Table Of ContentBLUMEA 41 (1996) 455-462
New taxaand combinationfor the Rubiaceae of
Sri Lanka
C.E. Ridsdale
Summary
In the familyRubiaceae four newtaxa inHedyotis and three newtaxa inLasianthus aredescribed;
tenother combinations arealso made.
Thisshortpaper validatesnomenclaturechanges andnewtaxa intheRubiaceaefor
theaccount ofthefamily fortheFloraofCeylonProject.
Hedyotis gartmorensis Ridsdale, spec. nov.
Hedyotis quinquinervisThwaitesaffinisfoliis suprascabridis infrapubescentibusdiffert.—
Typus: Weerasooriya,Jayasekera& Ridsdale 2060PDA (holoK; isoL), Sri Lanka.
Shrub. Stems densely pubescent. Stipules distinctly sheathing, persistent, on the
flowering shootsthebasalpartofthestipules 3-5mmlong,outsidepubescent, api-
cal rim fibriate, fimbriae9-13, short, 0.5—2(—3) mm long. Leaves broadly ovate,
1-3.5x0.8-2cm,abovescabrid, particularly onthenerves, belowpubescent, den-
sely so onthe nerves, margins revolute,apex acute, basesubcordate, lateralnerves
3pairs, appearing 5-nerved atthebase. Petiole 1-3 mm, densely pubescent. Inflo-
rescence terminal, subcapitate, 1-2 cm long, axis up to 3 cm. Flowers unknown.
Hypanthium 1-2mm long, pubescent, calyx tube short, lobes narrowly triangular
2-3 x0.5-1 mm, margins ciliate. Corollaunknown. Capsule ellipsoidal, 2-4 x
1.5-2mm, surmountedwith calyx remnants, seeds4-6 per placenta, angled to
planoconvex, brownishblack.
Distribution — SriLanka: Kandy District. Only known fromthe typecollected
aboveGartmoreEstate
Ecology — Uppermontane shrubberies.
Note—Closely relatedtoH. quinquinervia Thwaites.Furthercollectionrequired,
particularly floweringmaterial.
1) c/oRijksherbarium/Hortus Botanicus,P.O.Box9514,2300 RA Leiden. TheNetherlands.
456 BLUMEA Vol. 41, No. 2, 1996
Hedyotis neolessertianaRidsdale, spec. nov.
Hedyotis lessertiana auct. nonAm.,Pug. PI.Ind. Or. (1836)21; Nova ActaAcad. Caes.Leop.-
Carol. Nat.Cur. 18(1837)339;Thwaites,Enum. PI.Zeyl. (1859) 141;Hook.f.,Fl.Brit. India
3(1880)52;Trimen,Handb.Fl.Ceylon2(1894)309.—Oldenlandia lessertiana (Arn.)Kuntze,
Rev.Gen. PI. 1 (1891)292.
Hedyotislessertianae Arn. affinis caulibus gracilibus,stipulis separatisinternodiis longiori-
bus, inflorescentiis laxis pedicellis3-5 mm longisdiffert.— Typus: Thwaites CP 99(hole
PDA; iso K), Sri Lanka.
Shruborsmall tree up to3 m,stems slender.Stipules connate, distinctlysheathing,
persistent, and becoming scarious, widely separated fromeach other by the inter-
nodes, thebasal part ofthe stipules 0.7-1.2 cm long,outside glabrous, apical rim
fimbriate,fimbriae9-13,short, 1-6mm long,densely floccose attheapex.Leaves
lanceolatetoelliptic,(4—)7—10(—12)x(1—)1.5—2(—2.5) cm,aboveglabrousorsparse-
ly hairy onthemidrib,belowsparsely pallidly hairy particularlyon thenerves, apex
acute toacuminate, baseacute,sometimesdecurrent, lateralnerves4-6pairs. Petiole
up to 1.5cm long, glabrous. Inflorescence aterminalcyme, axis up to 10cm long,
lateralaxes upto 8 cm.Flowers heterostylous. Hypanthium 0.8-1.5mm long, gla-
brous orpubescent, pedicels 3-5 mm,calyx tubeshort, lobes deltoidto narrowly
triangular,0.7-1.5 x0.5-0.8mm. Corollainfundibular, tube4-5mmlong,outside
glabrous, insidedensely pubescent particularly atthethroat, lobes ovateto oblong,
2-3 mm long, incurvedatthe apex,outsideglabrous or with afew scattered hairs,
insidedensely pubescent. Filamentsshort, 0.1-0.3mm, or long,2-4 mm,anthers
linear, 0.5mm long,thin. Style 3-4mmor6-8mm,stigma bilobed.Capsule ellip-
soidal, 2-3x 1-2mm,surmountedwith calyx remnants,seeds6-12 perplacenta,
angled toplanoconvex, blackish.
Distribution— SriLanka; Kandy District (5 coll.), Ratnapura District(1 coll.).
Ecology —Upper montane shrubberies, ofteninopenandsecondary area.
Note — Twaites, Hooker, and Trimen did not examine Arnott's typespecimen
whichrepresentsthetaxon known toTrimenasH.confertiflorum.The taxon thought
torepresentH. lessertianaby previous authorshasremainedundescribed
Hedyotis tridentataRidsdale,spec. nov.
HedyotiscoprosmoidesTrimen,Handb. Fl.Ceylon 2(1894)306,p.p. CP 91.
?Hedyotis obscura var.minorThwaites exHook,f.,Fl.Brit.India 3 (1880)52,nonTrimen (1894).
Type:SriLanka,notindicated.
Hedyotis obscura var.BThwaites,Enum.PI.Zeyl. (1859) 141,p.p.CP 91.
Hedyotis obscura Thwaites affinis stipulispartibusapicalibus 3fimbriis differt.—Typus:
SriLaka, Weerasooriya, Jayasekera<6 Ridsdale 2064 (holoPDA;iso K,L).
Stragling trailingshrubup to3m.Lateralbranchesdistinctly supra-axillary inorigin.
Stipules distinctly sheathing, ontheflowering shoots thebasal partof thestipules
1-2 mmlong, hoop-like, apical part fimbriate,2-5 mmlong, fimbriae1 with2lat-
eralfimbriae, onthevegetativeorthotropic shootsthebasal part2-4mmlong, apical
part5-8mmlong. Leaves ovate tobroadly elliptic,3-5x 1-3cm, thoseofthe or-
thotropic vegetativeshoots larger upto 6x 4cm, aboveglabrous, below glabrous,
C.E.Ridsdale: New taxa for the Rubiaceae ofSri Lanka 457
apex acute,baseacute, often decurrent,lateralnerves 3-4pairs. Petioleupto 1 cm
long. Inflorescencelateral, axis distinctly supra-axillary, up to4cm long, the lateral
branches short, upto 1 cm, few-flowered.Flowers heterostylous, subsessile orwith
apedicel upto 2mm long. Hypanthium 0.7-1.2mmlong, glabrous,calyx tubeshort,
lobes deltoidto narrowly triangular (0.6-)l-2 x 0.5-1 mm, glabrous. Corollain-
fundibular, tube4-5 mm long, outside glabrous, inside densely pubescent atthe
throat,lobesovate tooblong, 3-4 mmlong, incurvedattheapex, outsidesparsely
pubescent, inside densely pubescent. Filaments short, 0.1-0.5mm, or long, 1-2
mm, anthers linear-oblong, 1-2 mm long. Style 3-4 mmor6-8 mm, stigma bi-
lobed.Capsule ovoidal, 4-6x 3-4mm, apex domedbut surroundedwithincurved
calyx remnants, seeds l(-few?) perplacenta, angled, blackish.
Distribution—SriLanka: Kandy District(6coll.).
Ecology —Open montane shrubberiesandforestedges.
Note—Thereis someconfusionaround 'var.minor’.Hooker's description proba-
bly applies tothisspecies. NoPeradeniya specimens bearalabel'H.ovata var.minor’.
Hedyotis trimeniivar. orbicularifoliaRidsdale, var. nov.
A Hedyotistrimenii Deb& Dutta var.trimeniifoliis orbicularibus apicibus etbasibus ob-
tusis, marginibusrevolutis,nervis lateralibus indistinctis differt.-—Typus: Weerasooriya,
Jayasekera&Ridsdale 2057(holoPDA;iso K,L),Sri Lanka.
Leavesorbicular, apexobtuse,base obtuse,marginsrevolute, lateralnervesindistinct,
notvisible.
Distribution—SriLanka:Kandy District(3 coll.).
Ecology —Open montaneforestin Adam'sPeakarea.
Thetaxoncanbe distinguished fromvar. trimeniiasfollows:
- Leaves ovate to elliptic, apex acute, base acute to cuneate, margins usually not
revolute, lateralnervesobscurebutusually visable var. trimenii
- Leaves orbicular, apex obtuse, baseobtuse, margins revolute, lateralnerves in-
distinct, not visable var. orbicularifolia
Knoxiaspicata (Thwaites ex Trimen) Ridsdale,comb. nov.
Knoxia platycarpavar.spicata Thwaites exTrimen,[Syst. Cat. (1885)44, nom. nud.] Handb. Fl.
Ceylon 2:(1894)342. Type:Thwaites CP 3996 (PDA),Sri Lanka.
Distribution— SriLanka:MataleDistrict(1 coll.),Kandy District(1 coll.),Badul-
laDistrict (2coll.), NuwaraEliya District(29coll.).
Ecology —Upper montane areas, inforest andopen sites.
Lasianthus chrysocaulis Ridsdale, spec. nov.
Lasianthusneolanceolato Ridsdale maxime similis.Ramulis glabrisvel sparsepubescenti-
bus saepe siccitate flaveo-brunneis,stipulis angustetriangulatis,floribus subsessilibus.
calycium lobis 0-1 mm longis, corollae tubo 3-4 mm longis extus glabro, fructibus
ovoideis 5-8 mm. —Typus: Alston 1428 (PDA),Sri Lanka.
458 BLUMEA Vol. 41, No. 2, 1996
Shrub. Branchlets glabrous, rarely sparsely hairy and rapidly becoming glabrous,
often drying yellow-brown. Stipules narrowly triangular, 1-2x0.5 mm, sparsely
hairy. Leaves lanceolate, (3-)4-8(-12) x (0.7-) 1.3 cm, above glabrous, below
glabrous or with a few scattered hairs, apex long-acuminate, base acute, lateral
nerves 3-4(-6) pairs, prominently raisedbelow, distinctfrom the tertiary nerves.
Petioleup to0.7 cm long, glabrous to sparsely hairy. Flowers 1-3. Hypanthium
ovoid, 0.7-1.5 mm long, glabrous. Calyx campanulate, tube short, 0.2-0.5 mm,
outside glabrous or withafewscattered hairs,lobestriangular, 0.5-0.7mmlong,
margins witha few scattered hairs. Corollatube 3-4mm, outsideglabrous, lobes
1.3-2mmlong,outsidewithafew sc attered hairs.Fruits 5-8mm.
Distribution—Sri Lanka: Kandy District(3 coll.), Ratnapura District (3 coll.),
Galle District (1 coll.).
Ecology —Montaneshrubbery.
Note—Thiselementwithglabrous stems, oftendrying yellowish,was mixedby
Thwaites andTrimenintheirconceptof ‘L. walkerianus’.Furthercollectionsarere-
quired toevaluatetherelationship toL.neolanceolatus.
Lasianthus foetulentusRidsdale, spec. nov.
Lasianthus walkerianus auct.nonWight: Hook,f.,Fl.Brit.India3(1880) 186.
Lasianthus varians (Thwaites)Thwaites,Enum. PI.Zeyl. (1824)420,p.p.
Lasianthus varianti ( Thwaites)Thwaites emend Hook.f. affinisplantis foetidis,glabratis,
stipulis lanceolatis,floribus subsessilibus nonpedunculis,calycium lobis 1-1.3 mmlon-
gis, corollis glabris,fructibus ovoideis 3-5 mm,nigris differt.—Typus: Weeasooriya,
Jayasekera&Ridsdale 1973 (holoPDA;iso K,L),Sri Lanka.
Shrub.Branchletsglabrous, crushedplantsfoetid.Stipules lanceolate,3-5x0.5mm,
glabrous orwitha fewrusset hairs atthetop.Leaves elliptic toobovate 3-8x 1-3
cm,above andbelowglabrous, lateralandtertiary nerves prominently raisedabove
andbelow, apex shallowly acuteandabruptly apiculate, baseacute, lateralnerves
3-5(-8)pairs, scarcely distinctfromtheprominent, reticulate, tertiary nerves. Peti-
ole up to 1.5 cm long, densely hairy. Flowers 2-6 per axil. Flypanthium ovoid,
0.8-1 mm long, glabrous. Calyx campanulate, tube c. 1 mm,outside glabrous or
withafew scattered hairs,lobestriangular, 1-1.3mmlong, margins withafewscat-
teredhairs. Corollainfundibular, tube3 mm, outside glabrous, lobesovate, 2mm
long, reflexed,abovehairy,hairscurly. Style5 mm exserted.Fruits ovoid, 3-5mm,
black.
Distribution— SriLanka.
Ecology — Montaneshrubberies.
Notes— Thewholecomplex, L. varians-walkerianus-stigosus, has beencontin-
ually confused in thepast. Ihave,therefore, takenthechoice oflectotypifying L.
walkerianthusby theright handelementwhichclearly is identicaltoL. strigosus, and
describingthecommon widespread plantofHortonPlains as anewspeciesbasedon
amoderncollection.
Thethirdelement'c' is treatedas aseparatevariety.
Lasianthus walkerianianusWight is clearly basedonmixedelements,two sheets
atKew are marked"Type" inblack ink. Sheet 1, Herb. Wight Ceylon 1836, consists
C.E.Ridsdale: New taxa for the Rubiaceae ofSri Lanka 459
oftwoelements,theright handelementhaving anadditionallabelwitha description
inpencil. Left 'a' hasa glabrous stem,corolla(immature) withafew scattered hairs;
right 'b' thestemis adpressed hairy, strigose,corollaimmature,with afewscattered
hairs.
Sheet2, Walkers.n.,consists ofleft+ rightspecimen marked 'c'. Stem, sparse-
ly adpressed hairy, stipules 3-4 x 0.5mm, adpressed russethairy, leaves variable
3-7 by 1.5-3cm, midribbelow sparsely pubescent, petiole up to6 mm,sparsely
hairy. Flowers 4-7perleafaxil, corollatube3 mm, sparsely hairy, lobes 1.5mm,
withafewscatterd hairsoutsideattop,anthers 1 mm.
Athird specimen, lowercentre, 'd', stem glabrous, youngbuds, corollanot ob-
served.
KEY TO THE VARIETIES
- Corollaglabrous var. foetulentus
Corollaoutsidepubescent var. pseudovarians
-
var. foetulentus
Distribution— SriLanka:NuwaraEliyaDistrict(16 coll.),apparently commonon
HortonPlains.
var. pseudovarians Ridsdale, var. nov.
A var.foetulento Ridsdale corollis extus pubescentibusdiffert. Typus: Walker s.n.'c'
(K),Sri Lanka.
Distribution—SriLanka,locality unknown,only known fromthetype.
Note—Differsonly inthepubescent corolla.
Lasianthus neolanceolatusRidsdale, nom. nov.
Lasianthus walkerianus var.lanceolata Hook,f.,Fl.Brit.India 3(1880) 186. Syntypes:Macrae
(458)s.n.;Thwaites CP80,340, 1728.Lectotype(mihi):Macrae 458 (K);all Sri Lanka.
Shrub. Branchlets sparsely hairy. Stipules lanceolate, 3-5 x0.5 mm, russet hairy.
Leaves lanceolate,4-15 x 1-3cm, aboveglabrous, below very sparsely hairy, lat-
eral nerves prominently raised aboveandbelow,apex long-acuminate, baseacute,
lateralnerves 3-4(-6) pairs, distinctfromthetertiary nerves. Petioleupto 0.5 cm
long, hairy. Flowers 3 or 4, only known fromimmaturestage. Hypanthium ovoid,
0.5-1 mmlong, glabrous. Calyx campanulate, tube0.7-1 mm,outside glabrous or
with a few scattered hairs, lobestriangular, 0.5-0.7 mmlong, margins with a few
scattered hairs. Corolla tube2-3 mm,outside very sparsely hairy, lobes c. 2 mm
long,outsidewitha few scattered hairs. Fruits c.5 mm.
Distribution— SriLanka: Kandy District (8 coll.), BadullaDistrict(2 coll.),Rat-
napuraDistrict(1 coll.), NuwaraEliyaDistrict (7 col.).
Ecology —Montaneshrubbery.
460 BLUMEA —Vol. 41,No. 2, 1996
Metabolus decipiens (Thwaites) Ridsdale, comb. nov.
AllaeophaniadecipiensThwaites,Enum.PI.Zeyl.(1859) 147;Hook,f.,Fl. Brit.India 3(1880)48;
Trimen,Handb. Fl.Ceylon 2(1894)301; Illus. Fl.Ceylon (1894)t. 48. Metabolus rugosa
var. decipiens (Thwaites)Hochr., Candollea 5 (1934)280;Puff& Igersheim, Bull. Jard. Bot.
Nat.Belg.63 (1994)257. Type:Thwaites CP3093, Sri Lanka.
Allaeophaniadecipiens var. flavescens Thwaites,Enum. PI.Zeyl. (1859) 147; Trimen,Handb. Fl.
Ceylon2(1894)301. Metabolus rugosa var. flavescens(Thwaites)Hochr.,Candollea 5(1934)
279 (inchforma zeylanica, nomilleg.). Type:Thwaites CP 3094, Sri Lanka.
Allaeophaniaarnottii Hook,f.,Fl.Brit.India 3(1880)48. Allaeophaniadecipiensvar. arnottii
(Hook,f.)Trimen,Handb.Fl.Ceylon 2(1894)301. Type: Thwaites CP87,Sri Lanka.
Distribution—Sri Lanka:MataleDistrict(1 coll.), Kandy District(2 coll.), Ba-
dullaDistrict (1 coll.),Ratnapura District(1 coll.),NuwaraEliya District(21 coll.).
Ecology —Montanevegetation inopenandsecondary sites. Common.
Note—Vegetatively very variable, var.flavescens withitssubsessile ovatetoro-
tundateleaves atfirstsight appearstobe distinct,butawholerangeofintermediates
exist.
Oldenlandiaerecta (Manilal & Sivar.) Ridsdale, comb. nov.
Hedyotiserecta Manilal& Sivar., Bot.Notiser 129(1975) 191;Sivarajan & Biju,Taxon 39 (1990)
671. —Type: Sivarajan491 (isoL),Kerala.
Hedyotiscorymbosavarlinearis auct.[nonOldenlandialinearis DC.]; Deb & Dutta,J.Bomb. Nat.
Hist Soc. 83 (1986)692,p.p.
Erectannualherbs.Stemsangular, ridges prominent, glabrousto scabrous.Stipules
2-3 mmlong, pectinate, bristles 2-5. Leaves suberect, linear-lanceolate, 10-40x
4-6mm, margins often recurved, aboveglabrous, below scabrous particularly on
themidrib,apexandbaseacute,lateralnervesinconspicuous. Inflorescenceaxillary
cyme,peduncles 0.5-2cmlong, usually atright angles tothestem.Flowers (1 or)2
or 3, pedicel 2-5 mm long. Hypanthium ovoid, 0.7-1 mm long, calyx tube short,
lobestriangular, 0.7-2mmlong.Corollatube 1.5-2mmlong,outsideglabrous, in-
sidewith aring ofhairsinthethroat,lobestriangular, 1-1.2 mmlong. Stamensin-
sertedjustbelow thecorollathroat, filamentsshort,anthers0.3-0.5mm. Style0.5-
0.7 mm,stigma bifid. Capsule globose, 2.5-3x 1.5-2mm.
Distribution— Sri Lanka(exact locality unknown), continentalSoutheastAsia,
Java.
Ecology —Weedy species ofdrygrasslandsandwaste places.
Note Sivarajan & Biju (1.e.)maintainthattheIndianmaterials they haveexam-
inedare distinct fromtheAfricanmaterialsofOldenlandiacorymbosa var. linearis
(DC.) Verde, whilstDeb& Dutta(1.e.) considerthem tobeconspecific. As thereis
no widespread revisionofthegroupthroughout SoutheastAsiaandMalesiaIhave
nominally recognised O. erecta.
Ophiorrhiza rugosaWall.
Ophiorrhiza rugosa Wall, in Roxb., Fl.Ind. 2(1824)547;Deb & Mondal,Bull. Bot. Surv.India
24 (1982)227. —Type:Wallich Cat.6255,Nepal.
C.E.Ridsdale: New taxa for the Rubiaceae ofSri Lanka 461
KEY TOTHE VARIETIES
la. Stem succulentbelow, procumbent, rooting atnodes var. prostrata
b. Stem woody below 2
2a. Leaves broadly elliptic toobovate, generally less than4cm long
var. decumbens
b. Leaves elliptic, generally over4cmlong 3
3a. Leaves dryingbrownish below var. angustifolia
b. Leaves dryingsilverybelow var. argentea
var. angustifolia (Thwaites) Ridsdale, comb. nov.
OphiorrhizaangustifoliaThwaites,Enum. PI.Zeyl. (1859)140;Alston,Handb.Fl.Ceylon6 (1931)
149. Ophiorrhizamungos var.angustifolia(Thwaites) Hook, f.,Fl.Brit.India 3(1880)77;
Trimen,Handb. Fl.Ceylon 2(1894)321. Type: Thwaites CP 431 (holoPDA; iso K), Sri
Lanka.
Distribution— SriLanka:Kandy District(10 coll.), BadullaDistrict (1 coll.),Ka-
lutaraDistrict(1 coll.), Ratnapura District(3 coll.),NuwaraEliya District(1 coll.).
?PeninsularIndia.
Ecology —Primary andsecondary forestand dampplaces inthewet zoneofSri
Lanka.
Pavettabadullensis Ridsdale,nom. nov.
Pavetta indica var.MontanaThwaites exHook.f.,Fl.Brit.India 3 (1880)150.—Type:Thwaites
(CP2512),Sri Lanka.
Distribution — SriLanka:BadullaDistrict (3 coll.),NuwaraEliyaDistrict (1 col-
lection).
Ecology —Primary andsecondary montane forest.
Notes Hookerbased var. montana on aThwaitescollectionnoting andadopt-
ing amanuscript name; CP2512 is theonly sheet withsuch anannotationandmust
be consideredasthetype.
Bremekamp attempted to createa newspecies forthistaxon and choseCP 2456
as thetypeofhisP. blanda. Ibelievethatthecollectionswithashortcorollatubeare
separatefrom P. blanda.Theepithet P.montana already exists, hencethenewname.
Fruiting materialis difficulttoseparatefromP.blanda.
Psydrax dicoccos Gaertn. var. lanceolatum(Arn.) Ridsdale, comb. nov.
[Webera lanceolatum Moon,Cat. (1824) 19,nom. nud.] Canthium lanceolatum Arn.,Pug.PI.
Ind. Or. (1836)24; NovaActaAcad. Caes.Leop.-Carol.Nat.Cur. 18(1837)342. - Canthium
didymumvar.lanceolatum (Arn.)Thwaites,Enum.PI.Zeyl. (1859) 152;Hook,f.,Fl.Brit.India
3(1880) 132;Trimen,Handb.Fl.Ceylon2(1894)343. Type:SriLanka,notindicated.
Distribution— SriLanka: Kandy District(3 coll.);possibly alsoPeninsularIndia.
Ecology —Rupiculous, sometimesrheophytic.
462 BLUMEA —Vol. 41, No. 2, 1996
Psydrax grandifolia(Thwaites) Ridsdale, comb. nov.
Canthium didymumvar. grandifoliumThwaites,Enum. PI.Zeyl. (1859) 152. Type: Thwaites
CP 3419(holoPDA;K iso), Sri Lanka.
Distribution —Endemic, only known withcertainty fromtwounlocalizedhistori-
calcollections.
Ecology — Unknown.
Psydrax montanum(Thwaites) Ridsdale, comb. nov.
Canthium montanumThwaites,Enum.PI.Zeyl. (1859) 152;Hook,f.,Fl.Brit.India3(1880) 132;
Handb. Fl.Ceylon 2(1894)343.—Plectronia montana(Thwaites)Bedd.,Fl.Sylv. For. Man.
5 (1874)cxxxiv.—Type: Thwaites CP 1685 (holoPDA;iso K), Sri Lanka.
Distribution— SriLanka: Kandy District(3 coll.),BadullaDistrict(1 coll.),Nu-
waraEliya District(3 coll.),Ratnapura District(1 coll.).
Ecology —Upper montaneforest.
Psydrax pergracile (Bourd.)Ridsdale, comb. nov.
Canthium pergracile Bourd.,J.Bomb. Nat. Hist. Soc. 12(1899)352, t. 5.—Type:notindicated
[Bourdillon(K)].
Canthium didymumvar.rostrata Thwaites,Enum.PI.Zeyl.(1859)152.—Type:Thwaites CP2607,
Sri Lanka.
Distribution—SriLanka: Ratnapura District(2coll.);PeninsularIndia.
Ecology —Evergreen forest.
Index
(Synonymsin italics)
Allaeophaniaarnottii460 (Hedyotis)tridentata456 Oldenlandia erecta460
decipiens460 trimenii lessertiana456
var.flavescens460 var. orbicularifolia 457 Ophiorrhizaangustifolia461
var. trimenii 457
Canthiumdidymum mungos
var.grandifolium462 Knoxiaplatycarpa var.angustifolia461
var.lanceolatum 461 var.spicata457 rugosa 460
var.rostrata462 spicata457 var.angustifolia461
lanceolatum461 Lasianthus chrysocaulis457 var.argentea461
montanum462 foetulentus458 var.prostrata461
pergracile462 var.foetulentus 459 var.decumbens 461
Hedyotiscoprosmoides456 var.pseudovarians459 Pavettabadullensis 461
corymbosa neolanceolatus 459 indica var.montana461
varlinearis auct. 460 varians458 Plectronia montana462
walkerianus auct.458
erecta460
walkerianus Psydrax dicoccos
gartmorensis455
lessertiana auct.456 var.lanceolata 459 var.lanceolatum 461
grandifolia462
neolessertiana 456 Metabolus decipiens460
montanum462
obscura rugosa
pergracile462
var.B 456 var.decipiens460
var.minor456 var.flavescens460 Weberalanceolatum 461
