Table Of ContentGenus Vol. 20(3): 435-484 Wrocław, 15 X 2009
New records of Asian and Australopapuan tortoise beetles
(Coleoptera: Chrysomelidae: Cassidinae)
Lech Borowiec
Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Sienkiewicza 21,
50-335 Wrocław, Poland, e-mail: [email protected]
ABstrAct. New records of 173 Cassidinae species from Asia, New Guinea, and Australia
are given (Australia 6, Bangladesh 1, Burma 13, Cambodia 6, China 41, India 25, Indonesia 51,
Japan 6, Laos 49, Malaysia 19, Mongolia 2, Nepal 9, Papua New Guinea 10, Philippines 11, Sri
Lanka 3, Taiwan 6, Thailand 44, Vietnam 10). Cassida langeri (Myanmar), C. ictericiformis
(NE India), and C. isarogensis (Philippines) are described as new to science.
Key words: entomology, taxonomy, zoogeography, new records, new synonyms, new species,
Coleoptera, Chrysomelidae, Cassidinae, E Palaearctic Region, Oriental Region, Austra-
lian Region, Pauan Subregion.
INTRODUCTION
The paper is my sixth contribution to the knowledge of Cassidinae from Asia and
Australopapuan area (Borowiec 1985, 1990, 1996, 2001; Borowiec and tAkizAwA 1991).
New faunistic records are presented for 173 Cassidinae species (Australia 6, Bangladesh
1, Burma 13, Cambodia 6, China 41, India 25, Indonesia 51, Japan 6, Laos 49, Malaysia
19, Mongolia 2, Nepal 9, Papua New Guinea 10, Philippines 11, Sri Lanka 3, Taiwan 6,
Thailand 44, Vietnam 10). Three species: Cassida langeri (Myanmar), C. ictericiformis
(NE India), and C. isarogensis (Philippines) are described as new to science.
The nomenclature used follows my world catalogue (Borowiec 1999).
The material was obtained from the following collections (with abbreviations
used in the text):
ACO – Agriculture Canada (Ottawa, Canada);
AW – Andrzej WarchałoWski, Wrocław, Poland;
CMNC – Canadian Museum of Nature, Ottawa (L. LesAge);
436 LECh BOROWIEC
DBET – Department of Biodiversity and Evolutionary Taxonomy, Wrocław, Poland
(L. Borowiec);
FK – Frantisek kAntner, Lipi, Czech Republic;
IRSN - Institut Royal des Sciences Naturelles, Bruxelles, Belgium (M. cLudts);
JB – Jan Bezdek, Brno, Czech Republic;
JC – J. cooter;
Jh – Jiri hAvA, Prague, Czech Republic;
MBLSC – Monte L. Bean Life Science Museum, Provo, USA (S.M. cLArk);
MCSNM – Museo Civico di Storia Naturale, Milano, Italy (C. LeonArdi);
MD – Mauro dAccordi, Verona, Italy;
ML – coll. M. LAnger, Lichtenwalde, Germany;
MNM - Muséums nature de Montréal (Stéphane Le tirAnt);
MO – Michal oudA, Plasy, Czech Republic;
MS – Miroslav snižek, Ceske Budejovice, Czech Republic;
MTD – Museum für Tierkunde, Dresden;
NRS – Naturhistoriska Riksmuseet Stockholm (B. vikLund);
SMNS – Staatliches Museum für Tierkunde, Stuttgart, Germany (W. schAwALLer);
Uh – Uwe heinig, Berlin, Germany;
WG – Wolfram guidetti, Leipzig, Germany.
LIST OF SPECIES
1. Aspidimorpha adhaerens (Weber, 1801)
AUSTRALIA: Queensland, 3-15 mi Cairns, 3-4 XI 1972, 1, h.E. Evans (MBLSC);
Queensland, Port Douglas, 9-11 XI 1972, 1, h.E. evAns (MBLSC); Queensland, Yep-
poon, 3-6 II 1970, 1, h.E. Evans (MBLSC).
INDONESIA: Irian Jaya, Manokwari, Gn. Meja, 200 m, 30 XII 2000-1 I 2001,
1, A. Riedel (SMNS).
PAPUA NEW GUINEA: Gusika, 1944, 1, W.L. howe (MBLSC); Morobe Prov.,
Timini, 200-400 m, 7 IV 1998, 3, A. Riedel (SMNS).
Distribution: N Australia, Papuean Subregion, eastern part of Sunda Is. (Amboina,
Buru, N Sulawesi).
2. Aspidimorpha amabilis boheman, 1854
INDONESIA: W Java, Ujung Kulon N.P., 0-50 m, 20 IV-26 VII 1996, 1, S. Jakl
(MS); Sumba, East prov., Luku Melolo Nat. Park, 200-400 m, 7-9 II 2001, 1, S. Jakl
(MS).
Distribution: Indonesia: Java, Lombok, Sulawezi, Sumatra, Sumba, Sumbawa;
Philippines: Luzon, Mindoro, Palawan.
NEW RECORDS OF ASIAN AND AUSTRALOPAPUAN TORTOISE BEETLES 437
3. Aspidimorpha assimilis boheman, 1854
INDONESIA: W Java, Bogor reg., Mt. Salak, 800-1200 m, 22-25 VIII 1997, 1, S.
Jakl (MS); Mentawai Islands, S. Siberut Isl., 0-100 m, II 2005, 11, 50-100 m, VI 2005,
145, X 2005, 1, XII 2005, 4, S. Jakl (MS, DBET); W Sumatra, harau Valley, 500-800
m, III-IV 2005, 1, S. Jakl (MS); Sumatra, harau Valley, 20 km N Payakumbuh, 750 m,
II 2004, 1, S. Jakl (MS); W Sumatra, Mt. Singgalang, Annai Valley N.R., 500 m, IV
2005, 2, V 2005, 2, 15 X 2005, 3, 17 VI 2005, 4, IV 2006, 2, S. Jakl (MS)
MALAYSIA: Bukit Berapit, W of Kuala Kangsar, 20-24 III 2001, 2, A. Kudrna
(MO); Negeri Pulau Pinang, Penang, 27 I 1981, 1, T.-E. Leiler (NRS); Pahang, Tioman
Isl., Kampong Tekek-K Juara, 2.48 N 104.11 E, 7-25 II 2000, 1, E. Strba (FK).
ThAILAND: Betong, Gunung Cang dun vill., Yala distr., 25 III-22 IV 1993, 1,
J. horák (FK).
Distribution: Malay Peninsula, Sunda Is., and Thailand.
4. Aspidimorpha aurata (montrouzier, 1855)
INDONESIA: Irian Jaya, Japen Is., E Seraui, Wawuti, 100-300 m, 17 XII 2000,
2, A. Riedel (SMNS); Irian Jaya, Japen Is., W Seraui, Panduamin, 50 m, 18 XII 2000,
1, A. Riedel (SMNS).
PAPUA NEW GUINEA: Madang Prov., Canopy mission, Baileta, 8 VI 1993, 1, O.
Missa (IRSN); Morobe Prov., Aseki, 1200-1400 m, 14 IV 1998, 3, A. Riedel (SMNS)
; Morobe Prov., 10 km S Garaina, Saureri, 1550-1800 m, 23-27 III 1998, 1, A. Riedel
(SMNS); Morobe Prov., Lakekamu, Tekadu, 300-400 m, 28 II-6 III 1998, 2, A. Riedel
(SMNS); Morobe Prov., W Pindiu, 100-1400 m, 23 IV 1998, 1, A. Riedel (SMNS);
Sandaun Prov., N Mianmin, 700-1100 m, 20 V 1998, 3, A. Riedel (SMNS).
Distribution: Papuan Subregion and New Caledonia.
5. Aspidimorpha australasiae (boisduval, 1835)
INDONESIA: Irian Jaya, Japen Is., Serui, Mantembu, 100-500 m, 16 XII 2000,
2, A. Riedel (SMNS); Irian Jaya, Jayawijaya Prov., Samboka, 200 m, 10-14 X 1996,
2, A. Riedel (SMNS); Irian Jaya, Jayawijaya Prov., Yalmabi, 1200-1400 m, 8 X 1996,
1, A. Riedel (SMNS); Irian Jaya, Wandamen Bay, Wondiwoi Mts., Wasior, 250-600
m, 4 I 2001, 1, A. Riedel (SMNS); Irian Jaya, Yapen Serui, 01.52.11 S/136.14.18 E,
22 XII 2006-9 I 2007, 3, S. Bilý (MS); Sorong Prov., Kec. Waigeo Sel., Gamang Is.,
Yembeser, 0-100 m, 16 XI 1996, 2, A. Riedel (SMNS).
PAPUA NEW GUINEA: Madang, Uho Village, 20 km SW Madang, 250 m, 29 I
2000, 1, R.S. Anderson (CMNC); Morobe Prov., Wau, Mt. Kaindi, 1400 m, 8 III 1998,
6, 20 III 1998, 1, 1 IV 1998, 1, A. Riedel (SMNS); E New Britain, Bainings Mts., Raun-
sepna, 28 VII-11 VIII 1999, 1, C. Mitparingi (CMNC); Western highland Prov., Wulum
Walne, Mt. hagen, 1535 m, 25 II 1997, 1, A. Michalczyk & D. Matzke (ML).
Distribution: New Guinea, Solomon Is., and N Australia.
438 LECh BOROWIEC
6. Aspidimorpha bataviana Świętojańska, 2001
INDONESIA: C Java, Blora env., XI 1997, 31, S. Jakl (MS, DBET); W Java, Ja-
karta-Bekasi, 20 m, 8 IX 1995, 13, S. Jakl (MS, DBET); Sulawesi, Mamasa, IV 2002,
2, S. Jakl (MS, DBET); Sumbawa, IV 2002, 7, S. Jakl (MS, DBET).
Distribution: Indonesia: Bali, Borneo, and Java.
7. Aspidimorpha bilobata boheman, 1856
PhILIPPINES: Luzon, Aurora, Sierra Madre, 15 VI 2007, 7, local collector (MNM,
DBET).
Distribution: endemic to Philippines: Luzon.
8. Aspidimorpha castaneipennis spaeth, 1912
INDONESIA: W Sumatra, Mt. Singgalang, Annai Valley N.R., 500 m, IV 2005,
1, 17 VI 2005, 1, IV 2006, 3, S. Jakl (MS, DBET).
MALAYSIA: Pahang, 30 km NE Raub, Lata Lambik, 300 m, 4 V 2002, 1, Jendek
& Šauša (MS); Pahang distr., 30 km NE Raub, Lata Lembik, 3°56’N/101°38’E, 200-
400 m, 22 IV-15 V 2002, 1, E. Jendek & O. Šauša (JB).
Distribution: Borneo, Nias, Sumatra, Luzon; single localities in Malay Peninsula
and Vietnam.
9. Aspidimorpha chandrika Maulik, 1918
ThAILAND: Nan Prov., Ban Bo Klua env., 13-26 V 2002, 1, Obořil & Prudek
(MS).
Distribution: widespread in continental part of the Oriental Region, from Uttar
Pradesh in India to Vietnam; in insular part of the region known only from Sumatra.
10. Aspidimorpha deusta (Fabricius, 1775)
AUSTRALIA: Queensland, Cairns, 22 II 1997, 10, h. & A. howden (CMNC).
INDONESIA: C Java, Blora env., XI 1997, 2, S. Jakl (MS); W Java, Jakarta-
Bekasi, 20 m, VIII-IX 1995, 5, S. Jakl (MS); Sumatra austr., Kotaagung, Lamuran,
Maudi Valley, 28 II 2000, 3, P. Moric (JB); Wetar Is., Anpala env., 0-20 m, 14 I 2007,
8, S. Jakl (MS, LB).
Distribution: N Australia, Papuan Subregion, Sunda Is., and Philippines.
11. Aspidimorpha difformis (Motschulsky, 1860)
ChINA: Gansu, Lazikou vall., 34.08 N/103.54 E, 27 VI 2005, 1, hajek, Kral &
Ruzicka (MS); W hubei, Muyuping env., 31.45 N/110.4 E, 1100 m, 17 VI 2002, 2,
NEW RECORDS OF ASIAN AND AUSTRALOPAPUAN TORTOISE BEETLES 439
J. Turna (MD); hunan, 30 km N Yanjiajie from Dayong, 27 V 2005, 1, 28 V 2003, 2,
Jenis (MS).
Distribution: China: Fujian, Guizhou, hebei, Manchuria, Shaanxi, Sichuan,
Zhejiang; Japan: Awashima, hokkaido, honshu, Kyushu, Okinawa, Shikoku; Korea;
Russian Far East; Taiwan.
12. Aspidimorpha dorsata (Fabricius, 1787)
ChINA: Tibet, Cha-Yu, 21-28 VII 2004, 1, Li Jingke (ACO).
INDONESIA: W Java, Mt. Salak, 1000 m, 16-22 VIII 1997, 1, S. Jakl (MS).
LAOS: Khammouan pr., 17°43’N/105°09E, 500-600 m, 22 V-8 VI 2001, 1, E. Jen-
dek & O. Šauša (JB); Kham Mouan prov., Nakai vill., 70 km NNE Muang Khammouan,
580 m, 7-25 V 2002, 1, M. Strba (FK); Vientiane Prov., Ban Vangheua, Phou Khao
Khouhay Nat. Park, 55 km N Vientiane, 1000 m, 4-18 V 2000, 1, P. Kresl (FK).
ThAILAND: Chiang Dao prov., Cave, 18-20 V 2004, 1, Z. hanc (MS); Chiang Mai,
Dot Pui, 1 V 1985, 1, S. Steinke (SMNS); Nan Prov., Doi Phuka Nat. Park, 28 IV-12
V 2002, 1, Obořil & Prudek (MS); Sakhon Nakhon Prov., Phu Pan N.P., 17°05’76.7’’
N/103°59’90.7’’ E, 7 VI 2008, 1, T. Ihle (ML).
Distribution: widespread in the Oriental Region, from Ceylon to Sulawesi.
13. Aspidimorpha dulcicula boheman, 1862
INDONESIA: W Sumatra, Mt. Singgalang, Annai Valley N.R., 500 m, IV 2005,
1, V 2006, 1, S. Jakl (MS); W Sumatra, Guguk, VIII 1991, 1, Widagdo (ML).
Distribution: Indonesia: Sumatra, Borneo; Malaysia: Malacca, Sabah, Sarawak;
Philippines: Luzon, Sembilan.
14. Aspidimorpha elevata (Fabricius, 1801)
INDONESIA: Mentawai Islands, S. Siberut Isl., 50-100 m, VI 2005, 1, S. Jakl
(MS); W Sumatra, harau Valley, 700 m, VI-VII 2004, 2, III-IV 2005, 1, S. Jakl (MS);
Sumatra, Lampung Krui: Sukanegara, 9 VIII 2001, 11, K. Smets (IRSN, DBET); Su-
matra, Lmlah Anai, 15 III 1973, 2, local collector (MNM).
ThAILAND: Betong, Gunung Cang dun vill., Yala distr., 25 III-22 IV 1993, 1 J.
horák (FK).
Distribution: W Sunda Is., Thailand, Malay Peninsula, and Philippines: Luzon.
15. Aspidimorpha furcata (thunberg, 1789)
BURMA (MYANMAR): Rangoon-Taikkyi, Nyaunggon, 19-31 V 2001, 1, M.
Klicha (UA).
CAMBODIA: Battambang Prov., Prek Toal, 6 VII 2005, 1, I. Var (IRSN).
ChINA: hongkong, Lantau Island, Ngong Ping env., Polin Monastery to North
County Park, 450-500 m, 14 VIII 2004, 1, Ružička & Vilimová (MS).
440 LECh BOROWIEC
INDIA: Goa Prov., 30 km S Margao, 18.00.47 N 74.01.58 E, 0-20 m, 12-14 VIII
2002, 2, P. Šipek & M. Fikáček (JB); Karnataka, Sulya, Medikeri, 12.32 N 75.29 E,
150 m, 24 IV 2005, 1, M. halada (MS); Madhya Pradesh, 120 km SE Bhopal, Matkuli
env., 22°35’N/78°28’E, 400 m, 28 VI 2006, 1, Z. Kejval (JB); Maharashtra Prov., 120
km NE Mumbai, Igatpuri env., 19.42.17 N 73.33.05 E, 600 m, 1-11 VIII 2002, 3, P.
Šipek & M. Fikáček (JB).
INDONESIA: W Sumatra, Mt. Sanggul, 1250 m, V 2004, 2, S. Jakl (MS); W
Sumatra, Padang Panjang, 10 IX 1991, 1, W. Guidettei (ML).
LAOS: Bolikhamsai Prov., Ban Nape env., 18.20 N/105.08 E, 400 m, 7-16 V
2004, 1, Jendek & Sausa (FK); Bolkhamsai Prov., Pakkading env., 18.18 N 103.59 E,
20-24 XI 2000, 1, E. Jendek & P. Pacholatko (MS); hua Phan Prov., Ban Saluei, Phu
Phan Mt. env., 20.13 N 103.59 E, 1300-2000 m, 6-18 V 2004, 1, F. & L. Kantner (FK);
Khammouan pr., 17°43’N/105°09E, 500-600 m, 22 V-8 VI 2001, 1, E. Jendek & O.
Šauša (JB); Khamouan Prov., Ban Khoun Ngeun env., 18.07 N 104.29 E, 250 m, 4-16 XI
2000, 1, E. Jendek & P. Pacholatko (MS); Khamouan Prov., Nakai env., 17.43 N 105.09
E, 550 m, 22 V-8 VI 2001, 3, Sausa (MS); Lao Pako env., 35 km NE Vientiane, 18.10
N/102.52 E, 50 m, 31 V-4 VI 2004, 1, Jendek & Sausa (FK); Phongsali Prov., Phongsali
env., 21.41-2 N 102.06-8 E, 1500 m, 28 V-20 VI 2003, 2, C. holzchuh (FK); Vientiane
Prov., Lao Pako, 35 km NE Vientiane, 200 m, 1-4 V 2005, 1, P. Kresl (FK).
TAIWAN: Meishan env., Kaohsiung, 100 km NW Taitung City, 23.15.9 N/120.49.5
E, 950 m, 12 VI 2008, 3, F. & L. Kantner (FK); Pingdong County, 8-15 km NE hen-
gchung, 22.05.4 N/120.49.5 E, 120 m, 16-20 VI 2008, 3, F. & L. Kantner (FK); Yilan
County, 20 km N Yilan City, 24.48 N/121.45 E, 120 m, 16-20 VI 2008, 4, F. & L.
Kantner (FK).
ThAILAND: Chumphon Prov., Pha To env., 1-11 V 1998, 2, P. Prudek & R. Šigut
(RS); Doi Phu Kha Nat. Park, 50 km NE Nan, 28 VI-1 VII 1997, 1, J. Rejsek (SMNS);
Mae hong Son Prov., Pai, 16-24 VII 1995, 3, R. Šigut (RS); Nakhon Si Thammarai
Prov., Khao Luang N.P., Krung Chiu waterfall, 13-22 V 1998, 1, P. Prudek & R. Šigut
(RS); Nan Prov., Ban huay Kon, 27 V-10 VI 2002, 1, Obořil & Prudek (MS) ; Ranon
Prov., Andaman Peace Resort, N Laem Son N.P., 09°37’03.8’’ N/098°27’94.2’’ E, 500
m, 5-6 VV 2008, 1, M. Langer (ML).
VIETNAM: Long Binh Army Post, 8 IX 1972, 1, h.J. hArLAn (MBLSC).
Distribution: one of the most common Oriental species, occuring from Ceylon to
Java, Borneo and Palawan, north to S China.
16. Aspidimorpha fuscopunctata boheman, 1854
ChINA: Tibet, Cha-Yu, 21-28 VII 2004, 2, Li Jingke (ACO).
INDIA: Assam, Bhalunpong, 27.02/92.35, 150 m, 1 VI 2006, 1, Pacholatko (MS);
Karnataka, 40 km SW Shimoga, 13.36.74 N/75.07.98 E, 600 m, 10 V 2005, 1, M.
halada (MS).
INDONESIA: W Java, Bogor reg., Mt. Salak, 800-1200 m, 22-25 VIII 1997, 1, S.
Jakl (MS); SE Sulawesi, Buton Isl., South Lasalimu reg., VII 2005, 1, native collector
(MS); W Sumatra, harau Vall., 500-800 m, III-IV 2005, 1, S. Jakl (MS); W Sumatra,
NEW RECORDS OF ASIAN AND AUSTRALOPAPUAN TORTOISE BEETLES 441
Lembeh Anai Vall., Singgalang, 13 IX 1991, 1, Guidetti & Widagdo (ML), 500 m, IV
2005, 1, S. Jakl (MS); W Sumatra, harau Vall., 20 km N Payakumbuh, 750 m, II 2004,
3, S. Jakl (MS); W Sumatra, Mt. Sanggul, 1250 m, V 2004, 1, S. Jakl (MS); W Sumatra,
Mt. Singgalang, Annai Valley N.R., 500 m, IV 2005, 1, IV 2006, 2, S. Jakl (MS).
LAOS: Bolikhamsai Prov., Ban Nape env., 18.20 N/105.08 E, 400 m, 7-16 V 2004,
1, Jendek & Sausa (FK); Khammouan pr., 17°43’N/105°09E, 500-600 m, 22 V-8 VI
2001, 2, E. Jendek & O. Šauša (JB).
MALAYSIA: Kampung Ulu Dong, 25 km NE Raub, 26-31 III 2001, 1, A. Kudrna
Jr. (MO); Perak, 25 km NE Ipoh, Banjaran Titi Wangsa Mts., Korbu Mt., 2100 m, 4-13
III 1998, 1, P. Cechovsky (UA).
PhILIPPINES: Laguna, Los Banos, 22 XI 1953, 1, h.M. & D. Townes
(CMNC).
ThAILAND: Betong, Gunung Cang dun vill., Yala distr., 25 III-22 IV 1993, 1,
J. Strnad (UA), 1 J. horák (FK); Chumphon Prov., Pha To env., 1-11 V 1998, 1, P.
Prudek & R. Šigut (RS); Nakhon Si Thammarai Prov., Khao Luang N.P., Krung Chiu
waterfall, 13-22 V 1998, 2, P. Prudek & R. Šigut (RS); Nan Prov., Ban Bo Klua env.,
13-26 V 2002, 1, Obořil & Prudek (MS); Nan Prov., Doi Phuka Nat. Park, 28 IV-12 V
2002, 1, Obořil & Prudek (MS).
VIETNAM: Bachma, 7 III 1997, 2, V. Sesna (MS).
Distribution: widespread in almost whole Oriental Region, incl. Philippines, north
to S China, east to Sulawesi.
17. Aspidimorpha hexaspilota Baly, 1863
ThAILAND: Nan Prov., Ban Bo Klua env., 13-26 V 2002, 1, P. Prudek & M.
Obořil (JB); Nan Prov., Doi Phuka Nat. Park, 28 IV-12 V 2002, 2, P. Prudek & M.
Obořil (FK, JB).
Distribution: Cambodia, Laos, and Thailand.
18. Aspidimorpha indica boheman, 1854
ChINA: Yunnan, Tengchong, 24°59.4’N/98°28.9E, 1650 m, 5 VI 2007, 1, J. hájek
& J. Růžička (JB).
INDONESIA: W Sumatra, Padang Panjang, Singelang, 450 m, 15 II 1995, 1, A.
Michalczyk (ML); W Sumatra, Solok area, Mt. Talang, 1200-1500 m, III 2006, 2, S.
Jakl (MS).
LAOS: Bollkhamsai Prov., Ban Nape-Kaew Nua pass, 600 m, 18.22.3 N 105.09.1
E, 18 IV-1 V 1998, 1, E. Jendek & O. Sausa (FK); Phongsali Prov., Phongsali env.,
21.41-2 N 102.06-8 E, 1500 m, 28 V-20 VI 2003, 5, C. holzchuh (FK, MS).
MALAYSIA: Cameron highl., Pahang Prov., Ringlet env., 20-23 XI 2000,6, F. &
L. Kantner (FK, DBET).
NEPAL: himalaya, near Mt. Everest, 2900 m, 1-20 IX 1999, 1 (MS).
Distribution: common in continental part of the Oriental Region, north to S China;
in insular part of the region only on Sumatra and Palawan.
442 LECh BOROWIEC
19. Aspidimorpha lobata boheman, 1854
INDIA: Goa Prov., 30 km S Margao, 18.00.47 N 74.01.58 E, 0-20 m, 12-14 VIII
2002, 1, P. Šipek & M. Fikáček (JB).
Distribution: little known, by many years it was treated as synonym of A. sanctaecru-
cis (F.). Confirmed localities known from Ceylon, S and C India, and Bangladesh.
20. Aspidimorpha maffinbayensis boroWiec, 1992
INDONESIA: Irian Jaya, Japen Is., E Serui, Wawuti, 100-300 m, 17 XII 2000, 3,
A. Riedel (SMNS).
Distribution: recently described from the northern coast of New Guinea.
21. Aspidimorpha malaccana spaeth, 1937
MALAYSIA: Pahang Distr., Kampung Kualaboh, 04.27.9 N 101.34.8 E, 850-1050
m, 26 III-3 VI 2001, 1, P. Somody (FK).
ThAILAND: Khao Sok Nat. Park, 8.52 N/98.28 E, VII-VIII 2007, 1, Smoleń
(DBET).
Distribution: Burma: Tenasserim, Thailand, and Malay Peninsula.
22. Aspidimorpha miliaris (Fabricius, 1775)
BURMA (MYANMAR): Yangoon Pr., Airport hotel, Season of Yangoon, 16°51’
N/96°07’ E, 22 VI 2008, 1, M. Langer (ML); Yangon Prov., International Airport hotel
vic., 96.07 E/16.51 N, 21 V 2006, 3, Langer, Naumann & Löffler (ML).
CAMBODIA: Phnom Pen, 15 V 1974, 4 (MNM).
ChINA: hainan, Mt. Wuzhishan, 1-19 VI 2007, 15, local collector (MNM, DBET);
Tibet, Cha-Yu, 21-28 VII 2004, 2, Li Jingke (ACO); Yunnan, Lv-Chun County, Mt.
huanglianshan, 2-8 VI 2008, 7, local collector (MNM).
INDIA: Karnataka, 20 km SE Sagar, 14.06.37 N/75.08.93 E, 600 m, 12 V 2005,
1, M. halada (MS); Tamil Nadu, Madurai, 5 XII 1998, 1, S.C. Bonta (MBLSC); Tamil
Nadu, 25 mls. SE Theni, 26-29 XI 1998, 1, S.C. BontA (MBLSC).
INDONESIA: Bali, Bedugui reg., Tamblingan Lakes N. R., XI 2004, 1, S. Jakl
(MS); 1200 m, Bali, Soka Beach, 27 V 2000, 2, M.A. hielkema (RB); C Java, Blora
env., XI 1997, 2, S. Jakl (MS); Flores, Gunung Kelimutu, 2 II 2002, 1, M. hoffmann
& R. Richter (ML); SE Sulawesi, Buton Is., South Lasalimu reg., VII 2005, 1, native
collector (MS); Sulawesi, Mamasa, IV 2002, 2, S. Jakl (MS); S Sulawesi, Puncak,
Palopo env., V 2000, 1, Benes & Secky (MS); S Sulawesi, 20 km NE Sabbang, 2.28.56
S 120.12.00 E, 400 m, 5-7 VII 2001, 1, Bolm (SMNS); Sumatra, Lampung Krui: Su-
kanegara, 9 VIII 2001, 5, K. Smets (IRSN); Sumatra, Lmlah Anai, 15 III 1973, 1, local
collector (MNM); W Sumatra, Padang Panjang, 16 IX 1991, 1, W. Guidettei (ML), 3
III 1995, 1, U. Arnold (ML); Sumbawa, IV 2002, 1, S. Jakl (MS).
NEW RECORDS OF ASIAN AND AUSTRALOPAPUAN TORTOISE BEETLES 443
LAOS: Bollkhamsai Prov., Ban Nape-Kaew Nua pass, 600 m, 18.22.3 N 105.09.1
E, 18 IV-1 V 1998, 1, E. Jendek & O. Sausa (FK); Khammouan pr., 17°43’N/105°09E,
500-600 m, 22 V-8 VI 2001, 1, E. Jendek & O. Šauša (JB); Phongsali Prov., Phongsali
env., 21.41-2 N 102.06-8 E, 1500 m, 28 V-20 VI 2003, 2, C. holzchuh (FK).
PhILIPPINES: Laguna, Los Banos, 13 XII 1953, 1, h.M. & D. Townes (CMNC);
Luzon, Aurora, Sierra Madre, 15 VI 2007, 9, local collector (MNM, DBET); Mindoro
occ., San Jose, 9 IV 1953, 1, h. Townes (CMNC); Mountain Province, Benaue, 31 XII
1953, 1, h.M. & D. Townes (CMNC).
ThAILAND: Chiang Mai, Doi Pui, 3 V 1985, 1, 4 VI 1985, 1, 15 V 1985, 1, S.
Steinke (SMNS); Loei, Dan Sai, 11 V 2001, 3, Constant & Grootaert (IRSN); Prachuap
Khiri Khan, Sam Roi Yot, 22 III 2001, 4, P. Grootaert (IRSN); Sakhon Nakhon Prov.,
Phu Pan N.P., 17°05’76.7’’ N/103°59’90.7’’ E, 7 VI 2008, 1, T. Ihle (ML).
Distribution: the most common and widespread species, known from the whole
Oriental Region and New Guinea.
23. Aspidimorpha mutilata boheman, 1854
INDONESIA: Bali, Beduqui reg., Tambilingan Lakes N.R., 1200-1300 m, VII
2004, 27, XI 2004, 6, I-II 2005, 6, V 2005, 5, S. Jakl (FK, MS, DBET); W Java, Puncak
pass, 17 km of Cipanas, 1250-1600 m, 8-10 X, 1, Bolm (MS).
Distribution: Indonesia: Bali, Java, Lombok, Sumatra; Malaysia: Pahang.
24. Aspidimorpha novaeguineensis (boisduval, 1835)
INDONESIA: Irian Jaya, Jayawijaya Prov., Samboka, 200 m, 10-14 X 1996, 5, A.
Riedel (SMNS); Irian Jaya, Timika, VI 2001, 1 (MS).
PAPUA NEW GUINEA: Morobe Prov., Tekadu, 300-400 m, 28 II-6 III 1998, 1,
A. Riedel (SMNS).
Distribution: New Guinea and adjacents islands.
25. Aspidimorpha pacalis spaeth, 1926
INDONESIA: Tanimbar Isl., S. Yamdena Isl., 21 km NE Saumlaki, Mam’s vill.,
150 m, 27 XI-11 XII 2005, 2, J. horák (MS, DBET); Tanimbar Isl., S. Yamdena Isl.,
Lorulun Vill., 20 km NE Saumlaki, Mam’s vill., 150 m, XII 2006, 1, Obořil & Bourna,
15 XII 2006-10 I 2007, 3, 10 I-5 II 2007, 6, S. Jakl (MS, DBET).
Distribution: hitherto known only from holotype of Tanimbar Island.
26. Aspidimorpha punctum (Fabricius, 1801)
INDONESIA: Irian Jaya, Jayawijaya Prov., Samboka, 200 m, 10-14 X 1996, 4,
A. Riedel (SMNS); Irian Jaya, Jayawijaya Prov., Yalmabi-Samboka, 200-650 m, 9 X
1996, 1, A. Riedel (SMNS); Irian Jaya, Waigeo Is., Urbinasopen, 0-150 m, 24 I 2001,
444 LECh BOROWIEC
8, A. Riedel (SMNS); Irian Jaya, Yapen Serui, 01.52.11 S/136.14.18 E, 22 XII 2006-9
I 2007, 3, S. BiLý (MS); Maluku, Seram, 12 km SE Wahai, Solea, 31 X-4 XI 1998, 1,
J. horák (FK); Seram, 35 km E Pasahari, 24-30 X 1998, 1, J. horák (FK).
PAPUA NEW GUINEA: Morobe Prov., Lakekamu, Tekadu, 300-400 m, 28 II-6
III 1998, 4, A. Riedel (SMNS); Popondetta, VIII 1946, 1, Lippert (MBLSC).
Distribution: New Guinea and N Australia: Queensland.
27. Aspidimorpha sanctaecrucis (Fabricius, 1792)
BURMA (MYANMAR): Shan state, nr hsipaw, 22.37 N/97.15 E, 500 m, 4 VII
2006, 1, M. homburg (Uh); Shan State, northern Inie Lake, Khaung Daing hotel gar-
den, 900 m, 14-15 II 2005, 4, S. Löffler (ML); Yangoon Pr., Airport hotel, Season of
Yangoon, 16°51’ N/96°07’ E, 22 VI 2008, 2, 4 VII 2008, 1, M. Langer (ML).
CAMBODIA: Phnom Pen, 15 V 1974, 1 (MNM).
ChINA: hainan, Mt. Wuzhishan, 1-19 VI 2007, 15, local collector (MNM, DBET);
Tibet, Cha-Yu, 21-28 VII 2004, 3, Li Jingke (ACO).
INDIA: Assam, Bhalunpong, 27.02/92.35, 150 m, 1 VI 2006, 2, Pacholatko
(MS).
INDONESIA: W Sumatra, harau Vall., 700 m, VI-VII 2004, 1, S. Jakl (MS).
LAOS: Bollkhamsai Prov., Ban Nape-Kaew Nua pass, 600 m, 18.22.3 N 105.09.1
E, 18 IV-1 V 1998, 1, E. Jendek & O. Sausa (FK); Khammouan pr., 17°43’N/105°09E,
500-600 m, 22 V-8 VI 2001, 1, E. Jendek & O. Šauša (JB); Khamouan Prov., Nakai env.,
17.43 N 105.09 E, 550 m, 22 V-8 VI 2001, 3, Sausa (MS); Phongsali Prov., Phongsali
env., 21.41-2 N 102.06-8 E, 1500 m, 28 V-20 VI 2003, 1, C. holzchuh (FK).
ThAILAND: Chiang Mai prov., Doi Suthep, 4 VI 2004, 1, Bily (MS); 10 km
W Kalabin, 20 IX 2002, 1, native collector (ML); Khao Sok Nat. Park. env., 9-11 IV
2001, 1, A. Kudrna Jr. (MO); Rakam env., 80 km N Chanthaburi, 22-24 V 2003, 1, O.
Safranek (MS); Sakhon Nakhon, Phu Pan Nat. Park, 4-18 VII 2002, 1 (ML) ; Sakhon
Nakhon Prov., Phu Pan N.P., 17°05’76.7’’ N/103°59’90.7’’ E, 7 VI 2008, 10, T. Ihle
(ML); Soppong, Pai, 16 IV-1 V 1997, 1, M. Zyka (UA).
VIETNAM: Tuyen Quang Prov., Na hang Reserve, 300 m, 16-24 V 1997, 1, S.
Peck (CMNC).
Distribution: widespread in continental part of the Oriental Region, north to S
China; in insular part of the region rare, only in Sumatra, Java and Borneo.
28. Aspidimorpha sarawacensis spaeth, 1904
MALAYSIA: Pahang, Cameron highlands, peak of Gunong Jasar, NW of Tanah
Rata, 7 II 1985, 1, J. Settele (ML); Pahang, Cameron highlands, Tanah Rata env.,
1500-1700 m, 1-13 II 2003, 1, M. Obořil (FK).
Distribution: Borneo and Malaysia: Pahang.
