Table Of ContentTThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan
uetwh 1)p6-toGa 45(3):I57-168O,ctober1994
New and little-known clearwing moths from Central Asia
(LepidopterSaes,iidae)
OlegG.GoRBuNOV'
Institut eof Evolutionary Morphology and Ecolegy of Animals, Russian Academy of Sciences,
Leninsky prospect 33, 117071, Moscow V-71, Russia
Abstract Two new Centra ]Asian species of the genus Bembecia are describe dand figure ;d
B. martensi sp. nov. (Kirghizs tAalna,shtau Mts), and B. 2onsteini sp. nov. (Kirghizstan,
Sarykamy･ sMhts) . New data of the distributio nof S,nanthedo ntosevslei igpaten k19a87, are
presentecl .A redescription of Bembecia tvagneri <Ptingel e1r9,12) coupled with notes on its
bionomics and habita tis given.
Key words Lepidopter aS,esiida eS,},nanthe dtoonsevsk iBiembeeia wagnen', B. maitensi n. sp.,
B. gonsteini n. sp., Centra lAsia, Kazakhstan, Kirghizstan, taxonomy.
The montainous areas of Central Asia (th eTian-Shang, the Pamirs-Alai ,the Pamirs) have
long been known as one of the richest in the entire Palearctic with respect to the fauna of
Sesiidae w,ith lot sof endemic taxa involved ,At the present, however, this region supports
only about 40 species, this being evidence of the very poor state of the part . To fi1 t1he gap
at least partly, this contribution puts on record only some results obtained in 1993 during
an internation aexlpedition to the Tian-Shang Mts of Kazakhstan and Kirghizsta nC,entral
Asia,headedby Dr S.Golovatch(Moscow,Russia) Prof. Dr J.Martens (Mainz,
and
Germany). Below,Idescribe two new, and redescribe one poorly-known, species of the
genus Be7ebeci aI'IUbner ,[181 9c]oupled with new data on the distributio nof Synanthedon
tosevsle iSipatenka ,1987. Material referred to in this article, includin gthe types, is
deposite din the followin gcollections i
ZISP-The Zoological Institut eR,ussian Academy ef Sciences ,St, Petersburg, Russia.
ZMMU-The ZoologicalMuseum, Moscow StateUniversityM,oscow, Russia.
ZMHB-Zoologisches Museum der Humboldt-Universitat, Berlin,Germany.
EMEM-Entomologisches Museum Dr U.EitschbergerM,arktleuthen, Germany,
CG-collection of the author, Moscow, Russia.
CL-collection of Z. Lagtdv kBarn,o, Bohemia.
CS-collection of K. Spatenka, Prague, Bohemia.
CT-collection I,Togevski,Novi Beograd,Yugoslavia.
of
Sgnantheden tosevskiivSpatenka,1987 (Fig23.)
gpatenka,
Synanthed otnosevski i 1987. ZL ArbGem. dst .Ent. 39:22, fig. 4.Type locality:Central Asia,
Uzbekistan ,Bolsho iTchimgan Mt. Holotype male, in CS.
Material examined. 1 0 (paraty pUeS)S,R, Uzbekistan ,Bolshoi Tchimgan, 90km NE
gpatenka
Tashkent, ca. 1,80 0m, 20. VI. 1986, leg .K. (CG) .4 6I, Kirghizstan, Sarykamysh
Mts, 2,OO Om, 41e55 'N, 74`03 'E, 24-26 .VII, 1993, leg .O, Gorbunov (CG ,CS).
Remarks. This species was describe don 15 males collected with a synthetic pheromone
* Visitin Sgcholar of Zoologica lLaboratory ,Faculty of Agriculture, Meijo University,
Tempaku-ku, Nagoya, 468Japan
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158 OlegG.GoRBuNOV
in three areas of Uzbekistan in the environs of Bolshoi Tchimgan Mt. (Bolsh oTichimgan
Mt., Burchmulla and Kara-Mazar). The exact host plant of this species is unknown, but
vSpatenka (198 7su)ggested that it may be a Stzg ispx. The above additional small series of
this species was taken with a synthetic pheromone developed by Dr Priesne r(Max-Planck-
InStitut ,Seewiesen ,Germany) for European populations of Synanthed aondnrenacfonne
(Laspeyre 1s8,01) and S. spuler i(Fuch s1,908) (Nr 10) on a dry and sunny slope supporting
a xerothermic vegetation. This slope is cut into small gorges with some shrubs, viz.
5birae sap,, Cotoneast sepr., Berberi ssp., Rosa sp., and Cerasu ssp. (Fig.23 )S.omewhat
Iower down the slope, Skel dsrp. and PoPzalzt ssp. flan kriver banks. Inspectio enf the shrubs
and trees in search of sesiid larval activity has been totally unsuccessful. However, {n my
opinion, the host plant of the species concerned might be one of the above shrubs. Because
of the directio nfrom which the wind blew, sesiids could fly toward the pheromone traps
only from the slope side.
Bembeciamartensi sp. nov. (Fig s1-5, 14-16)
Holotype .?,Kirghizst Aalna,shtau Mts,Ermendy,41015, N, 72"40 'E, 1,55e m, 25-26. V.
1
2
Figs1-4. Male genitalia of Bembecia martensl sp. nov., paratype (genita lpriecparation
No.93-13). 1) tegumen-uncus complex ; 2) valva; 3) saccus; 4> aedeagus.
Scale bar : O.5 mm.
Fig.5, Female genitalia of Bembecla martensz sp. nov., paratype (genita lpriepcaratien
No.94-03) .Scare bar: O.5mm.
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Clearwing Moths from Centra lAsia 159
1993, ex larva, leg ,O.Gorbunov (ZISP )P,aratypes ,1 di 2 \, same localit aynd date ,ex
larva,leg.0.Gorbunov (CG).
Description. \, ho]otype (Fi g1.4) . Alar expanse 18, Omm ; body Iength 9. 0mm ; forewing
7.8 mm ; antenna 4.2 mm.
Head: antenna dorsally black with greenish sheen, ventrally at apical quarter black ,at
basal 3/4 brown with admixture of individual yellow scales ; labial palpus yellow-orange
with a narrow black stripe both internall yand externally on joint 1s and 2 ; frons dark
yellow with a large grey-brown to black spot medially ; vertex black with individual
yellow-orange scales ; pericephalic hairs dorsall yblack ,laterall yyellow-orange.
Thorax : patagiurn black with blue-viole tsheen dorsall yand yellow-orange laterally;
tegula black with blue-viol estheen, with a few yellow-orange scales apically and with a
black blue-violet black
yellow-orange axillar spot ; mesothorax with sheen ; metathorax
with two small yellow-orange spots and two tufts of yellow-orange hair-lik escales;
pleura of thorax black with violet sheen with individua ylellow-orange scales.
Legs : fore coxa black with violet sheen on interna lhalf and yellow-orange on external
half; hind tibia orange-yellow with a narrow black ring with violet sheen near base of
apical spurs ; spurs orange-yellow,
Abdomen : black with bright greenish-viole tsheen ; dista lhalf of tergite s2 and 4 yellow-
orange ; tergite 6 with a few yellow-orange scales distal l; ysternites 2 and 4 each with a
narrow yellow stripe distall ; yanal tuft orange with a narrow black stripe laterally.
Forewing : costal margin, Cu-stem and narrow outer margin of apical area black with
greenish sheen with admixture of individual orange-yellow scales ; anal margin, veins Mi
and M2 as well as proximal part of apical area orange-yellow to orange with a few black
scales; disca lspot black with violet sheen, with a small orange-yellow spot distally;
transparent areas poorly developed ,covered with semitransparent scales with golden hue ;
posterior transparent area absent ; external transparent area small, oval, divide dinto
three cells, about as long as disca lspot ; cilia dark grey to black with bronzed sheen.
Hindwing: transparent but covered with hyaline scales with golden hue (basal lwyith
orange scales) ; veins and. disca slpot black ; outer margin black with brenzed-viol eshteen,
about twice as narrow as cilia; disca lspot narrow, reaching base of common M3-Cui
stem ; cilia dark grey to brown, caudally orange.
di (Fig 1.5) .Alar expanse 17.0 mm ; body length 10.0 mm ; forewing 7.5 mm ; antenna 5.7
mm,
Head: antenna black with greenish sheen, with a broad yellow stripe at 4/5 from base
externally, with a yellow apical tuft ; labia lpalpus yellow to yellow-orange with a broad
black stripe ventro-externally ; frons dark yellow with a large black to dark brown spot
medially ; vertex black with a few yellow scales ; pericephalic hairs orange.
Thorax : patagium black with bronzed-green sheen dorsally and orange-yellow laterall y;
tegula black with greenish sheen, with an orange-yellow tip and an orange--yellow axillar
spot ; meso- and metathorax black with greenish sheen ; beside sthat, tegula ,meso- and
metathorax densel ycovered with pale yellow hair-like scales ; pleura of thorax black with
greenish sheen, with a small orange-yellow spot.
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Legs: fore coxa black with a narrow orange-yellow stripe externally, densel ycovered
with pale yellow hair-lik secales ; hind tibia orange-yellow with a broad black ring both
basally and near base of apical spurs ; spurs orange-yellow.
Abdomen : blac kwith blue-green isshehen ; tergites 2 and 4 each with a narrow orange-
yellew stripe distall ; ytergite 7 in dista lhalf orange-yellow ; sternites 2, 4 and 7 with a
narrow orange-yellow stripe distal (lQyn tergite 2 broader ); anal tuft orange-yellow mixed
with black scales with blu esheen.
Forewing: black individual Cu-stem
costal margin with admixture of yellow scales;
yellow-orange mixed with black scales ; anal margin orange ; apical area and veins within
external transparent area yellow-orange; narrow outer margin of apical area black;
disca lspot black with a narrow, yellow-orange, dista lmargin; transparent areas well-
developed ,densel ycovered with colourless scales with bluis hhue ; external transparent
area elongate, divide dinto four cells, about 2.5 times as long as discal spot ; cilia brown-
grey with bronzed sheen.
Hindwing:transparent;veins discal black
and spot mixed with yellow scales;outer
margin narrowly black ; disca lspot narrow, reaching base of vein Mz ; cilia brown-grey
with bronzed sheen.
Variabilit y.Coloration virtually invariable ,slight variation concerning size only (Fig 1,6),
Male genitalia (preparati oNne,93-13) ,Tegumen-uncus complex (Fig. 1)relatively
broad; lateralis broad,
scopula androconialis well-developed; crista gnathi relatively
long ,subtriangular ; crista gnathi medialis undeveloped ; valva (Fig 2.) trapeziform oval ;
Iong distally
crista sacculi simple, and narrow, proximal part with pointed setae, with a
group of lappet-Iike ,fiat-tepped setae ; saccus (Fig 3.) with a flat base, about 1.5 times as
long as vinculum ; aedeagus (Fi g4). relatively thin, somewhat longer than valva ; vesica
with rows of small cornuti.
Female genitalia (preparat iNoon.94-03) (Fig 5.) , Eighth tergite relatively broad; apo-
long bursae
physis posteriori sas as apophysis anterioris; ostium narrow, membranous;
long bursae
antrum about as as apophysis anterioris, membranous ; corpus ovoid, membra-
nous, without signa.
gpatenka
Diagnosis .Closes tperhaps to B 2ebo et Gorbunov, 1992 (Fig s18-19) ,but can be
distinguish efdrom it by the yellow-orange coloration of scales on the head, thorax, wings
and abdomen (lemo nyellow in B. zebo), by the orange-yellow hind tibia (lemo nyellow in
the species compared>, and by the somewhat smaller size as a whole. From B. zuvandica
Gorbunov, 1987, this new species differ isn the orange-yellow or yellow-orange scales both
on the thorax and abdomen. From B. zonsteini sp. nov,, B. martensi sp. nov. is clearly
distinguish abbyl tehe coloration of the forewing ,legs and abdomen,
Biongmics. The host isOnobnyychis (FabaceaeT)h.e larvalivesinside
plant cho7ussanica
the root for ene year, and it makes there a tunnel about 3-6cm long . Sometimes,
especially in a thin roQt, larva lactivity promotes a gall-lik egrowth of the root. Before
pupation, the caterpillar makes an exit tunnel 2-7 cm long up to the ground level a,nd then
it makes a cobweb cocoon at the base of this tunnel. In the laboratory conditions, larvae
were pupated in the middle of Jul yand appeared in the beginning of August.
Habitat . The larvae of the type series were collected in the roots of the host plant
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CrcarwingMoths from CentralAsia 161
growing on a small but steep slope of a hil lwith a soft sandy seil. It was very diMcult to
find because hills floodlands
similarly open slopes everywhere around all and were covered
with wallnut-fruit forest s.In smarl clearings in this forest I, collected also ChamaesPhecia
mtftilata (Staudin g1e88r7,) moths as well as Bembecia tshi"agana (Sheljuzh k1o9,35) larvae.
Etymolegy. With great pleasure I name this beautifu lspecies after my frien dProf. Dr
J .Martens (Mainz G,emnany), who contributed to my travel to Kirghizsta nand Kazakhstan
in1993.
Bembeciaxonsteini sp. nov. (Fig s69, 17)
Holotype.di,Central Asia,Kirghizstan,Sarykamysh Mts,2,OOO 41"55'N, 74"03'E,24-
m,
26.VII,1993,leg.O.Gorbunov(CG).
Description,6n, holotype (Fig 1,7).Alarexpanse 18. 0mm ; body length 11.2 mm ; fore-
wing 8.5 mm ; antenna 5.0 mm.
Head: antenna black with greenish sheen, with a few pale yellow scales externally and
subapically; labia lpalpus yellow to pale yellow with a narrow black stripe externally on
joint 1s and 2 ; frons pale yellow with a few grey scales lateral l; yvertex black rnixed with
pale yellow hair-like scales ; pericephalic hair slemon-ye]lew,
Thorax: patagium dorsall yblack with bronzed sheen, with a few pale yellow scales
basally ,laterall ylemon-yellow ; tegula black with greenish sheen, with a lemon-yellow tip
and a large lemon-yellow axillar spot ; meso- and metathorax black with greenish sheen ;
beside sthat, tegula, meso- and metathorax densel ycovered with long ,hair-lik ep,ale
yellow scales; pleura of thorax black with green{sh sheen, with a small yellow spot
,:ny.,..Itg" "'g.vt/tttt
''/ttt
JA･･
'r.'t/t/t/t.t''tttt.tttttt//tt.tttt'nt.tt-t.tt"'t..el.8t{.t't.'t/.tttt.tt/'tttttll..t/.tt't'tttttt.t.4'itttttt/tt
7
9
Figs6-9. Male genitalia of Bembecia zonstetm sp. holotype (genita lpriecparation
No.93-14). 6) tegumen-uncus complex ;n7o) v.va,lva; 8} saccus; 9) aedeagus.
Scale bar : O.5 mm.
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cranially,
Legs : fore coxa black with a broad yellow stripe externally, densel ycovered with long,
pale yellow hair-lik secales ; hind tibia yellow with a broad black ring both basall yand
near base of apical spurs; spurs yellow.
Abdomen : black with bright greenis hsheen ; 2nd tergite with an extremely narrow yellow
stripe distall y4;th ,6th and 7th tergite scompletely yellow; 4th sternite yellow, 5-7th
sternites with admixture of individu aylellow scales ; anal tuft yellow-orange medially and
blacklaterally.
Forewing: costal margin black mixed with yellow scales ; Cu-stem and space between
veins M3-Cu2 yellow with a few black scales ; anal margin, posterior transparent area and
apical area, except for narrow black outer margin, yellow ; disca lspot black with narrow,
yellow, dista lmargin; transparent areas poorly developed ,covered with coiourless and
yellow scales with golden hue ; external transparent area very small, divide dinto three
cells, about twice as short as disca lspot ; cilia grey-brown with bronzed sheen.
Hindwing : transparent ; veins dark brown to black ; outer margin dark brown to black,
broad, about as broad as cilia ; disca lspot black with a few yellow scales, broad, triangu-
lar, base M3-Cui bronzed
reaching of common stem ; cilia grey-brown with sheen.
Female. Unknown.
Variability. Unknown,
Male (preparatioNno.93-l4).Tegumen-uncus (Fig.6)
genitalia complex relatively
bread;scopula androconialis well-developed;crista gnathi laterali s trapeziform,
(Fig,7)
relativery narrow and short;crista gnathi medialis very narrow;valva
trapeziform oval ; crista sacculi double ; dorsa lpart narrow, long, with rows of pointed
setae proximally ; ventral part oval, broader and shorter than dorsa lpart, bein gconnected
with it by a row of lappet-lik ef,lat-topp esdetae ; saccus (Fig 8,) with small broadening
basall yand a slightly rounded base ,about 1,5 times as long as vinculum ; aedeagus (Fi g9.)
long
as as valva ; vesica with rows of small cornuti.
Diagnosis ,Closes tto B. 2ebo vSpatenka et Gorbunov, 1992 (Fig s18-19) ,but it can be
distinguish ebdy the coloration of the forewing ,thorax and abdomen. From B. martensi
sp. nov., B. zonst'eini sp, nov, differ sclearly in the yellow or lemon-yellow scales on the
head, thorax, abdomen and legs .Besides that, the new species is easily distinguishab blye
the structure of the male genitalia ,especially by the form of crista gnathi andlor crista
sacculi.
Bionomics : The host plant is unknown. The holotype was taken in the end of Jul ywith
a synthetic pheromone developed by Dr Priesne r(Max-Planck-Inst iSteuetw,iesen ,Ger-
many) for European populations of Synanthed aondnrenaof)rme (Laspeyr e18s0,1) and S,
spzalen('Fuchs1,908)(Nrle),
Habitat. Dry, warm and sunny slopes supporting a xerothermic vegetation (Fig 2.3).
Etymology: I am pleased to name this new species after my frien dDr S. Zonstein
(Bishke kK,irghizstan), another participant of the 1993 expedition to Central Asia,
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ClearwingMothsfromCentralAsia 163
11
13
Figs10-13. Male genitalia of Bembecin tvagneri (Pttnge l1e9r1,2) (genita lpriecparatron
No,93-16) .10) tegumen-uncus complex;11) valva;12) saccus;13) aedoeagus.
Scale bar : O.5 mm.
Bembecia u7agneri (PUngel e19r1,2) (Fig s10-13, 20-22)
Dipsosphec iua'clgneri Pttngele r1,912 ,in Seitz ,Gross-Schmet tE.ide 2 : 395 , Type localit y: Ccntra lAsia,
II iRiver district .Lectotype, [P, in ZMHB {designati obny Capuge, 1973) ; Capuge ,1971 : 247, fig 4., pl.
gpatenka
I ,figsG-H ; and Lagtavka, 1988 :337.
Bembecia ts .stn) wagnen':Capuge, 1973: 143;gpatenka et al., 1993: 101.
Material examined, 1di (lectoty pe"1/)18",, "Cotype", "Pr, gen Nr. 3103 I.Capuse"
(ZMHB) ; 1 9 (paralectoty p"Ilei-G)ebi,et 1911", "Type" (ZMHB) ; 35 di" ,Centra lAsia,
Kazakhstan, Il iRiver, 50 km upstream of Bakanas, 4, VI. 1993, leg .O. Gorbunov (ZISP,
ZMMU, EMEM, CG, CL, CS, CT) ; 31 c{T 2 ?, Centra Alsia, Kazakhstan, Temirlik River,
43"18'N, 79=12'E,24-25,VI,1993,leg,O,Gorbunov (ZISPE,MEM, CG, CL, CS,CT).
Diagnosis ." (Fig 2.1) ,Alar expanse 20.0-27 .mm0 ; body length 12,O-16 .mm0 ; forewing
10.0-12 .m5m ; antenna 7.0-8. m5m.
Head: black brightbluish labial black;frons
antenna with sheen; palpus grey-brown
with a white stripe Iaterall yv;ertex black with admixture of silvery hair-lik escales;
hairsblack,
pericephalic
Thorax : dorsall yblack with bluis hsheen, covered with silvery hair-lik secales ; pleura of
thorax black with bluis hsheen, with a small white spot anteriorly.
Legs: fore coxa black ,covered with silvery hair-lik escales ; hind tibia brick-orange;
black.
spurs
Abdomen : completely black with bright greenish-blue sheen ; anal tuft black with green-
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164 ()Ieg(i{/}ORIILINOV
Figs14 -16 .Eigmbecia ewartens!l sp.nvv. I, t5L,holDt} ,pAlea.rexpanse 18.(Pmiv t.I5, ",
parat,ype , AIar expanse 17.0 inm. I5. 1+'re.c tehniieyrged female tparatype).
Fig ,17. Ben･thE' (:･.on'siteain!/ gspp. aTitov,e, i,fi', ihkelaot},p e.Alu. rexpEn)se 'IlS'.Rf )di"jtin.kistari,
Figs }8-l9, Bentbeci ax･ebo et Cjerbuuov .1992. 18 .- ,t, Turkestttn
IV'It KsL,im}.)e lPttss ,10, VT. 1'9I9'1u, rtk･xe isntia'nva ,It.g .O. (.}orhunoL '(CG). t'X]nr c,xpansc 2.,4,t')
mm. 19. ll,'1'ac]Mkistan, ?v'I tKsu,nibel Pass. 10, V'I ,IV91. ex larx' al,eg.
CZ).Gorbunov (CG) ,.'ig .elxp2anns'..e 21,5]']Ln].
ish-blue laterally.
sheen ee.ntrally iind orari..T.e
'f]'om
Fc)rewEn g: costal margin black with g't'eeni sshheen ; anal rnargin bnse to disca lspet
brick-orang e; apical area narrow proximally, a few scales at disca lspot distal} ay$ well
as veins Mi and N'1 w2ithin external transparent area brick-orange ; transparent areas well-
developed ,covered with light ,brownish semitransparent scales ; external transparent area
divide dinto fiv ･ceells, seTne"rhat les sthan t"･,i caes brc}a das disca lspQt.
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ClearwingMoths tro:nCentralAsia 165
20
'`yeUexV'
Figs2e-'2I2' c･mRicr"l7ibkec'i aza'dagneri {I'timge Te1r9,12) 20 tin , forni ,KAzakhstan,
Rixpr, 43'lg 'N, 79'12 'E, 24 25. Vg. 1993, teg .O.Gorbunev (C(' llA)lar
expnnse 24.e rnn] 2i. cs","b]ack" form, KAiakhstan, Teniirlik River, ･t 3IS' N, 79'
iZ' E, 24-ZJ VI. !(.}Y 3l,eg .'Or.cG,umiibruTni{k."･ fCe). :Xlat expanse 2150innL 22. if.
"veiluw" forni ,Kuiakhstan River 4:'IS 'N, 79'12 'E, 24 25, VI 1993, ieg
O, Gorbunov {CG) . tllar expanse 2T', Omm.
glpatenka,
Fig. 2: 1 II:]bitat osf Si'nanthetf" jtiosE:,sk ii IY87 and Benibc･ ica ..:cnisf stp, iunovi
Kirg]'tirstt mS,ar} lcan)y,h Mts, 2,(M) Om. 41 [}5 'N. 7S03' E.
Figg 24 25. Elflbltat est' Bc'nsbt,c iLat./a,gnen' (PUngeler l,912) 24. Kazakhstan, 11 iRivur, ,!}e
km upstream of ISakanas. Lt5 ,Kazakhstaii, Teinirlik Feiver ,43'IS 'N, 79'IL 'E.
Hindwing: transparent; veins. and disca lspot b}ack; outer margin about twice as narrew
as cilili, black ,basall yorlinge i disca lspet trtangultt ri,eaching base ef vehis peI 3Cui ; cilia
black,
9 (FIg2.2), Alar Z6.0-27.0 body lengthi･1.0-15.5 ferewingl2.0-12.5
expEinse rniv ; n)fn ;
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mm ; antenna 6.5-7. m0m,
Head: antenna black ,covered with sparse whitish scales ; labia lpalpus orange-yellow;
frons pale yollow ; vertex orange ; pericephalic hairs pale yellow.
Thorax : patagium black with bluish sheen dorsall yand yellow lateral l; ytegula yellow
with admixture of individua blrown to black scales posteriorly; meso- and metathorax
black bluish
with sheen.
Legs : fore coxa brown to black with a few orange and yellow scales at outer margin ; hind
tibia orange-yellow ; spurs erange-yellow.
Abdomen : dorsall yblack with brigh tgreenish-blue sheen ; 4th tergite yellow, 2nd, 5th and
6th tergites yellow in dista l2/3, 3rd tergite with a narrow yellow stripe medially ; ventrally
yellow ; anal tuft yellow with black scales medially at base.
Forewing : costal margin black with bluis hsheen ; anal margin, proximal half of apical
area, veins within external transparent area and dista lthird of disca lspot orange-yellow ;
posterior transparent area very small, anterior and external transparent areas covered
with brownish semitransparent scales ; external transparent area rounded, divide dinto five
cells, about twice as broad as disca lspot.
Hindwing : transparent, veins and disca lspot orange-yellow with admixture of individual
black scales ; outer rnargin about twice as narrow as cilia, black ,basall yorange-yellow ;
disca lspot triangular ,reaching base of veins M3-Cui.
Variabilit y.This species is extremely variable, The abeve redescription concerns also
the extreme "black" and "yeliow" forms involving both males and females (Fig.20),
However, in contrast to B. tshinngana ,where one can observe a gradual change in colora-
tion from a "yellow" to a "black" form, the situation with B. wagneri is differen t.Thus,
the "black" form varies from the absence of coloured stripes on the abdomen dorsally to
the presence of a very narrow yellowish stripe on each of the 2nd, 4th and 6th tergite sbut
the abdomen remains always black ventrally. On the other hand, the "yellow" form varies
in the number of yellow and black scales on the labia lpalpus, vertex, thorax, and abdomen.
Besides ,there are specimens of the "yellow" forr nwith,black abdominal sternites with a
narrow yellow stripe distall y.Males always have orange latera lparts of the anal tuft.
Male genitalia (preparat iNoon. 93-16) . Tegumen-uncus complex (Fig 1,0) broad; scopula
androconialis well-developed, Iong ; crista gnathi laterali sshort, relatively narrow ; crista
gnathi medialis long and broad; valva (Fig 1.1) trapeziform; crista sacculi double: dorsal
leng
part relatively and narrow, covered with rows of strong pointed setae ; ventral part
small, oval, connected with dorsal part by rows of lappet-lik ef,lat-topp esdetae ; saccus
(Fig 1.2) gradually tapering toward base, with a rounded base, about 1.5 times as Iong as
vinculum ; aedeagus (Fi g1.3) thin, slightly shorter than valva ; vesica with rows of small
cornutl,
Female genitalia Not studied,
Diagnosis .The "black" form is habituall ysimilar to B, aimca (Pttnge l1e91r2,) ,but it
clearly differ sin the well-developed transparent areas of the forewing and in the black
discal spot of the hindwing (wit rhed-orange scales in the species cornpared). The "yel-
low" form is simi]ar to B. vigzarea (PUngele 1r9,12) ,and B Polyzon a(PUngele 1r9,12) .From
males of the firs tspecies, B. wcrgnen' can be distinguish ebdy the presence of the yellow
stripes on the 2nd, 5th ,6th ,and 7th abdominal tergites (eccurr ionnlyg on the 4th tergite in
the species compared). From the female of B. vigztrea and B. Polyzona t,his species differs
in the coioration of the abdomen.
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