Scottotympana, a new Cicad genus from New Guinea, with the description of three new species, their taxonomy and biogeography (Homoptera, Tibicinidae)

11 Pages·1991·1 MB·English

by  De Boer, A J

Beaufortia
INSTITUTE OF TAXONOMICZOOLOGY (ZOOLOGICAL MUSEUM)
UNIVERSITY OF AMSTERDAM
Vol. 42, no. 1 May 7, 1991
Scottotympana, a newCicad genus from New Guinea, with the
description of three new species, their taxonomy and
biogeography (Homoptera, Tibicinidae)
A.J. de Boer
Institute of Taxonomic Zoology (Zoôlogisch Museum), University ofAmsterdam,
P.O. Box 4766, 1009AT Amsterdam, The Netherlands
Abstract
Thegenus Scottotympanais erected forthree species: S. biardae n.sp. and S.sahebdivanii n. sp. fromnorthern New Guinea
and S. huibregtsaen. sp. from North-East New Guinea and New Britain. The genus belongs toa complex ofgenera
groupedaroundBaeturia Ståland is mostclosely related tothegenera Gymnotympana Ståland VenustriaCoding&Froggatt
ofthis complex. Akey tothe males and a map of distributionare presented. The distribution ofthe genus suggests
anOuter Melanesian Arc pattern.
INTRODUCTION
Material of three undescribed and closely PHYLOGENY
related species was found in the collections of
the Bernice P. Bishop Museum (BPBM), The monophyly of Scottotympana
Honolulu and the Natural History Museum Scottotympana is characterizedby three supposed
(BMNH), London. Thesespecies belong to the apomorphies, all concerning male characters:
“Baeturia and related genera complex" as (1) Apical part of clasper very smooth and
defined earlier(De Boer, 1990), but could not oblong-shaped in lateral view; and supporting
be placed inany ofthegeneraattributedto that aedeagus in upright position with its rounded
complex. Scottotympana is erected to accomodate and inwards protruding proximodorsal corner
these species. The monophyly of the genus is (figs. 15, 22, 28 arrows). Both, shape and pro-
established and its taxonomic position within trusion, are considered synapomorphic.
the "Baeturia and related genera complex" is (2) Lateralcrestsof aedeagus verybroadand
discussed. rectangular at theirproximal end (figs. 12, 13,
The present paper is part of a taxonomic 20, 29).
revision of the “Baeturia and related genera (3) Hind margin of8th tergite strongly con-
complex", which is undertaken in connection vex close to 7th sternite, forming a small hind-
with a historical biogeographic study of New wards directed lobe (compare figs. 1 and 2,
Guinea and the West Pacific islands. arrows).
1
Fig. 5. left tegmen and wing, Scottotympana huibregtsae.
paratype Bululo.
The species of Scottotympana most resemble
Figs. 1-4. 1, caudal part ofabdomen,Scottotympana saheb- Gymnotympana and Venustria in general aspect.
divanii;2, caudal part ofabdomen, Baeturia spec; 3, male The species of these three genera share: (1)
operculum, Scottotympana sahebdivanii;4, male operculum, tegmina with reddish venation and a broad
Aedeastria sepia.
hyaline border along the hindmargin (fig. 5),
(2) a colourpattern of darklongitudinal streaks
The taxonomic position of Scottotympana on thorax and dark spots on head, (3) an
An S-curved aedeagus with wing-shaped lateral extremely long proximal spine on fore femur,
crests is the supposed synapomorphy for the often longer than the distance between prox-
“Baeturia and related genera complex" (De imal and middle spine (fig. 9), and (4) a very
Boer, 1990). Scottotympana is attributed to this short meracanthus. Furthermore, Scottotympana
compex, since the aedeagi of thethree Scottotym- shares with Gymnotympana a strongly curved
pana species are distinctly S-curved and pro- proximal margin of tergite 2, betweenauditory
vided with large lateralcrests. capsule and sternite 2, arching over tymbal
The phylogenetic relationships between the cavity (fig. 3).
various generaand species groups of the com- Only the long femoral spine might possibly
plex are still obscure. And since its sister group be asynapomorpy for thesegenera. The phylo-
is unknown, no decisions about apomorphy or genetic value of the other characters is uncer-
plesiomorphy of more widely distributed tain. Tegmen venation is often ochraceous,
character-states can be made. A major subdivi- though red venation sporadically occurs
sion of the complex is suggested, however, on throughout the generacomplex and is common
account of size and shape ofmale opercula. In in many other groups of cicadas. The hyaline
Baeturia Stâl, 1866, Gymnotympana Stâl, 1861, border along hind margin of tegmen, though
Venustria Goding& Froggatt, 1904and Scottotym- variable in width, is generally much narrower
pana, the male operculum extends medially in the other genera of the complex. However,
beyond meracanthus, whereas the medial both wide and narrow borders are widely
margin ofoperculum, or the greater part of it, distributed outside the complex.
lies laterally of the meracanthusin AedeastriaDe Species of the "Baeturia and related genera
Boer, 1990, Chlorocysta Westwood, 1851, complex" are usually unicoloredochraceous or
Cystosoma Westwood, 1842, Thaumastopsaltria greenish, though many Baeturia species are
Kirkaldy, 1900 and several species erroneously covered with densebrown speckling. A colour
placed in Baeturia (compare figs. 3 and 4). pattern as describedabove for Scottotympana, is
2
also found in Baeturia loriae Distant and some
related species and, although less distinct, in
some species ofChlorocysta. This pattern is very
similar to that foundin several other groupsof
cicadas.
The meracanthusis very short in Gymnolym-
pana and Venustria, slightly longer in Scottotym-
pana, but usually longer and more slender in
other groupsofthe complex. Cicadettinihave a
similar short meracanthus.
The arched proximal margin of tergite 2 is
probably a plesiomorph character, since a very
similarly shaped 2nd tergite is widely
distributede.g. in Cicadettini and Prasiini. In
ohter groups of the complex, this margin is
straight betweenthe auditory capsule and ster-
nite 2 (compare figs. 3 and 4). Figs. 6-9. Scottotympanabiardae,holotype: 6,headin dorsal
view; 7, postclypeus in lateral view; 8, first and second
The phylogeny of the species of Scottotympana abdominal segments in dorsal view; 9, fore femur.
It is most probable that S. biardae and S.
huibregtsae formthe sister taxon ofS. sahebdivanii.
Possible synapomorphies for biardae and margin convex, continuous with anterior
huibregtsae are the lateral plates on the claspers margins of vertex lobes. Postclypeus in lateral
and thelaminiform lobeat thedistal margins of view (fig. 7) slightly swollen, anterior margin
pygofer. Furthermore, these species share (lateral view) slightly convex. Vertex broad,
lateral rows of very light spots onthe abdomen. 1.4-1.5 x as broad as postclypeus, and
S. sahebdivanii and S. huibregtsae share an oval smoothly vaulted. Vertex with distinct medial
operculum, but this character is found in furrow, butwithout diverging furrows between
several related genera e.g. Venustria and frontal and lateralocelli. Ocelli small and wide
Baeturia. apart, distance betweenlateral ocelli distinctly
longer thandistancebetweenlateralocellusand
eye. Tegmina with eight apical areas, wings
with six (fig. 5). Tegmen venation, especially
TAXONOMY
costa, red. Tegmina and wings with broad
hyaline border along hind margins. Fore femur
Scottotympana n. gen. with row of three erect spines, diminishing in
Type species: Scottotympana biardae n. sp. length towards tibia; proximal spine longer
than distance to middle spine (fig. 9). Tym-
Description. — Body ochraceous to brown, panum with 5-7 parallel sclerotized ridges.
with pattern of dark spots on head, and dark Distal partofoperculum partly covering tymbal
streaks on thorax. Distal part of abdomen cavity in ventral view and reaching medially
darkened. Head (fig. 6) broad and short, beyond meracanthus. Meracanthus very short,
broader thananterior margin ofpronotumand reaching to about half the operculum length.
about as broad as anterior margin of Abdomen inflated. Proximal margin of second
mesonotum. Distance betweeneyes longer than tergite strongly curved, mesiad to auditory cap-
length ofhead. Postclypeus in dorsal view only sule, arching over tymbal cavity. Eighth tergite
slightly protruding beyond vertex lobes and forming asmall lobe at distoventralcorner (fig.
smoothly rounded anteriorly. Its anterior 1 arrow). Pygofer with short and sharply
3
pointed caudodorsal beak. Ventralmargins of Etymology. — Early last year, I was asked by
pygofer converge in sharp angle to base of theDutch branch oftheWorldWildlifeFund to
pygofer opening and continue as parallel ridges name threenew species ofcicadas after threeof
over underside of pygofer (fig. 18). Claspers theirjunior members(Rangers), the winnersof
parallel, not fused at theirbases, and thus not a fund raising competition, related to their
forming a ring aroundanal valves. Apical part biodiversity programme. The new species des-
of clasper very smoothly rounded, with large cribed in this paper are named after Patricia
and sharp-edged ventral hollow; proximodorsal Biard, Lindy Huibregtse and Sahand Saheb-
corner of this apical part serves as support for divani, honouring theireffords in raising funds
aedeagus. Aedeagus S-curved, and with broad for the WWF.
angular lateral crests. The genus was named in honourofSir Peter
Scott, founderofthe WorldWildlifeFund, The
Distribution. — Scottotympana is recorded suffix tympana is applied, to indicate the rela-
mainly from northern New Guinea. One tionship with Gymnotympana.
specimen ofS. huibregtsae probably comes from
New Britain, one specimen ofS. biardae comes
Key to the males
from central New Guinea(fig. 10).
la. Distal pygofer margin strongly convex, for-
ming a semi-circular laminiform protru-
sion. Lateral lobes of pygofer with
conical-shaped, slightly upwards curved,
protrusion. Ventralpygofer margin angu-
larly bent, just underneath protrusion of
lateral lobe of pygofer. Caudodorsal beak
erect. Clapser with distinct lateral plate at
base. Body length over 22 mm 2
b. Distal pygofer margin weakly convex,
without laminiform protrusion. Lateral
lobeof pygofer strongly bending outwards,
forming a rectangular lateral protrusion.
Ventral pygofer margin straight to apex of
protrusion of lateral lobe of pygofer (fig.
26). Caudodorsal beak strongly bent.
Clasper thickened, but withoutlateralplate
Fig. 10.Localities ofScottotympanabiardae,S. huibregtsaeand
at base. Body length under22 mm
S. sahebdwanii.
S. sahebdivanii.
2a. Body light brown. Middorsal part of first
New Britain and the northern mountain tergite totally hidden under metanotum
ranges of New Guinea are fragments of a (fig. 8). Distal part of operculum angular
historic island arc: the Outer Melanesian Arc (fig. 11). Clasper with large, square-
(see Hamilton, 1979; Holloway, 1979; 1984). shaped, lateral plate at base. Apical part of
Land masses originating from this arc have clasper elongate, finger-shaped, at prox-
been recognized as areas of endemism for imodorsal corner (fig. 15) S. biardae.
several groups of cicadas (Duffels, 1986; De b. Body colour, with very light parts con-
Boer, 1989; Duffels & De Boer, 1990). The geo- trasting with black-brown parts. Middorsal
graphic data of Scottolympana suggest, that this partof first tergite only partly hiddenunder
genus evolved within the Outer Melanesian metanotum (cf. fig. 25). Distal partofoper-
Arc. culumrounded, oval (fig. 19). Clasper with
4
small, triangular, lateral plate at base. end of medialband. In the paratype two pairs
Apical part of clasper slightly elongate, ofirregular shaped, broaddark bandsrun in an
broadly rounded, at proximodorsal corner along medial and lateral fissures; the lateral
(fig- 22) S. huibregtsae. bands only, reach margin of pronotal collar,
and fuse with dark brown streak along lateral
Description ofthe species partofthat margin. In theholotype, theselatter
markings are faded and hardly visible.
Mesonotum light brown with two large
Scottotympana biardae n. sp.
paramedian brown spots bordering pronotum,
(figs. 6-18)
and reticulate pattern of angular brown spots
Holotype: "New GuineaNE. Eliptamin Valley forming lateral bands from pronotum margin,
1200-1350 m. Aug. 16-30, 1959" (print); converging to corners of cruciform elevation.
"W.W. Brandt Collector BISHOP" (print), Two black spots infrontof cruciform elevation.
O", BPBM; Paratype: Lower Jimmi V., 1965, Tymbal organ: Holotype with 6 dark
native coll., 1 O", BPBM. sclerotized parallel ridges spanning tymbal
from dorsal to ventral margin. The most prox-
S. biardaeis easily recognized by thelarge lateral imal ridge narrows considerably towards ven-
plates on clapsers and its large, almost rec- tral margin and only just reaches it. Six short
tangular operculum. There are considerable and lighter coloured intercalary ridges seem to
differences in shape of clasper and aedeagus form a longitudinal band across tymbal.
between the two available specimens, so they Paratype has onecomplete ridge and one inter-
possibly represent two separate species. calary ridge less than holotype.
Description. — Body light brown, with distinct Operculum (fig. 11): Larger than in both
pattern of brown bands on thorax. Abdomen other species, almost completely covering tym-
longand slender, 1.2-1.3 x as long as headand bal cavity in ventral view. Basal part of oper-
thorax. Tegmina 1.1-1.3 x as long as body culum strongly vaulted, especially medially,
length. wherethe vaulting abruptly bendsdown tobase
Head (fig. 6): Ochraceous, slightly darker ofmeracanthus. Basalpart darkbrown suffused
brown on postclypeus, vertex lobes and at distolateral corner and on slope of medial
between ocelli. Vertex lobe with two, almost vaulting. Distal part of operculum plate-like,
round, dark spots between lateral ocellus and very angular, almost square-shaped and erect.
eye, and a triangular dark spot at prox- Distal partnarrower thanin both otherspecies,
imomedial edge of eye. Head 2.7-2.8 x as only just reaching medially beyond meracan-
broad as long. Postclypeus 2.5-3.2 x asbroad thus. Its lateralmargin longand straight, rising
as long. Sides of postclypeus (fig. 7) with 5-6 from close to distolateral corner of basal part.
vague and irregular rows of short parallel Distal margin shorter and slightly convex,
ridges. Vertex 1.6-1.8 x as broad as long and medialmargin still shorter and straight. Distal
1.1-1.2 x as broad as length of head. Distance margin making an almost right angle with
betweenlateralocelli 2.6-2.7 x widthof frontal lateral margin, distomedialcorner rounded.
ocellus, and 1.3-1.5 x distancebetween lateral Abdomen: Long and slender, light
ocellus and eye. oehraceous, but slightly darkenedin middorsal
Thorax: Pronotum light brown, with two band, onsides oftergites 6-8and onsternites 6-
narrow, dark brown longitudinal streaks, 7. Segmental hind margins slightly reddish.
bordering a broad and ochraceous middorsal Sides of abdomen with 2 parallel rows of light
band. These streaks broaden anteriorly to patches, and lateroventral row of very slightly
triangular spots on both sides of middorsal darkened spots, on segments 3-7. First tergite
band. A pair of triangle-shaped paramedian (fig. 8) very short, its medial part completely
spots, at margin of pronotal collar, marks the hidden under metanotum. Second tergite very
5
Figs. 11-18.Scottotympanabiardae: 11,operculum, holotype; 12, aedeagus,holotype;13, aedeagus, paratype; 14,pygofer
in lateral view, holotype; 15,claspers, detail apicalpart, holotype; 16, caudodorsal beak in dorsal view, holotype; 17,
pygofer inlateral view,paratype; 18,pygoferaslant,paratype. Lettering: bp=basal part ofoperculum;c=crest around
distolateral corner; cdb =caudodorsal beak; di=distal margin of pygofer; dm =distal margin of operculum;
dmc=distomedialcornerofoperculum; do=dorsal margin of pygofer; dp =distal part ofoperculum; lm =lateral
marginofoperculum; lp=lateral protuberance; ve=ventral margin ofpygofer.
long middorsally and reaching metanotum,its sharply pointed atapex. Lateral lobeof pygofer
proximal margin strongly convex middorsally. in holotype with conical, slightly upwards
Genitalia: Pygofer in lateral view (fig. 14, curved and distinctly hindwardsprojecting pro-
17): Dorsal margin concave, continuous with tuberance. In the paratype this protuberance
straight and erect caudodorsal beak. Distal much smaller, more angular and less far
margin concave from beak, but strongly convex projecting hindwards. Ventral margin of
at halfits length, forming an inwards curved pygofer convexly bent, forming angular corner
semi-circular laminiform protrusion in holo- under protuberance. Clasper very conspicuous
type (fig. 18); this protrusion is very sharply by its large laminose and square-shaped lateral
bent inwards, and therefore not visible in plate (in holotype larger than in paratype).
lateral view, in paratype (fig. 17). Caudodorsal Apical part ofclasper forming elongate, finger-
beak in dorsal view (fig. 16) triangular, very shaped, proximodorsal protrusion. This protru-
6
sion supports aedeagus in upright position. In ochraceous areas between lateral ocelli and
lateral view (fig. 14, 17) this protrusion lies eyes. Vertex with two pairs of dark spots within
partly beyond lateralplate, but is not fusedwith these light areas, and a triangle-shaped dark
it (fig. 15); lateral plate connected to ventral spot at proximomedial edge of eye. Head
part of clasper only. Aedeagus very long and broader than in both other species, 3.1-3.4 x
broad, strongly S-curved; apical part almost in as broad as long. Postclypeus 2.3-2.4 x as
right angle on medialpart. Aedeagus porevery broad as long. Sides of postclypeus with 5-7
large and oval. Size and shape of aedeagus, rows of short parallel ridges. Vertex very
however, quite different in the two available broad, 2.0-2.2 x asbroad as long and 1.4-1.5
specimens. Aedeagus of holotype (fig. 12) x asbroadaslength ofhead.Distance between
distinctly longer than that ofparatype (fig. 13), lateralocelli 3.0-3.3 x width offrontal ocellus,
with longer and oblong-shaped lateral crests. and 1.2-1.3 x distancebetweenlateral ocellus
These crests triangle-shaped, narrowing more and eye.
gradually towards aedeagus apex, in paratype. Thorax: Colour pattern almost the same as
Measurements: Body length: 23.2-26.2 mm; in biardae, but more strongly contrasting. Pro-
tegmen length: 30.0-31.0 mm; pronotum notum light brown, with two narrow dark
length: 3.0-3.3 mm; mesonotum length: 4.9- brown longitudinal streaks, bordering a broad
6.0 mm; headlength; 2.1-2.4mm; headwidth: and ochraceous middorsalband. These streaks
6.0-6.6 mm; width of pronotal collar: 7.7- broaden anteriorly to triangular spots on both
8.5 mm. sides of middorsal band and diverge sharply
Distribution (fig. 10): Central west Papua near pronotal collar. A short dark middorsal
New Guinea. streak, along pronotal collar, marks the end of
medial band. Two pairs of irregular shaped,
broad dark bands run in and along medial and
Scottotympana huibregtsae n. sp.
lateral fissures; the lateral pair only, reaching
(figs. 5, 10, 19-24)
margin ofpronotal collar, and fusing withdark
Holotype: "New Guinea NE Lae, Singuawa brown streak bordering lateralparts ofpronotal
R. 147° 10' E 6° 45' S 80 m. 16.iv.1966" collar. Mesonotum light brown with two large
(print); "Pri. Forest" (print); "O.R. Wilkes paramedian brown spots bordering pronotum.
Light Trap BISHOP Mus." (print), cr, A reticulate pattern of angular brown spots
BPBM; Paratypes: PAPUA: NEW GUINEA formslateralbands frompronotum,converging
(NE): Bulolo, 700 m, 26.xii.1969, J. Sedlacek, to corners ofcruciform elevation. A broad light
lO*, BPBM; Lae, viii. 1944, F.E. Skinner, lc, brown streak lies lateral to these bands. Two
BPBM; BISMARCK ARCHIPELAGO: black spots in frontof cruciform elevation.
Bismarck Arch. Buololo, xi.1934, J.L. Frog- Tymbal organ: Six dark red-brown, parallel,
gatt, 1er, BMNH. sclerotized ridges spanning the tymbal from
S. huibregtsae is ofabout the same size asS. biar- dorsal to ventral margin and a 7th most prox-
dae, but is darkercolouredand has a relatively imal ridge spanning about 3/4 tymbal width.
shorter abdomen. S. huibregtsae is easily recog- Six verydistinct, light brown, intercalary ridges
nized by its dark brown abdomen with light, seem to form a longitudinal band across
almost transparent sides. tymbal.
Operculum (fig. 19): Distinctly shorter than
Description. — Body brown, with distinct pat- that of the foregoing species and less angular,
tern of light and dark bands on thorax. only partly covering tymbal cavity in ventral
Abdomen short and broad, 1.0-1.2 x as long view. Basalpart of operculum strongly vaulted
as headand thorax. Tegmina 1.2-1.4 x as long as inbiardae, witha similarabruptly downwards
as body length. curved medialvaulting. Basal part dark brown
Head: Dark brown, but with large suffused at distolateral corner and on slope of
7
Figs. 19-24.Scottotympana huibregtsae: 19,operculum, holotype;20, aedeagus,paratypeBulolo;21, pygoferaslant,holo-
type; 22, claspers, detail apical part, holotype; 23, caudodorsal,beak in dorsal view, holotype; 24, pygoferin lateral
view, holotype.
media] vaulting. Distal part of operculum spots on segments 2-8, even distinguishable on
rounded, oval and curved towards abdomen. lighter parts of abdomen. Middorsal part of
Its lateralmargin shortand almoststraight, ris- first tergite very short and partly hidden under
ing from close to distolateral corner of basal metanotum. Second tergite much shorter than
part and curving gradually into slightly convex in biardae (cf. fig. 25), its proximal margin
distalmargin. Medial margin strongly convex. slightly convex middorsally.
Abdomen: Shorter and rather stout com- Genitalia: Pygofer in lateral view (fig. 24):
pared to foregoing species, with more contrast Dorsal margin almost straight, angularly bent
in colouration. Second tergite dark brown into straight and erect caudodorsalbeak. Distal
laterally, but light ochraceous dorsally. Third, margin convex between base of caudodorsal
fourth and part of fifth tergite very light, and beak and lateral protrusion, and forming an
almost transparent laterally, but dark brown inwards curved protrusion, similar to that of
dorsally. Distal halfof 5th tergite and wholeof biardae, though smaller.Caudodorsalbeak very
6th and 7th tergite dark brown. Proximalpart long and slender, spine-shaped and sharply
of 8th tergite very light, distalpart dark. Ven- pointed at apex (fig. 23). Protrusion on lateral
tral side ofabdomen with large oval dark spot lobe of pygofer small, conical-shaped and
covering sternites 5-7. Hind margins of ster- curvedupwards, bluntly roundedatapex. Ven-
nites 3-4 slightly darkened. Lateral sides of tralmargin slightly convex and forming aright-
abdomen with 2 parallel rows of small light angled corner just underneath this pro-
8
tuberance. Clasper (fig. 22) very similarto that very vague markings on headand thorax. Hind
of biardae, but with a smaller lateral plate. margins of abdominal segments bright red.
Lateral plate more triangle-shaped, its heighth Abdomenin males 1.3 x as long as head and
not exceeding dorsal clasper margin (lateral thorax, in female 1.2 x . Male tegmen 1.2-1.3
view). Apical partofclasper broadly roundedat x as long as body length, in female 1.2 x .
its proximodorsal corner. This slightly pro- Head: Ochraceous, slightly darker brown on
truding corner supports aedeagus in upright postclypeus. Two specimens with a triangular
position. Aedeagus (fig. 20) long and broad, darkarea betweenocelli. Thetwo pairs of dark
though smaller than in biardae, and strongly S- spots between lateral ocelli and eyes, as des-
curved. Lateral crests of aedeagus very broad cribedforboth other species, are fused to single
and triangle-shaped (as in biardae paratype). dark streaks. A triangular dark spot at prox-
Aedeagus pore large and oval. imomedial edge of eye. Head 2.7-2.9 x as
Measurements: Body length: 22.0-25.6 mm; broad as long. Postclypeus 2.8-4.0 x as broad
tegmen length: 30.1-31.2 mm; pronotum as long. Sides of postclypeus rather smooth,
length: 3.1-3.4 mm; mesonotum length: 5.7- with some indistinct rows of short parallel
5.8 mm; headlength 1.9-2.0mm; head width: ridges. Vertex narrower thanin huibregtsae, 1.8-
6.6-6.8 mm; width of pronotal collar: 7.6-8.2 1.9 x asbroad as long and 1.3-1.4 x asbroad
mm. as length of head. Distance between lateral
Distribution (fig. 10): Northwestern part of ocelli 2.9-4.5 x width of frontal ocellus, and
Papuan Peninsulaand, probably, New Britain 1.2-1.5 x distancebetweenlateral ocellus and
(see remark). eye.
Remark: The locality "Buololo, Bismarck Thorax: With less distinct colour markings
Archipelago" couldnot be tracedBuololocould than both foregoing species. Pronotum
be a mis-spelling of Bulolo on New Guinea, ochraceous, with 2 small paramedian brown
which is the more likely, since the species is spots at margin of pronotal collar, and often
known from that locality too. However, the slightly darkenedalong fissures. One male with
label definitely mentioned Bismarck slightly reddish middorsal band on pronotum
Archipelago, and since Mr. Froggatt was sta- and one male with broad castaneous brown
tioned at Rabaul (New Britain), as Govern- middorsalband over anteriorhalfofpronotum
ment Entomologist (Van Steenis-Kruseman, only. Mesonotum light brown with two black
1950), the specimen probably comes from that spots in front of cruciform elevation. Three
island. specimens, as in foregoing species, with two
large paramedian brown spots bordering pro-
notum and a reticulate pattern of angular
Scottotympana sahebdivanii n. sp. brown spots forming lateral bands from pro-
(figs. 1, 3, 10, 25-33)
notum, converging to corners of cruciform
Holotype: "New Guinea Neth. Bodem, Sarmi elevation.
Areavii-10-1959"(print); "T.C. Maa Collec- Tymbal organ: Five light brown, parallel,
tor BISHOP" (print), O", BPBM; Paratypes: sclerotized ridges spanning tymbal from dorsal
same data as holotype 2C, 1Ç, BPBM; same to ventral margin, and a 6th and 7th proximal
data lO", Instituut voor Taxonomische ridge spanning about 4/5 and 2/3 of tymbal
Zoologie, Zoôlogisch Museum, Amsterdam. width respectively. Intercalary ridges not
clearly separated from main ridges.
5. sahebdivanii is distinctly smaller than both Opercular Male operculum (fig. 3) closely
foregoing species. It can be recognized by its resembling thatof huibregtsae, only partly cover-
strongly bent caudodorsal beak. ing tymbal cavity inventral view. Basal part of
operculum less vaulted than in foregoing
Description. — Body ochraceous, with only species, more gradually curved down mesiad.
9
Figs. 25-33. Scottotympanasahebdivanii: 25, first and second abdominal segments in dorsal view, holotype; 26, pygofer
aslant,holotype; 27,pygoferinlateral view, holotype;28,clasper, paratype;29,aedeagus,paratype;30, female genital
segment in lateral view; 31, male caudodorsal beak in dorsal view, holotype; 32, female operculum; 33, female
caudodorsal beak in dorsal view.
Distal part of male operculum oval and curved metanotum. Proximalmargin ofsecond tergite
towards abdomen, as in huibregtsae. Its lateral slightly convex, almost straight middorsally.
margin short and straight, rising from close to Femaleabdomenas in male, withits caudalhalf
distolateral corner of basal part and curving darkened. Femalegenital segment(fig. 30) long
gradually into slightly convex distal margin. and slender; greatest length longer than
Medial margin strongly convex. Femaleoper- greatest width. Ovipositor sheaths reaching
culum(fig. 32) very short. Its distalpartshorter well beyond apex of caudodorsalbeak. Female
thanbasal part, sickle-shaped and erect. caudodorsalbeak in dorsal view (fig. 33) stout,
Abdomen: Maleabdomen slenderas in biar- triangle-shaped and sharply pointed at apex.
dae, ochraceous, but segments 6-8 dark brown. Male genitalia: Pygofer in lateral view (fig.
Abdomen in one male entirely dark brown. 27): Dorsal margin almost straight, angularly
Segmental hind margins bright red. Abdomen bending into straight and hindwards directed
not darkenedmiddorsally, lateroventralrow of caudodorsal beak. Distal margin slightly con-
slightly darkened spots hardly visible. First vex between base of caudodorsal beak and
tergite (fig. 25) short and partly hidden under lateral protrusion, though not forming the
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