Table Of Content"
Gayana Zool. 55(3): 247 255. 1991 ISSN 0016-531X
THE AMERICAN GENERA OF ASILIDAE (DÍPTERA): KEYS FOR
IDENTIFICATION WITH AN ATLAS OF FEMALE SPERMATHECA
AND OTHER MORPHOLOGICAL DETAILS. VII. 2. SUBFAMILY
STENOPOGONINAE HULL TRIBES ACRONYCHINI,
-
BATHYPOGONINI (WITH DESCRIPTION OF A NEW GENUS) AND
CERATURGINI, AND A CATALOGUE OF THE NEOTROPICAL
SPECIES*
LOS GÉNEROS AMERICANOS DE ASILIDAE (DÍPTERA): CLA VES
PARA SU IDENTIFICACIÓN CON UN ATLAS DE LAS
ESPERMA TECAS DE LAS HEMBRAS Y OTROS DETALLES
MORFOLÓGICOS. VIL SUBFAMILIA STENOPOGONINAE HULL
2.
TRIBUS ACRONYCHINI, BA THYPOGONINI (CON DESCRIPCIÓN DE
UN NUEVO GENERO) Y CERA TURGINI, Y CA TALOGO DE LAS
ESPECIES NEOTROPICALES
Jorge N. Artigas** and Nelson Papavero*
ABSTRACT RESUMEN
A catalogue of the neotropical species of Acronyches Se presenta un catálogo de las especies neotropicales de
Williston. 1908. only representative of the Tribe Acronyches Williston. 1908, único representante de la tribu
Acronychini, is given. Carebaricus, gen. n. (type-species. Acronychini. Se describe Carebaricus, gen. n. lespecietipo,
Stenopogon rionegrensis Lamas. 1971). the first neotropical Stenopogon rionegrensis Lamas, 1971), el primer
Bathypogonini. isdescribed.A keytothetwonearcticgenera Bathypogonini neotropical. Una clave para los dos géneros
ofCeraturgini ispresented. lllustrationsofthespermathecae neárticos de Ceraturgini es dada, asi como ilustraciones de
and other morphological details are included. espermatecas y otros detalles morfológicos.
Keywords Insecta. Taxonomy. America. Asilidae. Palabras claves Insecta. Taxonomía. América. Asilidae.
Stenopogoninae. Acronychini. Bathypogonini. Ceraturgini. Stenopogoninae. Acronychini. Bathypogonini. Ceraturgini.
This research was supported by the Fundacao de INTRODUCTION
Amparoá PesquisadoEstadodeSaoPaulo(Grants85/1772-
5, 86/3227-1 and 87/3170-8).
Universidad de Concepción, Facultad de Ciencias Previous parts of this series of studies, dealing
Biológicas y de Recursos Naturales, Departamento de with the American genera of Asilidae, with
Zoología. Proyecto Dirección de Investigación. U. de illustrations of the female spermathecae and
Concepción, N" 203812. other morphological details, have been published
Museu de Zoología & Instituto de Estudos
Avancados, Universidade de Sao Paulo. Pesquisador do as follows: Part 1 (Key to subfamilies and
Conselho Nacional de Desenvolvimento Científico e subfamily Leptogastrinae) in Gayana, Zool.
Tecnológico iCNPq. Proc. n" 30.0994/79). 52(l-2):95-l 14, 1988; Part II (Dasypogoninae) in
247
GayanaZool. 55(3), 1991
Gayana, Zool., 52(3-4): 199-260, 1988; Part III alexanderi Papavero, 1971:147. Type-locality:
(Trigonomiminae) in Bol. Soc. Biol. Concepción Brazil, Sao Paulo, Mogi das Cruzes. Distr.
60:35-41, 1989; Part IV (Laphriinae, except Brazil (Rio de Janeiro, Sao Paulo). HT
Atomosiini) in Bol. Mus. paraense EmilioGoeldi, MZUSP.
Zool. 4(2):211-255, 1989; Part V fenestratulus Hermann, 1921:122. Type-locality:
(Stichopogoninae) in Bol. Soc. Biol. Concepción Paraguay, Santa Trinidad; Argentina.
61:39-47, 1990; Part VI (Laphriini (Atomosiini)) Distr.- Brazil (Amazonas) to Argentina.
in Gayana, Zool. 55(l):53-87, 1991; Part VII.1 (Figs. 1,2).
(Stenopogoninae, key to tribes) in Gayana, Zool. giganteus Hull, 1962:313, fig. 204 (Megonyx).
55(2):139-144, 1991. Type-locality: Brazil, Amazonas, Manaus.
The methodology employed in the HT AMNH.
dissection and preservation of the male geosarginus Papavero, 1971:148. Type-locality:
terminalia, female spermathecae and other Surinam, Paramaribo. HT MZUSP.
morphological parts is the same employed by imitator Hermann, 1921:122. Type-locality:
Artigas (1971). "Paraguay". Distr. Brazil (Mato Grosso),
Paraguay. HT MUN.
maya Martin, 1968:180. Type-locality: México,
Tribe ACRONYCHINI Artigas& Papavero Yucatán Chichen Itzá. HT CMNH.
meruuna Papavero, 1971:149. Type-locality:
GenusAcronychesWilliston Brazil, Espirito Santo, Linhares (Parque
Sooretama). HT MZUSP.
Acronyches Williston, 1908:388, fig. 103. Type- plutactites Papavero, 1971:150. Type-locality:
species, WiUistoni Hermann (sub. des., Costa Rica, Turrialba. HT MCZ.
Hermann, 1921:121). rarus Martin, 1968:181. Type-locality: México,
Megonyx Hull, 1962:312. Type-species, Morelos, Yautepec, HT CAS.
=
giganteus Hull (orig. des.) fenestratulus westcotti Martin, 1968:182. Type-locality:
Hermann. México, Nayarit, 20 mi. se. Ixtlán del Río.
HT CAS.
Ref. Papavero (1971) willistoni Hermann, 1921:121. Type-locality:
"Sunnam". HT MUN.
Tribe BATHYPOGONINI Artigas & Papavero
The two genera of this tribe may be recognized by the following key:
Facial gibbosity not occupying entire length of face, but leavinga short fíat área immediately
below antennae; proboscis about as long bristles of mystax, or subequal to length of
face. Occiput only with marginal bristles. Wing short, reaching only segment 5 of abdo-
men. Humeri without bristles. Femora incrassate. Pleura moreor less bare. Male termina-
lia small and slender (Australia) Bathypogon Loew, 1851.
Face gradually sloping from antennal base to oral margin. Proboscis very long and slender,
much longer than length ofbristlesofmystaxor lengthofface(Figs. 3-4). Occiput entirely
coveredby dense bristlesand bristle-like hairs. Wingslightly longerthan abdomen. Hume
ri with several strong bristles. Femora slender. Pleura pilóse, especially above second co-
xa. Male terminalia very large and globose (Figs. 7-9). Female spermathecae as in Fig. 10
(Argentina, Chile) Carebaricus, gen. n.
248
American generaof Asilidae. Stenopogoninae: Artigas,J.N.& N. Papavkro
Genus Carebaricus,gen. n. bristles, similar to the ones on the posterior
(Figs.3-10) callus. Mesopleura with fine hairs only.
Anepisternum bare. Katepisternum with strong
Body length, 25-35 mm. bristles.
Face broad, 1/4 or more width of head (Fig. Legs similar, with strong bristles similar to
4). Oral margin aswide asface (Fig. 3). Frons 1/3 the humeral ones. Claws acute. Pulvilli 3/4
wider than face. Facial gibbosity occupying 3/4 length of claws. Femora slender, not incrassate.
of face, gradually sloping from antennae to oral
Wing: Cell r, open; 2 submarginal cells
omraarlgimna.rgMiynsatnadxswhiotrhterloonng tbhreistulpepseornpacretntoefrthoef present; vein R4 slightly recurrent,ending before
gibbosity. Sides of oral margin bare, shining wing apex; cell r5 closed: anal cell closed at wing
margin or slightly open: cell m-, extremely wide,
posteriorly. Ocellar and frontal bristles and hairs
over twice as wide as high.
similar. Vértex and postocular área with
abundant bristles, mostly directed upwards and Abdomen: First abdominal segment as wide
backwards. Antennae situated on the upper 1/4 as thorax. Abdomen cylindrical, strongly
of head; pedicel and seape of about same length; tapered, 7 tergites visible from above in males, 8
first flagellomere subequal in length tocombined in females. Male terminalia (Figs. 7-9) not
length of scape and pedicel, attenuated on the rotated, epandrium small, divided into two
apical half; second flagellomere minute; third halves, curved in and back at theapex;gonopods
flagellomere pointed at apex, 1/5 length of first. elongated; hypandrium free, large, globose,
Proboscis long, oval in crosssection, straight. curved in on posterior border. Aedeagus small,
Palpus 2-segmented. Occiput entirely covered with only one tube (Figs. 5-6). Female ovipositor
with dense bristles and bristle-like hairs. with strong spines on the acanthophorites.
Thorax: Prosternum dissociated from Spermathecae with 3 elongated black capsules
proepisternum. Pronotum with abundant and fine capsular duets (Fig. 10).
bristles, as long and strong as the postoculars. Type-species, Stenopogon rionegrensis
Mesonotum more or less fíat, covered with fine Lamas, 1971.
scattered hairs. Dorsocentral bristles hair-like,
mostly apressed. Postsutural dorsocentrals
stronger. Bristles on humeral callus, prealar, Carebaricusrionegrensis(Lamas),n.comb.
postalar, postcallar bristles and bristles on
posterior margin of mesonotum long and strong, Stenopogon rionegrensis Lamas, 1971:15, figs. 1,
contrastingwith the fine vestiture ofthe anterior 2-4, 5. Type-locality: Argentina, Río
3/4 of the mesonotal disc. Scutellar disc with Negro, R. Colorado. Distr.- Argentina (Río
scattered fine hairs, 8-10 marginal scutellar Negro, Neuquén). HT MZUSP.
TribeCERATURGINI Artigas& Papavero
Only two genera, both nearctic. No Neotropical representative is known.
KEYTOGENERA:
1. Apex offirst flagellomere and ofsecond flagellomere with deep clefts, in which is received the
attachment ofthe next succeedingsegment Myelaphus Bigot, 1882
Antennae neverasabove. Spermathecaeasin Figs. 11-12 CeraturgusWiedemann, 1824
249
Gayaría Zool. 55(3). 1991
REFERENCES
Artigas. J.N. 1971. Las estructuras quitinizadas de la Lamas M.,G., 1971. DescriptionofthefirstSouthAmerican
spermathecayfundadelpenedelosasilidosysuvalor species of Stenopogon Loew (Díptera. Asilidae).
sistemático a través del estudio por taxonomía Papéisavulsos Zool.. S. Paulo 25(21:15-18, 5 figs.
numérica. Cayana(Zool.) 75:1-106, 138 figs. Martin, C.H., 1968. New Mexican Acronyches and
Hermann. F., 1921. ln L. Zürcher. Dipteren-Ausbeute aus Parataracticus (Diptera: Asilidae). Pan-Pacific
Paraguay: Mydaiden und Asiliden, Arch. Natur- Entomologist 44(4):179 183.
gesch. S7(A, 3).118 122. Papavero, N., 1971. Neotropical Acronyches (Diptera,
Huix. F.M., 1962. Robber flies of the world. The generaof Asilidael, new or otherwise. Papéis avulsos Zool.. S.
thefamilyAsilidae.Smithson. Inst.Bull. 224(I):1-432; Paulo 23\\6):\45-l5\.
(21:433-907, 2.536 figs. Williston. S.W., 1908. ManualofNorthAmericanDiptera
(3rded.l,405 pp., 163 figs. New Haven,Connecticut.
Manuscritoaceptadoenjuniode ¡991.
250
American genera ofAsilidae. Stenopogoninae: Artigas, J.N.& N. Papayero
Figuras 1-2: Acronychesfenestratulus Hermano. I, situation of ihe spermathecae ¡n the abdomen 2 spermathecae
(scales in mmi.
251
Gayaría Zool. 55(3), 1991
Figuras 3-5: Carebaricusrionegrensis(Lamasl, head, in lateral(3)andfrontal(41 views. 5,aedeagusindorsal view(scales
in mm).
252
American genera ofAsilidae. Stenopogoninae: Artigas,J.N.& N. Papavero
Figuras 6-9: Carebaricusrionegrensis(Lamas). Aedeagusin lateral view (61. Maleterminalia in lateral(7), dorsal (81and
ventral (9) views Iscales in mm).
253
Gayana Zool. 55(3), 1991
Figura 10: Carebaricus rionegrensis (Lamas). 10. spermalhecae (scale in mm).
254
AmericangeneraofAsilidae.Stenopogoninae: Artigas,J.N.& N. Papavf.ro
FlGlras 1 1-12: CeralurguscruciatusSay. 11, situaiion of thespermathecae m theabdomen. 12.spermathecae (scales in
mml.
255
