoire d ela
Anatoliecentrale,
Turquie. productionpastoraledanslarégion estcomplexeetmulti-facettes.
INTRODUCTION aparticularfocus onexaminingevidenceforsheep
andgo athus bandry in centralAnatol ia from the
Asoneofthemost importantsocioeconomicsystems Neolithicthrough theBronzeAge.Despitearapid
intheancientandmodernNearEast,pastoralism, increasein studiesde tailingthenatu reof a nimal
theherding ofdomestichoofed ungulates,whether economiesin centralAnatol ia( e.g.,Boess neck&
practicedbynomads,agro-pastoralists,within ega- Wiedemann1977,von denDriesch& Boessneck
litariantribesors tratifiedstates,hasbeenamajor 1981,Hongo1993;1998;1999;2004,Zeder&
focusofanthropologicalresearchformorethanahalf Arter1994,Buitenhuis1996; 1997; 1999,De
century(de Planhol1958,Barth1959,Irons1969, Cupere &Waelkens1998, Howell-Meurs2001;
Salzman1971;2004,Dyson-Hudson 1972,Irons Martinetal. 2002,Atıcı 2003;2005,Carr uthers
&Dyson-Hudson 1972,Bates1973,Gart hwaite 2003,DeCupere&Dur u2003,von denDriesch
1978,Tapper1979,Beck 1 986,Black-Michaud &Pöllath2004,Russell &Martin2005,Russell, et
1986,Galaty &Johnson 1990,Borgerhoff Mulder al.2005,Martin&Russell 2006)therehaveb een
&Sellen1994,Azarya1996). Archaeologistshave fewattempts tolookatthebroadpatternsofchange
beenparticularlyinterestedinaddressingthreebroad overmultiplechronological/culturalperiodsandto
questionsrelatedtopastoralismincluding:1) the synthesizeresults frommultiplesites(although for
originsofsheepandgoatherding,whichrepresents theNeolithicperiodsee Martinetal. 2002).Asa
thefirstsuccessfuldomestication ofafoodanimal result,acentralcomponentofthec ulturalhistory
(Helmer1992,Harris1996,Horw itzetal. 1999, oftheregion remainspoorlykno wn.
Petersetal. 1999,Zeder&Hesse2000,V igneet Byfocusingontheevolutionofpastoralproduction
al.2005),2)thed evelopmentofhighlyproductive, inthelonguedurée,thegoalo fthispaperist opre-
milk and fiberproducing herds(Sherratt 1981, sentabroadpictureofthee volutionofsheepand
Greenfield1989), and3)the a ppearance oflarge goathusbandry patternsin centralAnatoliaandto
scale,mobilepastoralsystemskno wnfrom histori- addressimportanttransitionsincluding theinitial
calperiodsin theregion (Barth1961,Irons1975, appearance ofpastoraleconomiesas w ell as the
Matthews1978,Beck1986,Bar-Yosef&Khazanov developmentofmoreintensivepastoralmanagement
1992, Zeder1994b, Bar-Yosef&Meadow1995, strategiesincluding theuseofherdsfors econdary,
Grigson2000,Fleming 2004,Greenfield&Fowler orantemort em,products (i.e.,milk, f iber) (after
2005). Itis w idely recognized thateach o fthese Vigne&Helmer2007).Inaddition torepresenting
transitionsrepresentsmajortransformativeevents animportantincreasein theproductivepotential
in the culturehistory oftheNearEast (Sherratt ofancientherds,thislast featureisoftenseenasa
1981;1983,Bar-Yosef &Khazanov1992). necessarypreconditionforthedevelopmentoflarge
Inthispaperwedescribethebroadpatternsofchange scale,specializedandmobileformsofpastoralism,
overtimein thiscentralsocioeconomicsystemwith thedevelopmentofwhichi salsoaddressed.
130 ANTHROPOZOOLOGICA •2009•44 (1)
Theevolutionofsheepandgoathusbandry incentralAnatolia
Thefollowingdiscussionisbased onabodyofnew BACKGROUND
andpublishedfaunaldatafromsevensitesin central
Anatoliarepresenting Neolithic,Chalcolithicand Inordertoaddress questionsconcerning pastoral
BronzeAgesettlements (Fig.1;Table1). Although productionwefocusonacombinationofageand
chronologicalcoverageisnotcomplete, thecurrent measurementdataforsheepandgoats.Aged ata,
sample doesprovide avaluablemeans t oaddress basedonthestateoff usionoftheepiphysesoflong
majorquestionsofchangeovertimein systemsof bonesandthe e ruption andwearofmandibular
caprinehusbandry inthisregion. teeth, canbe used to interpret the age composi-
FIG.1.—MapofTurkeyshowingthelocation ofsitesmentioned inthetext.A I=A ıklı;ERB =Erbaba;ÇA T=Çatalhöyük;
SUB=Suberde;K K=Kö kHöyük;GÜV =Güvercinkayası;ACE =Acemhöyük.
TABLE1.—Approximatechronologicalrelationshipsof theprimarysitesmentioned inthetextaswellasthefrequencyof caprines
andtheratioofsheeptogoatin eachassemblage (fromPerkins&Daly1968,Buitenhuis1997,Cessford2001,Thissen2002,Martin
&Russell2005,Arbuckle2006).
SiteApprox.d atecalBC %caprinesSheep:goatr atio
AıklıHöyük 8200-7500 84 4.0:1
Suberde 7400-7000 825.6:1
Çatalhöyükpre-XIItoIV 7400-6200 65-757.0:1
Erbaba 6500-6000 77 4.6:1
KökII-V6200-5500 60 3.5:1
KökI5200-4800 83 3.2:1
Güvercinkayası 5200-4800 81 4.3:1
AcemhöyükII-III 2000-1800 65 1.7:1
ANTHROPOZOOLOGICA •2009•44 (1) 131
S.ArbuckleB.,ÖztanA.&Gülçur S.
tionoftheanimalschosenfors laughter(Hesse& chosenforslaughterwithgeneralgoalsofpastoral
Wapnish1985,Davis1987,Lyman1994).Theseare production.Thelink betweenthesevariableshas
oftenpresentedintheformofsurvivorshipcurves, beendiscussed bymanyresearchers(e.g.Higham
whichvisuallyrepresentthefrequencyofanimals 1967,Ducos1968, Redding 1981;1984, Z eder
survivingincreasinglyoldagecategories,beginning 1991;2001,Helmer1992,Vigne&Helmer2007)
with100%atagec ategory1andendingwithnone but hasbeendescribed most influentiallyin the
survivingpastagecategoryX(Payne1973,Levine formofpredictivemodelsbyPayne(1973). These
1983,Lyman 1 994). Although there areseveral modelsdefinetheb asicrelationshipsbetw eenthe
methodsforgenerating survivorship curves,this goalsofherdmanagement—whetherfocused on
paperfollo ws thelife tablemethod c ommon in maximizing theproduction ofprimary (orpost-
thewildlife literature(Deevey1947,Quick1 963, mortem) products suchasmeator s econdary (or
Caughley1966,Lyman1 994)inwhicheachsur- antemortem)productssuchasmilk orfiber—a nd
vivorshipvalue(sometimesdenoted as“lx”)repre- thesexanda gecompositionoftheanimalschosen
sentsanestimateofthepercentageofindividuals forslaughter.
survivingatthebeginningofagivenagec ategory. Briefly,themeatmodelp redictsthatwhenthegoal
Inarecentpaperfocusing ontheinterpretation of ofproductionisprimarilymea t,mostyoungmales
mortality data,Vigne andHelmer(2007:20-21) willbekilledwhentheyreachanoptimumpoint
suggest that whencalculating mortality profiles, inweight-gain—between18-30months(Payne
a correction factor should be applied in order to 1973:281-82; alsoDigard1981,Redding 1981,
accountforthevaryinglengthsoftimerepresentedby Black-Michaud1986,Salzman2004). Theageat
themost commonlyusedagestages(Payne’s (1973) whichmaleswillbeslaughteredwithinthisrange
MandibularWearStagesA-I). However,sincemor- (oreven o utside ofit) m ay vary depending on a
talitydatain thispaperarepresented intheformof varietyoffactorsincludingthecost/availabilityof
survivorshipcurves,thissuggestedcorrectionfactor fodderandgraze, theavailabilityoflabortosupervise
isnot u tilized.Since eachsur vivorship (lx)value herds,theimmediate f inancial orotherneedsof
representsanestimateofthepercentageofa nimals theherder,environmentalconditions,andcultural/
survivingatthebeginningofagecategoryX,differences marketpreferencesforlambort hemeatofother
inthelengthofeachcategory (ifwithin reason)are specificdemographicgroups.Thismodelp redicts
relativelyunimportant.However,whenm ortality thatwhenmeatistheprimary goalofproduction,
values(dx v aluesin wildlife studies)representing theresulting survivorshipcurvedropsprecipitously
thenumberofd eathsin aparticularagecategory formalessometimein thefirst2.5yearswhilefemale
arepresented as the focus of analysis(as t heyare survivorshipdeclinesmuchmoregraduallythrough
in Vigne andHelmer 2 007)then i tmakesmore adulthood.Asaresult,in themeatmodelthevast
sensetoaddress thevary ing lengthsof e achage majority of animals surviving intoadulthood a re
category witha c orrection factor(alsosee Payne females(80-98%based onethnographicexamples
1973:Table3,f igures15 and16). (Bates1973:147,Redding 1981)).
Inthispaper,measur ementdataincluding those The fibermodel p redicts that whentheprimary
derivedfrombothfused andunfused specimensare goal ofproduction is wool orhair,herders shift
usedtoidentifysizediminutionassociatedwiththe theirmanagement s trategy towardsculling adult
process of domestication (Bökönyi 1969,Ducos individuals(Payne1973:282). Asaresult,this
1978,Uerpmann1979,Davis1987,Meadow1989, modelpredictsthatwhenfiberist heprimary goal
Zeder2006)and, incombinationwithsurvivorship ofproduction,themajority ofanimalswillsurvive
curves,todeterminetheproport ionsofmalesand wellintoadulthood, and,incontrasttothemeat
females slaughtered within specific agegroups model,the a dultpopulation will consist of both
(Hesse1978, Zeder&Hesse2000,Z eder2001). femalesandmales(oftencastrated)since bothare
Inordertousethesesour cesofdatatointerpretthe effectivefiberproducers.Identification ofbothof
goalsofpastoralp roduction,wefocusonmodels thesefeaturesiscriticaltotheidentification ofthe
thatlink the age andsexcomposition of animals fibermodelandf ordistinguishing fiberproduction
132 ANTHROPOZOOLOGICA •2009•44 (1)
Theevolutionofsheepandgoathusbandry incentralAnatolia
fromameatprofilein whichthejuvenileshavebeen thenon-intensiveproduction ofherdsformilk in
deletedbytaphonomicfactors. additiontomeatmaybe indistinguishablefromthe
Payne’sthirdmodeldescribesthemanagementexpec- rangeofvariation expectedwithinthemeatmodel.
tations whenthegoal i s theintensiveproduction Finally,theinterpretationofdairyproductionfrom
ofmilk. Thismodelpredictsthatwhenmilkisthe mortalityprofilesischaracterized bytheproblemof
primarygoalofproduction herderswillslaughter equifinality.Likeallarchaeologicalpatterns,mortality
mostmalesasyounglambssoastomaximizethe profilessimilartothemilk modelcouldpotentially
milkavailableforhumanconsumption. Thismodel beexplainedbyahost ofbehaviorsotherthandairy
isdistinguishablefromthemeatmodeli nitsfocus production(Halstead1998). Inparticular,produc-
ontheslaughterofmalesin theyoungest agecate- tion systemsfocused on very young tendermeat,
gories.Of all oftheproduction models therehas thepresenceofhigh infantmortality(asin casesof
beenthemost discussionconcerningtheapplication crowdedpenningandstall ing),or samplingfrom
ofthemilk modeltoarchaeologicalcontexts (see areascontaininghighproportionsofinfantremains
Halstead1998andreferencestherein).Arguments (e.g.,shrinesor t emples)couldproduce morta-
havetended tofall intothree categories. litypatternsthatmimicthemilk model.Halstead
First,althoughslaughterofsurplus animalsin the (1998:14)haspointed outthatwhiletaphonomic
first6-8weeksfollo wingbirthiscommon practice biases,particularlythoseaffectingtheremainsofthe
amongmoderndairy producersandactstomaxi- youngestindividuals,arelikelytomaskarchaeological
mizethemilk available forhumanconsumption, evidenceforintensivemilk production(e.g.,Munson
somehaveraised doubts as towhetherprimitive 2000),theyareless likelytoartificiallycreateit(also
domesticbreedscouldproduce milkintheabsence Vigne&Helmer2007:16). However,asdiscussed
of an infant(Clutton-Brock 1 981,McCormick above,convergentmortalitypatternsresultingfrom
1992,Balasse2002). IsotopicworkbyBalassea nd differentmanagement strategies( e.g.,subsistence
Tresset(2002) has suggested that thismayhave productionofmilk vsintensivemeatproduction)
beenthecaseforNeolithiccattlein Europe,which remainamajorobstacletotheinterpretation ofthe
appeartohavebeenweaned at6-9monthsrather goalsofpastoralp roduction.
thaninthefirstweeks.However,thisissueofmilk Moreover,Halstead hasalsorightlypointed out
let-down maybe aless serious issue forcaprines the fundamentalweakness in mortality evidence
(Halstead1998:5-6). fordairy ing:that w hiletheintensive c ulling of
Secondly,Payne’s milk modeldescribesasystem surpluslambsdoes“implythatherdmanagement
ofhighlyintensivemilk production focused on enhancedthepotentialforproductionofmilk rather
producingforlarge-scalemarkets andits applica- than[otherproducts]”(Halstead 1998:7,original
bility to prehistoricsocioeconomic c ontexts has emphasis),itdoes notprovethatmilk wasactually
beenquestioned.Halstead (1998) has suggested exploited.Asaresult,multipleinterpretationsare
that this typeofmanagement s ystem isprobably alwayspossibleandmilk productionremainsoneof
mostlikelyunderconditionsin whichherdersare themost difficultmanagementsystemstoidentify
highlydependentuponanimalproducts,havelarge archaeologically.
herds,andarehighlyintegrated intoamarketeco- Payne’sthreemodelsrepresenttheoreticalconstructs
nomy(i.e.,specialized pastoralists).Smallerscale, thatdescribepatternsofherdmanagementexpected
subsistence-oriented producers aremorelikely t o underoptimizing conditions.Asaresult,theyare
space theslaughterofsurplus ramsout overan best viewed asheuristic d evices,whichserv e asa
extended periodofmonthsorevenyears sacrifi- startingpointfrom whichtointerpretarchaeolo-
cing thequantity ofmilk available but providing gicalpatterns,whichthemselvesarenotexpected
a“walkinglarder”asahedgea gainstfuturepoten- torepresentoptimization (Halstead1998:4-5).As
tialinsecurities(see Redding(1981)foradetailed Payne(1973:282)n oted,herdersrarelyfocus ona
discussionofriskreduction asaherding strategy). singleproductandmust insteadbalancebetween
Thisdelayed slaughterofsurplus lambsislikelyt o conflicting requirements represented bymanage-
lookverymuchlikePayne’s meatmodelandthus mentformultipleproducts,a swellasin response
ANTHROPOZOOLOGICA •2009•44 (1) 133
S.ArbuckleB.,ÖztanA.&Gülçur S.
toahost ofsocial,political, a nd environmen- a“tendermeat”model.IncontrasttoPayne’smodel
talvariables.However,instead ofrepresenting a (termedthe“typeAmilk”model),the“t ypeBmilk”
weakness,thetheoretical n atureofthesemodels model ischaracterized by the delayedslaughterof
isalsotheirgreatest strength, asitis t hrough the lambs throughout theirfirst year similar tothe
process ofidentifying deviationsfrom optimality situationdescribedabove fornon-intensivevillage
thatwebegintointerpretthecomplexmanagement herders(Halstead1998:9,alsoPayne1973:282).
behaviorsofherders. Thismodelproducesmortalitypeaksamong older
Althoughrarelydiscussed, Payne(1973:282)p ro- lambs,from6-12months,andthenagain foradult
videsseveralexamplesoftheinterpretation ofdevia- femalesbetween2-4 years.The“tendermeat”model
tionsfrom hisoptimization modelsillustrating how predictsakill-off ofyoung ramsbetweentheagesof
multipleproduction goalsmightbe balanced.For 6-12monthsratherthanbetween18-30monthsas
example,ifbothmeatandmilk areproduced and inPayne’smeatmodel(alsosee Payne1973:282).
milkisofprimary importanceandeitherlaboror Thesemodelsarepresented tofill perceivedgaps
wintergraze/fodderisr estrictedorexpensive, then inthePaynemodelsandrepresent“ intermediate”
mostmalelambswillslaughteredbeforetheirfirst management strategies,whicharethenused by
winterat6-9months.Iftheproduction ofmeatis Vigne andHelmer t o interpretcomplex “ mix-
ofgreaterimport ance,ifwinterfeeding posesfew tures”ofdifferenttypesofproduction goalsinclu-
problems,andiflaborisavailablethensurplus males ding tendermeat + t ypeBmilk,orfiber +meat,
arelikelytobe slaughteredintheirsecondandeven based largelyon thelocation ofmodal m ortality
thirdyears.However,ifwinterfeed isexpensiveor age.Althou gh the developmentofmore d etailed
unavailableandlaborisin shortsupplythensurplus models withwhichto i nterpretmort ality data
lambsmayagain be slaughtered beforetheirfirst isbeneficial,onemust always be mindful ofthe
winter.Inthis w ay, deviationsfrom the expected inherent weaknessesin theuseofmort ality data
modelscanbe interpreted in termsofvariables to interpretmanagement s trategies.Perhaps t he
includingmixedproductiongoals,costsoffodder, mostimportantofthese—a sdiscussed earlieras
availabilityofwinterpastur e,availabilityoflabor, acriticismofPayne’s milkmodel—isequifinality ,
aswellastheinfluence ofmarkets. boththrough thelikelihoodthatmultiplestrategies
Vigne andHelmer(2007 andreferences therein) withdifferentproductiongoalscanproducesimilar
haverecentlymade an importantcontribution to mortalityprofiles(e.g.,tendermeat,typeBmilk,
theinterpretationofmanagementstrategiesbypro- mixedmeatandmilk) asherdersmakecomplexand
posingtwonewmodelsin additiontothosedefined oftenconflictingmanagementdecisionsin response
byPayne.Theseinclude a“typeBmilk ”modeland todynamicsocialande nvironmentalconditions,as
Table2.—Summary oftherelationshipbetweenmodalslaughterage andthegoalsof herdmanagement.
Modalslaughterage(months)
0-2 2-6 6-12 12-24 24-48 48+
Fiber
Dairy
TypeBmilk
Meatandmilk
Tender
Meat
meat
134 ANTHROPOZOOLOGICA •2009•44 (1)
Theevolutionofsheepandgoathusbandry incentralAnatolia
wellasthroughthealterationofmortalityprofiles SITESANDDATA
throughtaphonomicprocesses.
Thesuiteofmodelsdiscussed aboveprovidesthe Thesitesexamined inthispaperrepresentarange
frameworkforinterpreting managementgoalsfrom ofsettlement t ypesincluding large andsmall
mortalityprofileswiththefullacknowledgement Neolithic andChalcolithic a griculturalvillages
oftheirin herentshortcomings.Togethertheypro- as well asonelargeBronzeAge c enter(Fig. 1;
videseveralpointsalongacontinuumofpossible Table1). Ageandmeasur ementdataf romthese
managementstrategieseachfocused onincreasingly sitesallo wustoaddress thecharacterofsystems
aged animals(see Table2). Thesemodelsarenot of caprinehusbandry in theregion andtheir
viewedasthefinalanswerstoquestionsofpastoral changeover t ime from the earliest Neolithicto
productionbutinsteadareused assuggestedpoints theBronzeAge.
ofentryintotheprocess ofinterpreting themana-
gementdecisionsofa ncientherders. A5IKLIHÖYÜK
There aretwoadditional i ssues thatmust alsobe AıklıHöyük i salargemoundsitelocated in
addressedwheninterpretingmortalityevidencefor themountainous region ofCappadocia(Esin &
herdmanagement.Thefirstistheissueofmobility Harmankaya1999) (Fig.1).Thesiterepresents an
andthefactthatmortalitydataderivedfromone earlyphaseofthec olonizationofcentralAnatolia
sitemay r eflectan i ncompletesampleof ahus- bysedentaryfarmersintheAceramicNeolithicand
bandry system (Meadow1980). Ifmanagement datestothelateninthande arlyeighthmillennia
strategiesinclude seasonalmovementstoandfrom calBC (middlePPNBin theLevant) ( Table1)
summerpastures,forexample,thenthearchaeolo- (Thissen2002).
gicalpatternsderivedfromeithersummerorwinter Buitenhuis(1997)foundthatt heanimaleconomy
residencesiteswillrepresentatr uncatedmortality atAıklıwasdominated bycaprines(Table1).
profilereflecting onlyaport ion ofthe c omplete However,questionsremainconcerninghowcaprines
system(e.g.,Cribb 1984,Vigne&Helmer2007: wereexploitedatthissite,andwhethertheywere
22). AsVigne andHelmer(2007) p ointout this herded orhunted (Vigne etal. 1999,Mart in et
representsasignificant“trap”forarchaeologists that al. 2002). Survivorship curvesbased on epiphy-
canbe addressedonly throughregionalsampling sealfusion indicatethat t heAıklı c aprines were
ofdifferenttypesofsites( e.g.,Helmer etal. 2005). slaughteredprimarilybetweentheagesof1-3y ears,
Thesecondissueisoneofprovisioning. Particularlyin withrelativelyfe wanimalssurvivingpasttheageof
thecontextofcomplexsocieties,theprovisioning of fusionofthedistalradius (c.36 months)(Fig.2).
settlementswithmeat,oftenbyspecializedproducers, Inaddition,Buitenhuisnoted thepresence of a
canhaveasignificantimpacton theinterpretationof
mortalitydata(e.g.,Stein 1987,Wapnish&Hesse
1988;1991,Crabtr ee 1990; 1996, Zeder1991). 100
Urbansystemsareoftencharacterized bysystemsin 90 GGaannjjDDaarreehh
80
whichruralproducersprovisionurbanconsumers.
70
Stein(1987)hascontrasted “consumer”mortality (%) 60 KKööskk AAsııkkllıı
otpofroainfgieclesluascndhdea“rapardcoteedarruitzcheedro”fbmpyaaatrrtkeeelratn-tsia,vgweelhdyincaahnrarirmeeoalwxsp.Treachntuegsde survivorship 345000 SSuubbeerrddee
20
themovementofanimalsofparticulardemographic
10
groups toand f rom producerand consumer sites
0
has thepotentialtocreatemort ality profiles that 0 3 6 12 24 36
age(months)
primarily represent systemsof d istribution rather
FIG.2.–Survivorship curvesbased onepiphysealfusionfor
thanproduction(Zeder1991) andmust betaken
sheep/goatsforAıklıHöyük(n=2806)(fromBuitenhuis1997),
intoaccountwheninterpretingmortalitydata,par- KökHöyükI-V(n =473),Suberde (n=95)(from Arbuckle
2008a)and wildsheepfromGanjDareh(n=2112)(fromHesse
ticularlyin (butnotlimited to)complexsocieties.
1978).
ANTHROPOZOOLOGICA •2009•44 (1) 135
S.ArbuckleB.,ÖztanA.&Gülçur S.
significantnumberofneonatalcaprineremains inconcentrationsoflargemales(see Fig.3:Ganj
in the assemblage.The f ocus on culling animals Dareh) (Helmer1988, Helmer etal. 2005,Rus-
withinanarrowagerangeiscommonlyassociated sell&Martin2005).Thissuggeststhatsheepand
withhumanmanagement,whilehunting often(but goats werelikely s ubject todifferentexploitation
certainlynotalways)results inhigherproportions strategiesatAıklı.
of“prime-aged ”adultindividuals(Stiner1990) Recentanalysisof c arbon andnitr ogen isotope
(comparecurvesforKö4kandGanj DarehinFig.2). valuesfrom Aıklı caprineremainshasindicated
Moreover,theremainsofneonatalanimalsare thatbothsheepandgoats werecharacterizedbya
notuncommonatNeolithicandlaterperiodsites homogeneousand“restricted”diet(Pearson etal.
characterizedbyherding economies,buttheycan 2007:2178),whichwasinterpreted aspossibly
alsobe anindicatorofspring hunting(e.g.,Davis reflectinganearlystagein themanagementofthe
&Fischer1990). Together,thesemort ality data caprinepopulations.However,thelimited variation
havebeenused tosuggest thattheAıklıcaprines in CandNvaluesin theAıklı c aprinesisiden-
were“appropriated”resources(Vigne&Buitenhuis tified as “unusual” whencompared tothewider
1999:58, Martinetal. 2002)andweresubjectt o spreadofvaluescharacteristicoflaterNeolithicsites
anearlyformo fherdmanagementt hatBuitenhuis includingÇatalhöyük(Pearsonetal. 2007:2178),
(1997)referredtoas“proto-domestication”. whileasimilarpatterno flimited variationhasbeen
Moreover,Buitenhuishasargued that theAıklı identifiedinancientwildungulatepopulationsin
caprinesrepresentamorphologicallyw ildpopula- thesouthernLevant(Makarewicz2 007;personal
tion,i.e.,theyexhibitno evidenceforadecreasein communication 2008).Thus isotope data add to
sizeorothermorphologicalchangesoftenassociated thepicturethatcaprinee xploitationatAıklıwas
withtheprocess ofdomestication,throughout the significantlydifferentfrom laterNeolithicmana-
occupationalsequence ofthesite, aperiodspan- gementstrategiesbut donotclarifythenatureof
ningc.400years.Although morphologicalchanges thosestrategies,whichm ayhavebeenmoresimilar
arenotexpected tocharacterizethee arlieststages towildlife managementthanintensivehusbandry.
ofhumanmanagementoveranimalspopulations
(Zeder&Hesse2000, Z eder 2006),thelack o f SUBERDE
evidence for the developmentofmorphological ThesiteofSub erdeislocatedinaninter-montaine
changesoverthisextended periodoftimeisinte- basinintheBeysehir-Suglaregionofsouthwestern
restingandsuggests thatanimalsmaynothaveb een centralAnatolia(Fig. 1). Suberde represents the
underintensivehumanm anagementand/orwere remainsof asmall villagesettlementofthelatest
notreproductivelyisolated fromlocalwildcaprine AceramicNeolithic a ndis t he earliest excavated
populations(Arbuckle2005). Neolithicsettlementin theBeyehirregion.Radio-
Inaddition,Buitenhuis(1997:659) reports that carbondatesindicatethattheNeolithicoccupation
thesexratioforsheepatAıklıiss lightlybiased in ofSuberde spannedthesecondhalfofthee ighth
favoroffemales,whileforgoats itisskewedtowards millenniumcalBC (Bordaz1965;1966;1969;1973,
males.Theseresults areevidentin thedistribution Bordaz&Alper-Bordaz1977,Arbuckle2008a).
ofastragalusmeasurementspresentedinFigures3 Although originallyinterpreted as representing a
and4. Forsheep,thesemeasur ementsareskewed “Neolithichunters’village”(Perkins&Daly1968),
slightly towards theleftindicating anabundance recent reanalysisoftheSuberde assemblagehas
ofsmallerindividuals(females),whilethegraph shownthatbothsur vivorshipcurvesandmetrical
forgoats isskewedtowardstherightindicating a datasuggest that sheepandpossiblygoats were
concentrationoflargermales.Thesetwop atterns underhuman m anagementat t his site(Arbuckle
areoftenassociated withdifferentexploitation sys- 2008a).SurvivorshipcurvesfortheSuberdecaprines
temswithherdingpracticesoften(butnotalways) indicatethat t hevast majority of animals were
producingsexratiosskewedtowardssmallerfemales selectedforslaughterbetweentheagesof1-3y ears,
(seeFigs3:Çatalhöyük,Kö4k;4:Ganj Dareh),and withdentalweardataindicating apeakbetw een
huntingstrategiesoften(butnotalways)resulting 12-24monthsaspredi ctedbymodelsofmea tor
136 ANTHROPOZOOLOGICA •2009•44 (1)
Theevolutionofsheepandgoathusbandry incentralAnatolia
Fig.3.—Greatest lengthofthe astragalus(GLl)forsheepfromGanjDareh(n=34) (fromHesse1978),AıklıHöyük(n=470),
Suberde(n=21),Çatalhöyükpre-XIItoIV (n=56)(fromRussell&Martin2005),Erbaba(n=95)andKökI-V(n =137).Triangles
representmeanvalues.
ANTHROPOZOOLOGICA •2009•44 (1) 137
S.ArbuckleB.,ÖztanA.&Gülçur S.
FIG.4.–Greatest lengthofthe astragalus(GLl)forgoats fromGanjDareh(n=170)(fromHesse1978),AıklıHöyük(n=87),Suberde
(n=4),Çatalhöyükpre-XIItoIV (n=10)(fromRussell&Martin2005),Erbaba(n=30),andKökI-V(n =50).Trianglesrepresent
meanvalues.
138 ANTHROPOZOOLOGICA •2009•44 (1)
Description:in the management of sheep and goats as well as synchronic differences in herding strategies, have .. surplus lambs does “imply that herd management enhanced the potential .. living and breeding under conditions of human.