Table Of ContentNEW SPEACNIDEA SN EWR ECORODFE CNOMMUCSL ACHL(ATNR ICHOPTER1A :M ay 1998
ECNOMIDFAREO)MP APUNAE WG UINAEAN D[ RIJAANY A
http.osr:g///d1o0i.2419.90/5j.mmv.1998.57
DAVIIDC. ARTWRIGHT
13B roClrgeas WcaenndtHa,en iagV hitcst3,o2 r1Ai6ua,s tralia
Abstract
Cartw0r.i11g9.h9,Nt8 e,.sw p eacniaden ser we coofErc dn nMmcusl ac(hTlrpaitnce hroa:
EncomifdraPoeam)p NueaGw u ianneldar Jiaa.Myn ea mooiftr hsMe u seoufVim c to5r7i:a
73-87.
Dersicptkieaoynnsadsc ,h ecakrploerivi sdfoter md a loefts w entsyp-eoocfEnic een so mus
inlcudsienvge nnetswep eenfc riPoeamsp NueaGw u ianneldar JiaayTnah .e nAoui:shem
traslpieaEcnci neoslm aursaC kairrati wg1h9t9r,0e ciosfrr odPmea dpN ueaGw u mfeoatr h e
fitrismte .
Introduction potashsyidurmo Mxaildaeens.d f emales can
seldboeam s socpioastietudin vlceeolslysl ected
Thrsepee coifEe csn omhuasvb ee edne scribed
inc opourlb ar etdh rofruogmlh a rvae. Females
previfrooumPs alpyNu eaw G uin(ePaN G) and
havneo bte edne scriintb hepidas p beerc ause
liraJna y(aEc. y cloKpiimcmui1sn9 s6E,2.a ; d di
oft hper obolfep mo siatsisvoec iwahtiicohn,
Malic1k9y9E,3. ;p apuaUnlumse1 r9,3 E8)..
isi ncrewahsemenod rteh aonn es peciises
cycloapniEdc. pu sa puaanrhuees r ree drawn
collfreocmats e idt e.
frotmho er igfiignuarNleot s y.op rne e mwa teria)
Deposifotrso preiceiasmr aeebn ebsvr iaatsed
oft hetsweso p echiaebsse esne eKnu.m ansk1
folloTwhsNe:a tuHriasltM ourys euLmo,n don
(19r7e9c)o trwdmoea dls ep ecifmreoPnmNs G ,
(BMNHB).P; . B ishMoups euHmo,n olulu,
whihceih d entaisEf .cyi celdo pKiicmumsi ns.
Hawa(iBiP BMN)a;tn iaoNla tuHriaslt ory
Mosotft hmea tearviaailfl oasbrtl uewd ays
MuseuSmofi,Ha u,n g(aNrHyM SM)u;s eoufm
colldeucrttiehndI9eg 6 0asnd de posaisd treide d
VictMoerlibao,(u NrnMeY N)a;t iMounsaelu m
pinnsepde ciimnte hnBes.P .Bis hoMpu seumo,fN atuHriasltL oirey(d,Re Mn NHZ)o;o logisch
HonolHualwua(,iB iP BM).
MuseuUmn,i vervsaAinmt setietr( dZaMmA );
At otofa6 l7m alweesre ex amidnuertdih nigs
ZoologMiusscehueUmsn, i veorfsC iotpye n
study, wphliacichne1 d8sw peercoeifw e hshi c
. hagen, C(oZpMeUnCh)ag.e n
17w ernee wto s cieAnlcmeoh.sa ttl hfse p eci
Figusrpeedc iamreien dse ntbiyfit ehde
menasr ree fetrort ewdos pec(iEecsn omus
authnoort'esb nouomkb perre,f CiTx-o ;c
skruismpn .o va.nE d. m ilnesnpns.oi vst. h)e,
casiPoTnn-aulm breerftseot r hn eo teubsoeodk
remai3n7si pnegc irmeepnrse s1e6sn pteicnige s.
byD rA .N ebo(iNsMsV A)bb.r evifoart ions
ThPea pNueaw GuinJeaayt-aol otraf2il 1a n
geniptaaraltrisecs haosiw nFn i Iga sn 2d.
speccioemsp awriettshh A eu strtaoltoiafaln
40s pec(iCeasr twIr9 i9g0ah�nt dw, e �tem
EcnomMucsl achlan
Indonteostaialalosn fo 2 1s pec(iCeasr twnght,
1994). . EcnoMmucsL ac1h8l63a40n:., - Car1t9w9r4i:ght,
Thoejb eivcetosft hsitsu wdeyrt eop rovaid4e4 7.
keayn dde scrifoprtt him�oa nlosef Esc nomus
. T;,sppee cPiheisl.o ptoetnameRulaslm ubsu r,
specfrioemPs a pNueawG utneaanl dn aJna y�.
184(2bo yr igdiensaiiolgn n)a.t
Botmha laensfd e maolfEe csn omsupse cies
arbee sitd eifnitebdyd iffeirneg necneist aRleima�r,Afa r.e vidsieadg nwoaspsir so vbiyded
whiucshu arlelqyuc ilreeastr hiaenb gd o1m1en1C artw(r9 i19)g4.h t
Ketyo m aloefPs a pNueaw G uinaenaldnr ian sJpaeycoaifEne c s1 1011111s
Supieoarp i i
r
J. �.�������.����.���.�.:.��.���.;�pI..�:. ����:� .���.��.��'.�����.:�t:p�-�
s�;�ri·�·�w·i;t·hlpoapurPertgno edc ?aengvs eessn mtarnrl(g �iin3g ,5s. ) . �. ..
.
2()1. Baspaallto efi nferior Wa)patpp heo nmdtda�ogdre ssa (lF i3lg5,o.s) b. .e-'. ...
.
Baspallao tfie ni foearp penwdiatghaepo sou mttd eodrl soa(blFe i 7g9,.s) . 4
r
73
1
74 D. 1. CARTWRIGHT
3(2). Inferior appendages in ventral view dilated, with strongly bifid apices
(Fig.4) E. masalaisp. nov.
Inferior appendages in ventral view with weakly bifid apices (Fig. 6)
E. cyclopicus Kimmins
4(2). Superior appendages in lateral view with apical finger-like projection
(Fig. 7) E. digitulus sp. nov.
Superior appendages in lateral view without apical finger-like projection
(Fig. 9) 5
5(4). Superiorappendages in lateral view short and broadbased, length about 2 *
width(Fig. 9) E. larakiaCartwright
Superior appendages in lateral view long and robust, length > 2.5 x width
(Figs II, 13) 6
6(5). Inferiorappendages in lateralviewdeeply incised(Figs 11, 13, 15) 7
Inferiorappendagesin lateral viewnotdeeplyincised(Figs 17, 19) 9
>6"
7(6). Superior appendages very long and slender, length width (Fig. 1 1)....
E. addi Malicky
Superiorappendagesmorerobust, length<5 y width (Figs 13, 15) 8
8(7). Inferior appendages in ventral with dorsomesal process (Fig. 14)
E. oriomo sp nov.
Inferiorappendages inventralwithoutprocess(Fig. \6)....E. cavatus sp. nov.
9(6). Inferiorappendages in ventral view with mesal digitiform process (Fig. 18),
in lateral view with greatly enlarged basal segment of inferior appendages
(Fig. 17) E. illugisp. nov.
Inferiorappendages in ventral viewwithoutprocess(Fig. 20), basal segment
ofinferiorappendagesnotenlarged(Fig. 19) 10
10(9). Parameres branched (Fig. 19) E. skruim sp. nov.
Parameresnotbranched(Figs21,23,41) 1
1 1(10). Superior appendages in lateral view dilated strongly apically (Figs 21, 23)
12
Superiorappendages in lateral view not obviously dilated or weakly dilated
apically(Figs 25, 27) 13
12(11). Inferior appendages in ventral view slender and fairly straight (Fig. 22)
E. masongsp.nov.
Inferiorappendages in ventral view robust and strongly bent (Fig. 24)
E. tamiok sp. nov.
13(1 1). Parameresin lateral viewdownturnedapically(Figs25, 27) 14
Parameresin lateral view notdownturnedapically(Figs 35, 37,41) 18
14(13). Inferiorappendagesin ventral view,taperedin apicalhalf(Figs26,28) 15
Inferiorappendages in ventral view, not tapered in apical half(Figs 30, 32)
16
15(14). Superior appendages with a swelling on dorsobasal margin (Fig. 25)
E. bullatus sp. nov.
Superior appendages without a swelling on dorsobasal margin (Fig. 27)
E. aliceae sp. nov.
16(14). Inferior appendages in ventral view, tapered strongly to narrow rounded
apices(Fig. 30) E. laensis sp. nov.
Inferiorappendages in ventral view, not tapered strongly, with short pointed
apices (Figs 32, 34) 17
17(16). Inferior appendages in lateral view, tapered in apical half, with pointed
apices(Fig. 31) E. ulap sp. nov.
Inferiorappendages in lateral view, not tapered in apical half, with rounded
apices(Fig. 33) E. wamena sp. nov.
18(13). Superiorappendages in lateral view witha small mesally directed digitiform
projection on ventrobasal margin(Fig. 35) E. milnensissp. nov.
Superiorappendages in lateral view without digitiform projection (Fig. 37)
19
ECNOMUS(TRICHOPTERA) FROM NEW GUINEA 75
19(18). Superior appendages very long and slender, length > 6 * width (Fig. 37)....
E. dadi sp. nov.
Superiorappendagesmore robust, length <5 * width(Figs39,41) 20
20(19). Superiorappendages tapered slightly distally, parameres simple and slender
(Fig.39) E.iomarisp. nov.
Superior appendages not tapered distally, parameres complex (Fig. 41)
E.papuanus Ulmer
Ecnomus spin sp. nov. G.A. Samuelson (genitalia prep. PT-1281 figured,
BPBM).
Figures 1, 2
Description. Male. Wings fawn, venation charac-
Type material. Holotype male, Papua New Guinea, teristic of genus. Ventrolateral processes of
Mamai Plantation, EofPortGlasgow, 150m,lighttrap, segment 10 short, broadbased, with 3 short setae
27 Feb 1965, R. Straatman (genitalia prep. CT-142 apically. Superior appendages in lateral view
figured, BP-8324, BPBM).
long, length about 5 x width (Fig. 3), in ventral
Description. Male. Wings fawn. Ventrolateral view with field of mesally directed spiny setae
processes of segment 10 reduced to inconspicu- apically; inferior appendages in ventral view,
ous rounded swellings each with 3 small setae length about 2.5 * width, apices dilated, shal-
apically. Superiorappendages inlateral view long lowly bifid (Fig. 4), in lateral view robust, length
with a large ventrally directed, digitiform process about 2 x width; basal plate of the inferior
mesally on ventral margin (Fig. 1), in ventral appendages with a large pointed dorsal lobe;
view with a field ofmesally directed spiny setae parameres in lateral view robust, dilated and
apically; inferiorappendages inventral and lateral slightly down turned apically; phallus with apex
views with a subapical dorsally directed digiti- extended into a short projection (Fig. 3).
form projection (Figs 1, 2); parameres straight Female unknown.
and robust in lateral view; phallus acute apically Length offorewing: male4.4-5.7 mm.
(Fig. 1). Etymology. PapuaNewGuineanwordforspiritof
Female unknown.
the waterways, noun in apposition.
Length offorewing: male 3.4 mm.
Distribution. Northeast PapuaNew Guinea.
Etymology. Papua New Guinean word for spear,
referring to the projection on the superior Remarks. Ecnomus masalai groups with E.
appendages, noun in apposition. cyclopicus Kimmins on the basis ofthe pointed
dorsal lobe on the basal plate of the inferior
Distribution. Southeast Papua New Guinea appendages, and can be distinguished by the
(known only fromtype locality). dilated, bifid apices on the inferior appendages,
Remarks. The single male specimen is very which are similarin appearance to the Australian
distinctive and distinguished readily from all species E. turrbalCartwright.
other known Ecnomus species by the very large
Ecnomuscyclopicus Kimmins
process on each superior appendage.
Figures 5, 6
Ecnomusmasalaisp. nov. Ecnomus cyclopicus Kimmins, 1962: 134, figs
Figures 3, 4 35A-C.
GTuyipneea,Wmaatue,ria1l1.50-1H3o0l0otmy,peMalamiaslee,trap,P2a9puDaec 1N96e5w, M(TyBtpMeNCHmyac)tl.eorpisa,l.3H5o0l0otyftp,e Mmaalre,1I9r3i6an. JLa.yEa.(ICnhdoeneessimaa)n,
JK.aSsPeasrdaalmta,ycpe4ek8s:(kgemniEItalKiaaimnaparlneetp,u.,CT1P-3a15p40u8ma,,B7P-NN8eo3vw241,95BG9Pu.iBnTMe.)Ca.., SabPraorna,typceammpale2,, 2Ir0i0a0n Jfat,yaJu(lIn1d9o3n6e,sia()L,.EC.ycClhoepessmMatns),
NMaeaw(Ggueniinteaal,iaMporerpo.bePT-Di1s6t0r8ic,t,BPMBtM)M;iss1imma,le1,30P0apmu,a (BBMMNNHH).staTtyepsetshantotmyseerne,praoldtuhcotuigohnDorfPK.iBmamrinnarsdloaftetrhale
7°13'S, 146°20'E, 15-21 Dec 1966, G.A. Samuelson view drawingmatchesthe holotype specimen prepared
(Ggueinnietaal,iaWaprue,p.MoCTr-o1b2e2,DisBtPriBcMt,);12010maml,e,MaPlaapiusaeNtreapw, oOnthaerslimdaete.rial. 2 males, Papua New Guinea, Tele-
26 Oct 1961, J. Sedlacek (genitalia prep. CT-147, fomin, West Sepic Province, c. 1600 m,25 Jul? 3 Aug
BPBM); 1 male, Papua New Guinea, Umboi Island, 1 1975, P. Beron and/or P. Chapman (NHMS). Material
km N Awelkom, 600 m, light trap, 21 28 Feb 1967, notseen.
CARTWRIGHT
76 D. I.
Figures 1-10. Ecnomus spp. Male genitalia in lateral and ventral views. 1, 2: E. spia sp. nov. 3, 4:
E. masalai sp. nov. 5, 6: E. cyclopicus Kimmins (copied from Kimmins, 1962, Figs 35A-C). 7, 8:
E. digitulus sp. nov. 9, 10: E. larakia Cartwright.
Abbreviations: i.a., inferiorappendages; par, parameres; pha, phallus; plO, ventrolateral processes of
.segment 10; s.a., superior appendages. All scale lines 0.1 mm.
ECNOMUS(TRICHOPTERA) FROM NEW GUINEA 77
Description (revisedafterKimmins, 1962). Male. Etymology. Latin, a little finger, referring to the
Wings, forewing with sparse golden pubescence, projection on the superiorappendages.
brownish membrane with hyaline spots. Ventro-
lateral processes of segment 10 short, digitate, Distribution. Central-west Papua New Guinea.
with several small setae apically. Superior Remarks. Ecnomus digitulus can be separated
appendages in lateral view long, length about 4 * from all otherEcnomus species by the distinctive
width (Fig. 5), in ventral view with field of apices on the superior appendages which are
mesally directed spiny setae apically; inferior downturned, constricted and have a finger-like
appendages in ventral view, robust, slightly projection. The phallus has an unusual embedded
incurved (Fig. 6), weakly bifid in dorsal view process.
(Kimmins, 1962, Fig. 35B), in lateral viewrobust,
narrowed slightly in middle, apices slightly Ecnomuslarakia Cartwright
dilated and truncated; basal plate of the inferior
appendages with a large pointed dorsal lobe; Figures 9, 10
parameres in lateral view robust, dilated and Ecnomuslarakia Cartwright, 1990: 25, figs77-78.
down turned apically; phallus slightly dilated Type material. Holotype male, Australia, Northern
subapically with acute apex (Fig. 5). Territory, Howard Springs, 9 Sep 1980, D. King
Female unknown. (NMV).
Length offorewing: male 5 mm. Other material. 1 male, Papua New Guinea, Morobe
Distribution. Irian Jaya (Indonesia) and western Province, Wau Ecology Institute, 24 May 1986, A.
Papua New Guinea. Wells (genitaliaprep. CT-242 figured,NMV).
Remarks. Ecnomus cyclopicus groups with E. Description, (revised after Cartwright, 1990).
masalai on the basis of having a pointed dorsal Male. Wingspale fawn, venation characteristic of
lobe onthe basal plate ofthe inferior appendages genus. Ventrolateral processes of segment 10
The two species can be separated by the form of very short, with 3 small setae apically. Superior
the inferior appendages, which are truncated and appendages in lateral view short, broad based,
weakly bifid apically in E. cyclopicus. Kimmins tapered strongly distally, length about 2 * width
(1962) figures are reproduced to allow direct (Fig. 9), in ventral view with field of mesally
comparisons and to accompany the description directed spiny setae apically; inferiorappendages
that is revised in light of new interpretations of in ventral view, length about 3 x width, slightly
Ecnomus genitalic structures. constricted medially, dilated subapically, tapered
topointed apices (Fig. 10), in lateral viewrobust,
Ecnomusdigitulus sp. nov. straight, apices rounded; parameres in lateral
view robust, dilated and down turned slightly
Figures 7, 8 apically; phallus slightly dilated subapically with
Type material. Holotype male, Papua New Guinea, acute apex (Fig. 9).
Muller Plateau (Duna Sands, Atem Karanda?), 5 Aug Femaleunknown.
1978, G.S. (genitaliaprep. CT-260figured, RMNH). Length offorewing: male 2.9-3.5 mm.
Description. Male. Wings pale fawn, venation Distribution. Northern Australia and northeast
characteristic ofgenus. Ventrolateral processes of Papua New Guinea.
segment 10 short. Superior appendages in lateral
Remarks. Ecnomus larakia can be distinguished
vcoineswtrliocntge,d barnodadd-obawsnedt,urlneendgtshubaabpoiuctal2ly,x wwiitdhth,a from all other PNG and Irian Jaya species by the
short and broad-based superiorappendages.
finger-like projection apically (Figs 7, 8), in
ventral view with field ofmesally directed spiny
setae apically; inferior appendages in ventral Ecnomus addiMalicky
view, length about 3 * width, tapered to pointed
apices (Fig. 8); in lateral view, length about 4 * Figures 11, 12
width,taperedtopointedapices; parameres in lat- Ecnomusaddi Malicky, 1993: 1120,pi8, 3 figs.
eral view short, straight, dilated apically; phallus Typematerial. Holotypemale,PapuaNewGuinea,Bis-
robust, with ventral subapical projection, acute marck Island, date unknown, Noona Dan Expedition,
apex and embedded slender, elongated, down (ZMK).
turned process (Fig. 7). Paratypes. 8 males, 3 females, samedataasholotype
Female unknown. (ZMK). One male paratype seen and figured (now
Length offorewing: male 6.6 mm. lodged in theNMV).
78 D. I. CARTWRIGHT
Description, (revised after Malicky, 1993). Male. Remarks. Ecnomus oriomo groups with E. addi
Wings fawn. Ventrolateral processes ofsegment and E. cavatus sp. nov. on the basis ofthe incised
10 long, broad based, with 3 small setae apically. form of the inferior appendages, but can be
Superior appendages in lateral view very long, distinguished by the small, truncate dorsomesal
slender, length about 6 * width (Fig. 11), in projection on the inferior appendages and the
ventral view with field ofmesally directed spiny slender, downturned parameres.
setae apically; inferior appendages in ventral
view, length about 3 x width, taperedto inflexed, Ecnomus cavatus sp. nov.
pointed apices (Fig. 12), in lateral view robust,
incised indistal half, apicesupturnedandpointed; Figures 15, 16
parameres in lateral view robust, straight; phallus Type material. Holotype male, Papua New Guinea,
slightlv dilated subapically with acute apex Wau, Morobe District, 1050 m, Malaise trap, 4 Nov
(Fig. 11). 1961, .1. Sedlacek (genitalia prep. CT-149 figured. BP-
Female known (but not described by Malicky, 8324, BPBM).
1993). Description. Male. Wings fawn. Ventrolateral
Length offorewing: male 4mm, female 5 mm. processes ofsegment 1 long, with 3 small setae
Distribution. Northeast Papua New Guinea (type apically. Superior appendages in lateral view
locality only). long, length about 3.5 * width, tapered slightly
distally (Fig.15), in ventral view with field of
Remarks. Ecnomus addi can be distinguished mesally directed spiny setae apically; inferior
from all other PNG and Irian Jaya species by the appendages in ventral view robust, length about
very long and slender superior appendages com- 2.5 x width,taperedto pointedapices (Fig. 16). in
binedwith the incised inferiorappendages, in lat- lateral view incised sub-apically with upturned
eral view. Malicky's (1993) figures are redrawn apices; parameres in lateral view robust, dilated
to allow direct comparisons and to accompany anddownturnedapically; phalluswithacuteapex
the description that is revised in light of new extended into a short projection (Fig. 15).
interpretations ofEcnomus genitalic structures. Female unknown.
Length offorewing: male 5.2 mm.
Ecnomusoriomo sp. now Etynwlog}'. Latin, hollowed out, referring to
appearance ofinferiorappendages.
Figures 13, 14
Distribution. Northeast Papua New Guinea (type
Type material. Holotype male, Papua New Guinea.
locality only).
Western District.OriomoRiver.3 m,8°50'S, 143°1l'E,
light trap, 4 Aug 1964. H. Clissold (genitalia prep. Remarks. Ecnomus cavatus groups with E. addi
CT-121 figured, BP-8324. BPBM). and E. oriomo on the basis of incised inferior
Paratvpe male, same data as holotype. 1 Aug 1964 appendages, but can be distinguished by the
(genitalia prep. CT-131, BPBM).
absence of a mesal projection on the inferior
Description. Male. Wings fawn. Ventrolateral appendages and the robust, downturned
processes ofsegment 10 short, broad based, with parameres.
3 small setaeapically. Superiorappendagesin lat-
eral view long, length about 4 x width (Fig. 13). Ecnomus illugi sp. nov.
in ventral view with field of mesally directed
spiny setae apically; inferior appendages robust, Figures 17, 18
length about 2.5 x width, with small truncate, Type material. Holotype male. Papua New Guinea,
dorsomesal projection near middle, incised New Britain, Gazelle Peninsula, Upper Warangoi.
distally. tapered to pointed apices (Figs 13. 14): Illugi, 220 m. Malaise trap. 15 Dec 1962, J. Sedlacek
parameres in lateral view slender, down turned (genitalia prep. CT-132 figured. BP-8324. BPBM).
apically; phallus slightly dilated subapically with Paratvpe male, samedataasholotype(genitalia prep.
CT-136. BPBM).
acute apex (Fig. 13).
Female unknown. Description. Male. Wings fawn. Ventrolateral
Length offorewing: male 3.8 mm. processes ofsegment 10 long, with 3 small setae
Etymology. Named after type locality (Oriomo apically. Superior appendages in lateral view
River), noun in apposition. long, length about 5 width (Fig. 17), in ventral
view with field of mesally directed spiny setae
Distribution. Southeast Papua New Guinea. apically; inferior appendages in ventral view
ECNOMUS(TRICHOPTERA) FROM NEW GUINEA 79
Figures 11-18. Ecnomus spp. Male genitalia in lateral and ventral views. 11, 12: E. addi Malicky
(reproduced from Malicky, 1993, pi. 8, 3 figs). 13, 14: E. oriomo sp. nov. 15, 16: E. cavatus sp. nov.
mm.
17, 18: E. illugi sp. nov. All scale lines 0.1
HO I). I. CAKTWRKillT
robust, length aboul 1.5 • width, with mesal digi- Length offorewing: male 3.2 4.2 mm.
tiform projection subapically, tapered to pointed Etymology. Papua New (iuinean word meaningto
apices (Fig. IK), in lateral view slender, tapered join (arms), referring to the forked parameres.
slightly iii pointed apices, basal section ofinferior
appendages greatly enlarged, concealing para Distribution. Northeast and southeast Papua New
meres and phallus; parameres in lateral view Guinea,
robust, dilated and down turned apieally; phallus Remarks. Ecnomusskruim is separated from other
with acute apex extended into a short projection PN(i speeies on basis ofthe distinctive, branched
(Fig. 17). parameres. Similar branched parameres also
Female unknown,
Length offorewing: male 4.4 mm. occur in the Sulawesi (Indonesian) species E.
scltik Cartwright, E. tipis Cartwright and E. tang
Etymology. Named after type locality (Illugi), Cartwright.
noun in apposition.
Distribution. Northeast Papua New Guinea (type Ecnomus masong sp. nov.
locality only).
Remarks. Ecnomus illugi separates from other Figures 21, 22
PNG speeies on the basis of the distinctive,
garpepaetnldyageesn,laarngdeddigbialsiaflormsemcetsiaoln proofjectiinofnerioonr UTympbeo!maItselrainad,l aHbooluotty8pekmmalwe,nwPapLuaabNLeabw, G3u0i0nema,,
the inferiorappendages. Malaise trap, 8 ll> Feb 1967, G.A. Samuelson
(genitalia prep. (1-172 figured, BP-8324, BPBM).
Paralvpes: 1 male, similar data to holotype, 600 m,
lighl trap, 2I-2S Feb 1967 (genitalia prep. CT-137,
Ecnomusskruim sp, nov. BPBM); 2 males, Papua New Guinea, New Britain,
Gazelle Peninsula, Upper Warangoi, Illugi, 220 m.
figures I1). 20 CMTal-a1i3s5e,tPraTp-,141359D,eBeP1B%M2),.J. Sedlacek (genitalia preps.
Type material. Holotype male, Papua New Guinea,
Mainai Plantation, V. of Glasgow, 150 in, light trap, Description. Male. Wings fawn. Ventrolateral
27 Feb 1965, R, Straatman (genitalia prep. CT-153 processes ofsegment 10 long, broad based, with
figured, BP-8324, BI'UM). 3 small setae apieally. Superior appendages in
prePpasr,atCoTp-es1:557,m1a5l6e,s,1s6a5-m1e69d,atBaPaBsMh)o;lot2ypmeal(egse,nistaalmiea rlaotwereadl voiveewr lmoindg,dlleensgetchtiaobno,utdi.l3a.5texd wsiudbtahpi,canlalry-
dg1au1t1aa,waaBsPRhBiovMleo)rt,;yp6e°2,45m3'a0Sl,eJsa1.n47Pl°al1>p0(>u'5aE,(gNSeenOiwtma,lGiulaiignphreteapt,sra.pl,CacT1.-31SA4i3pn,r- (dFiirge.cte21d),spiinnyvesnettraaelapviieealwlyw;itihnfeHreiolrdaopfpemnedsaaglelsy
1966, O.K. Wilkes (genitalia preps. CT-123, 173, long and slender, length about 5 x width, tapered
UPHM); male, Papua New Guinea, Wau, Morohe slightly apieally (Figs 21, 22), basal section of
I
District, 1050 m, II Nov 1961, .1. Sedlacek (genitalia inferior appendages with dense group of long
prep. IT-1441. BPBM), hairs nearmid-dorsal margin; parameres in lateral
view robust,dilatedand slightly down turnedsub-
Description. Male. Wings fawn. Ventrolateral
apically; phallus with acute apex extended into a
processes of segment 10 long, broad based,
bilobed, with 3 small setae apieally. Superior long projection (Fig. 21).
Female unknown.
appendages in lateral view robust, length about length of forewing; male 3.3 3.6 mm.
3.5 v width (Fig.19), in ventral view with field of
niesally directed spiny setae apieally; inferior Etymology. Papua New Guinean word for line
appendages in ventral view robust, length about hairs, referring to dense group of long hairs on
3 * width, incised subapically, tapered to rounded basal section ofinferior appendages.
apiees (Fig. 20). in lateral view robust, straight, Distribution. Northeast Papua New Guinea.
length about 2.5 x width; parameres in lateral
view branched, with dorsal branch long and Remarks. I'.cnomus masong separates from other
slender, down curved apieally, ventral branch PN(i species on basis ofthe distinctive group of
shorter, slightly down turned apieally; phallus long hairs on basal section ofinferior appendages
with acute apex extended into a short projection and long slender inferior appendages. 1 have not
(Fig. 19), noted this group of hairs on any other Ecnomus
Female unknown speeies from Indonesia. PNG or Australia.
ECNOMUS(TRICHOPTERA) FROM NEW GUINEA
81
Ecnomus tamiok sp. nov. inferior appendages in ventral view short, length
Figures 23, 24 about 3 x width, broad based, narrowed at about
the middle, tapered to pointedapices (Fig. 26), in
Type material. Holotype maie, Papua New Guinea, lateral view short, straight, length about 3 x
Morobe District, Ulap, 800-1100 m, Sep 1968, width, tapered distally; parameres in lateral view
W.A. Steffan and Y.M.Huang (genitalia prep. CT-124 robust, dilatedanddownturnedsubapically; phal-
figured, BP-8324, BPBM).
p1r4e.Pp4asr.katmCyTp-eWs1:27L1,aem,BaPl2eB8,M-)3s0;amOe1ctmdaatl1ae9,6a5s,PahWpou.laoAt.yNpeSetwe(fgfGeaunniitnaaelnaid,a eluxstFewenimdtaehldeveinunttnrokanlaosswuhnob.ratpipcraoljescwteilolnin(gFiagn.d25a)c.uteapex
Y.M.Huang (genitalia prep. CT-139, BPBM); 1 male, Length offorewing: male 3.2-3.8 mm.
P1a99p0u,aNWe.wF.GuiHnuema,phMraedyasng,(g5e°ni1t6a'Sl,ia145p°re4p5.'E,C2T4-2M6a1y, Etymology. Latin, swelling, referring to superior
RMNH). appendages.
Description. Male. Wings fawn. Ventrolateral Distribution. Southeast Papua New Guinea.
processes ofsegment 10 long, broad based, with Remarks. Ecnomus bullatus separates from other
3smallsetaeapically. Superiorappendages in lat- PNG species on the distinctive superior
eral view long, dilated strongly apically, length appendages with the swelling on the dorsobasal
about 2.5 x maximum width (Fig. 23), in ventral margin.
view with field of mesally directed spiny setae
apically; inferior appendages relatively hairy,
in ventral view robust, length about 2 * width, Ecnomusaliceae sp. nov.
strongly inflexed at about the middle, tapered to Figures 27, 28
pointed apices (Fig. 24), in lateral view tapered
distally, length about 2 x width; parameres in lat- Type material. Holotype male, Papua New Guinea,
eral viewrobust, dilatedslightlyanddown curved Central Province, Eilogo River, near Sergeri, 9°27'S,
apically; phallus with acute apex extended into a C1T47-°24207'Ef,ignuerte,d,21T-M1a67y981,98N6M,VA)..Wells(genitalia prep.
shortprojection (Fig. 23). Paratype male, Papua New Guinea, Mamai
Female unknown. Plantation, E ofPortGlasgow, 150 m, 30Jan 1965, R.
Length offorewing: male 3.0-3.4 mm. Straatman (genitaliaprep. CT-140, BPBM).
Etymology. Papua New Guinean word for axe, Description. Male. Wings pale fawn, venation
referringto shape ofsuperiorappendages. characteristic ofgenus. Ventrolateral processes of
Distribution. Northeast Papua New Guinea. segment 10 broad based, with 3 small setae api-
cally. Superiorappendages in lateral view robust,
Remarks. Ecnomus tamiok separates from other length about 3.5 x width (Fig. 27), inventralview
PNG species on basis of the distinctive dilated with fieldofmesallydirectedspinysetaeapically;
apices on the superior appendages and the inferiorappendages in ventral view long, slender,
strongly inflexed inferiorappendages. length about 4 x width, narrowed or incised at
about the middle, tapered to pointed apices (Fig.
Ecnomus bullatus sp. nov. 28), in lateral view long, straight, length about
Figures 25, 26 4 x width; parameresin lateral view long, slender,
slightly dilated and down turned apically; phallus
Type material. Holotype male, Papua New Guinea, slender, with acute apex extended into a short
Mamai Plantation,EofPortGlasgow, 150m,lighttrap, projection (Fig. 27).
27 Feb 1965, R. Straatman (genitalia prep. CT-154 Female unknown.
figured, BP-8324, BPBM). Length offorewing: male 3.2-3.4 mm.
Paratypes: 1 male, same data as holotype, 27 Jan
1965 (genitalia prep. CT-152, BPBM); 1 male, same Etymology. The species is named for Alice Wells
data as holotype, 30 Jan 1965 (genitalia prep. CT-141, (collector).
BPBM).
Distribution. Central and southeast Papua New
Description. Male. Wings fawn. Ventrolateral
Guinea.
processes ofsegment 10 long, with 3 small setae
apically. Superior appendages in lateral view Remarks. Ecnomus aliceae separates from other
long, length about 4 x width, with a swelling on PNG species on the combination of the straight
dorsobasal margin (Fig. 25), in ventral view with superior appendages and the slender inferior
field of mesally directed spiny setae apically; appendages.
82 D. I. CARTWRIGHT
Figures 19-26. Ecnomus spp. Male genitalia in lateral and ventral views. 19, 20: E. skruitn sp. nov. 21,
22: E. masongsp. nov. (with basal segment ofinferior appendage showing hairs, lateral view). 23, 24:
E. tamiok sp. nov. (with inferior appendage showing hairs, lateral and ventral views). 25, 26:
E. bullatus sp. nov. All scale lines 0.1 mm.
