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Inventory of Australian Megaloptera
larvae and resulting implications for
the classification of the group
(Insecta: Neuropterida)
Günther THEISCHINGER
Abstract Archichauliodes. Only three of four spe-
cies groups of Archichauliodes (Rie-
kochauliodes) separable on the basis of
adult characters, are distinguished on lar-
Abstract: The information avai-
val morphology. A key to the larvae of the
lable on the larvae of the Australian Mega-
now recognised group taxa of the Austra-
loptera is presented and used for re-apprai-
lian Megaloptera and distributional details
sal of some supraspecific taxa. Apochauli-
of most species are given in order to initia-
odes THEISCHINGER, formerly described as
te more thorough studies.
a subgenus of Archichauliodes WEELE is
accorded full generic status. A new subge-
nus, Riekochauliodes, is established for Key words: Megaloptera, Aus-
the majority of the Australian species of tralia, larvae, classification.
Stapfia 60,
zugleich Kataloge des OÖ. Landesmuseums,
Neue Folge Nr. 138 (1999), 85-100
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Introduction Systematics
Some information on the taxonomy of the Family Corydalidae (Figs 1-47)
larvae of the Australian Megaloptera was
Remarks: All Australian Corydalidae
given in the first revision of the group (RlEK
belong to the subfamily Chauliodinae.
1954). Presentation of accumulated informati-
Larvae: Abdomen with 8 pairs of late-
on on the larvae was announced for the futu-
ral processes (gills); apical segment of abdo-
re in a more comprehensive treatment of the
men with pair of prolegs but without terminal
adults (THE1SCHINGER 1983). In the meantime
filament.
and afterwards, keys to the larvae of families
and genera of Australian Megaloptera larvae Adults: Three ocelli. Fourth tarsal seg-
were given (e.g. RlEK 1970, WILLIAMS 1980, ment simple. Conspicuous pigment spots in
THEISCHINGER 1991). wings.
In this paper the now available taxonomic Distribution in Australia:
information on the larvae of the Australian Eastern Australia, south-western Australia.
Megaloptera is presented. This, together with Genera: Apochauliodes THEISCHINGER,
some additional information on the adults, Archichauliodes WEELE, Protochauliodes WEELE.
makes it possible to re-appraise some of the
supraspecific taxa as established by THEI-
Archichauliodes group (Figs 1-42)
SCHINGER (1983) and should enable to iden-
tify larvae to the level of those groups and in Larvae: Head and pronotum rather
some cases to the species. smooth, with only scattered long hairs. Left
mandible with 4, right mandible with 3 teeth
on inner margin (in addition to apical tooth).
Apical two segments of antennae combined
Material and methods
about as long as segment 2. Thorax and abdo-
men without prominent intersegmental lobes.
Larval material of Megaloptera from
eastern and south-western Australia, all of it Posterior pair of spiracles each raised (general-
now in the collection of the author and in the ly) or not raised (rarely) on a definite stalk.
Australian National Insect Collection Lateral processes of abdominal segments 1-5
(ANIC), preserved in 70-80% alcohol, was longer than width of respective segment.
studied. The diagnostic tools available for Adults: 2A of forewing connected to
identification of the major taxa were used in 1A by a crossvein. Female valves without late-
conjunction with reared material of Archich' ral stylus.
auliodes guuiferus (WALKER), A. anagaurus
Distribution: Australia, New Zea-
RlEK, A. plomleyi KlMMlNS and Protochauliodes
land, Chile.
biconicus KlMMlNS and with various ways of
Genera: Apochauliodes THEISCHINGER,
elimination in order to correlate adults and
Archichauliodes WEELE.
larvae of most species and to produce a preli-
minary guide to the Australian Megaloptera
Genus Apochauliodes THEISCHINGER
larvae.
(Figs 1-10)
In the following only brief diagnoses of
Apochauliodes THEISCHiNGER 1983:80 (as subgenus
families, genera and species groups of the Aus-
of Archichauliodes WEELE).
tralian Megaloptera and a key to those taxa Type species: Archichauliodes (Apochauliodes) cervu-
are given. For identification attempts beyond lus THEISCHINGER 1983.
that level the distributional overviews may be Larvae (Figs 1-6): Labium not produ-
useful. ced medially, truncate. Segment 2 of labial
palp not much longer than segment 1. Poster-
ior pair of spiracles each raised on a long stalk.
Lateral abdominal processes with short hair,
those of segments 1-5 distinctly longer than
width of respective segment.
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Adults (Figs 7-10): Forewing with Distribution: Eastern Australia,
CuA simple; male anal claspers bifid, with New Zealand, Chile.
laterobasal callosity not protruding; aedeagus Species in Australia: Archich-
bifid, symmetrical. auliodes (A.) anagaurus RlEK, A. (A.) simpsoni
Distribution: South-western Aus- THEISCHINGER.
tralia. Distributions: A. anagaurus is
Species: Apochauliodes cervidus (THEI- widely distributed mainly in alpine Victoria
SCHINGER). and south-eastern New South Wales; A. simp-
Larvae studied: Western Australia: Barlee Brook, soni is known only from Mt Howitt in Victo-
S of Nannup; Carey Brook, nr Pemberton; Pemberton. ria.
Larvae studied: New South Wales: Brown Mt;
Kiandra; Kosciusko Hotel; Pipers Ck, Mt Kosciusko;
Sawpit Ck, Mt Kosciusko; Wragge's Ck, Mt Kosciusko;
Genus Archichauliodes WEELE
Wilsons Valley, Mt Kosciusko. Victoria: Mt Buffalo;
(Figs 11-42)
Watt's R., Healesville.
Archkhauliodes WEELE 1909: 258.
Type species: Hermes diversus WALKER 1853.
Subgenus Riekochauliodes sub-
Larvae (Figs 11-16, 25-42): Labium gen. nov. (Figs 21-42)
produced medially. Segment 2 of labial palp Type species: Hermes gumferus WALKER 1853.
much longer than segment 1. Posterior pair of Larvae (Figs 25-42): Posterior tooth of
spiracles each raised (generally) or not raised mandibles generally smaller than median
(rarely) on a definite stalk. Lateral abdominal tooth. Posterior pair of spiracles each raised on
processes generally with short, rarely with a definite stalk.
long hair, those of segments 1-5 generally Adults (Figs 21-24): Forewing with
slightly longer than width of respective seg- tows of spots mostly between longitudinal
ment. veins; pigment spots of hindwing variably but
Adults (Figs 17-24): Forewing with distinctly enlatged. Aedeagus simple or widely
CuA forked; male anal claspers simple, with bifid, slightly to strongly asymmetrical.
laterobasal callosity prominent; aedeagus sim- Name : Dedication to Dr E.F. Riek
ple or bifid, symmetrical or assymetrical. (Canberra).
Distribution: Eastern Australia, Distribution: Eastern Australia.
New Zealand, Chile. Species groups: Archichauliodes
Subgenera: Archichauliodes WEELE, (Riekochauliodes) guttiferus group, A. (R.)
Riekochauliodes subgen. nov. deceptor group, A. (R.) polypastus group.
Subgenus Archichauliodes WEELE Archichauliodes (Riekochauliodes)
(Figs 11-20)
guttiferus group (Figs 21-30)
Archichauliodes WEELE 1909: 258.
Larvae (Figs 25-30): Stalk of posterior
Type species: Hermes diversus WALKER 1853.
spiracles shorter than half the distance bet-
Remarks: It is assumed that all availa- ween them; posterior spiracles not markedly
ble larval material belongs to A. anagaurus larger than spiracles on preceding segments.
RlEK. Adults (Figs 21-24): Male anal claspers
Larvae (Figs 11-16): Posterior tooth of amost straight, much longer than tergum 9;
mandibles generally larger than median tooth. aedeagus simple, asymmetrical.
Posterior pair of spiracles each not raised on a Species: Archichauliodes (Riekochauli-
definite stalk. odes) guttiferus (WALKER), A. (R.) neoguttiferus
Adults (Figs 17-20): Forewing with THEISCHINGER, A. (R.) phaeoscius RIEK, A.
rows of dark spots mostly on longitudinal (R.) rieki THEISCHINGER.
veins; pigment spots of hindwing hardly enlar- Distributions:A. guttiferus is known
ged. Aedeagus bifid, symmetrical. to occur only south of the Hunter River. A.
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rieki seems to be endemic to Carnarvon Range A. (R.) piscator THEISCHINGER, A. (R.) unci-
in southern inland Queensland. A. neoguttife- natus THEISCHINGER.
nis is known only from north of the Hunter Distributions: Only one species, A.
River (excluding Carnarvon Range), whereas deceptor, is known to occur south of the Expe-
A. phaeoscius has been collected only north dition Range (24° S); all its published recotds
from Paluma where it coexists in places (rain- are from north of the Hunter River. Only
forest) with A. neogutnferus. recently it has also been found at Black Mtn,
Larvae studied: Queensland: Beatrice R., Palmer- A.C.T. Two species, A. cuspidatus and A. gbs-
ston N.P.; Behana Gorge, S of Gordonvale; The Boul- sa, are known from the Eungella area
ders, Babinda; Broken R., Eungella N.P.; Crystal Casca-
(2O°55'S) and Expedition Range (24°° S); of
des, nr Cairns; upper Curambin Ck; 16 km on Davies
Ck Rd, E of Mareeba; Dayboro; 2 km on Mt Edith Rd, these only A. cuspidatus extends to north of
Tinaroo Dam; Finch Hatton; Fishery Falls, S of Gor- Paluma. There are no records of the remaining
donvale; 30 km S of Gympie; Kirrama S.F. (Western
species (A. collifer, A. conversus, A. lewis, A.
Fall); Little Mulgrave R.; Millstream Falls. New South
Wales: Allyn R., at Eccleston; upper Allyn R.; Bobun- piscator, A. uncinatus) from localities south of
dra Ck, Maffra; Bolairo; Brindabella, A.C.T.; Cabbage Paluma.
Tree Ck, Clyde Mt; Chandler R.; Cotter R., A.C.T;
upper Cotter; Dorrigo; Falconer Ck, 3 mi. E of Guyra; Larvae studied: Queensland: Captain Billy Ck,
Gara R.; Goodradigby R., Wee Jasper; Kiandra; Major's CY; Cheetah Ck, Evelyn Tableland; Cunninghams
Ck, Ebor; McLaughlin R.; Monga; Moonbi; Murrum- Gap; 16 km on Davies Ck Rd, E of Mareeba; 2 km on
bidgee R.-Eumarella R. Jn; Paddy's R., 6 mi from Cot- Mt Edith Rd, Tinaroo Dam; Iron Range, Mt Tozer foo-
ter, A.C.T; Peel R.; 7 W Rosebank; Snowy R., Dalgety. thills; upper Lankelly Ck, Coen Dist.; McLeod R., 15
Victoria: Aberfeldy R., Wallhalla Rd; Bright; Mt Buf- km W of Mt Carbine; Mt Lewis; Mossman Gorge;
falo; Bullarook Ck, at Kingston; Wellington R., Licola Rocky Ck, Atherton.
Rd.
Archichauliodes (Riekochauliodes) Archichauliodes {Riekochauliodes)
deceptor group (Figs 31-36) polypastus group (Figs 37-42)
Remarks: On the basis of a more ot less Larvae (Figs 37-42): Stalk of posterior
elongate tergum 9 of the adult male an A. spiracles several times as long as wide, mar-
deceptor group (species A. cuspidatus, A. decep- kedly longer than half the distance between
tor, A. ghssa) and an A. uncinatus group (spe- them; posterior spiracles significantly larger
cies: A. collifer, A. conversus, A. lewis, A. pis-
than spiracles on preceding segments.
cator, A. uncinatus) were distinguished (THEI-
Adults: Male anal claspers strongly
SCHINGER 1983, 1988). It appears that these
bent, not longer than tergum 9; aedeagus
and other group specific differences in the
widely bifid, strongly asymmetrical.
adults are not reflected in the morphology of
the larvae. Only a single group (A. deceptor Species: Archichauliodes (Riekochauli-
group), including all above species, is now odes) isolatus THEISCHINGER, A. (R.) pictus
recognised. THEISCHINGER, A. (R.) pbmleyi KlMMiNS, A.
(R.) polypastus RiEK.
Larvae (Figs 31-36): Stalk of posterior
spiracles several times as long as wide, mar- Distributions: A. isolatus is known
kedly longer than half the distance between only from Fräser Island. Records of A. plomleyi
them; postetior spiracles not markedly larger are available from Victoria and from New
than spiracles on preceding segments. South Wales north to Barrington Tops. Both,
A. pictus and A. polypastus, are known from
Adults: Male anal claspets slightly cur-
north-eastern New South Wales and south-
ved, not longer than tergum 9; aedeagus sim-
eastern Queensland.
ple, slightly to strongly asymmettical
Larvae studied: Queensland: Lower Bellungui
Species: Archichauliodes (Riekochauli- Falls, Lamington N.P.; Binna Burra; Bunya Mts; Flaggy
odes) cuspidatus THEISCHINGER, A. (R.) decep- Ck, Mistake Mts, via Laidley; Joalah N.P., Tamborine
Mtn. New South Wales: Bowens Ck, Bell; Brown
tor KlMMiNS, A. (R.) gbssae THEISCHINGER, A.
Mtn; Cyde Mt, E slope; Clyde Mt, W slope; Kosciusko
(R.) collifer THEISCHINGER, A. (R.) conversus Hotel; upper reaches of Middle Harbour, Sydney;
THEISCHINGER, A. (R.) lewis THEISCHINGER, Minyon Falls, 10 W Rosebank; Moonbi; Rosebank.
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Genus Protochauliodes WEELE Major's Ck, Ebor; upper reaches of Middle Harbour,
(Figs 43-47) Sydney; Newell Falls; Styx R., at Hyatrs Flat.
Prowctauliooes WEELE 1909: 258.
Type species: Hermes diversus WALKER 1853.
Austrochauliodes RiEK 1954: 137. Family Sialidae (Figs 48-52)
Type species as indicated: Hermes dubiuuus WAL-
KER 1853; de facto: Protochfluliodes biconicus KlMMINS Remarks: The available larvae appear
1954. very homogeneous. They probably belong to a
Remarks: At present at least three single species (Stenosiaiis australiensis).
geographical groups of species are included in
Larvae (Figs 48-52): Abdomen with 7
ProwchauUodes. The Australian, South Ameri-
pairs of lateral processes (gills); apical segment
can and North American components may in
of abdomen without prolegs but with terminal
fact constitute discrete taxa. It is possible that
filament.
Austrochauliodes RiEK will have to be reinsta-
Adults: Ocelli wanting. Fourth tarsal
ted for the Australian group of species [after
segment prominently bilobed. Wings without
comprehensive study and decision by the
conspicuous pigment spots.
ICZN (International Commision on Zoologi-
cal Nomenclature) on the type species of Aus- Distribution in Australia:
trochauliodes)]. Eastern Australia.
Genera: Austrosiaiis TlLLYARD, Stenosia-
Larvae: (Figs 43-47): Head and prono-
Us TlLLYARD.
tum covered with fine hairs. Both mandibles
Larvae studied: Queensland: Beatrice R., Palmer-
with 3 teeth on inner margin (in addition to
ston N.P.; Lake Eacham; 30 km S of Gympie.
apical tooth). Labium slightly produced medi-
ally with apex slightly bilobed, very hairy. Seg-
ment 2 of labial palps almost twice as long as Genus Austrosiaiis TILLYARD
segment 1. Apical two segments of antennae
Austrosiaiis TlLLYARD 1919: 821.
combined much shorter than segment 2. Type species: Austrosiaiis ignicottis TlLLYARD 1919.
Thorax and abdomen generally with promi-
Larvae: Diagnostic characters not
nent intersegmental lobes. Spiracles on abdo-
available.
minal segment 8 only slightly raised (stalked).
Lateral processes of abdominal segments 1-5 Adults: Forks of Rs, MA, MP and CuA
shorter than respective segment. subeqal in length. Sternum 8 of female long,
significantly incised posteromedially.
A d u 11 s : 2 A of forewing fused to 1A for
Species: Austrosiaiis ignicollis
part of its length. Female valves with lateral
TlLLYARD, A. maxmouldsi THEISCHINGER.
stylus.
Distributions: Only a few speci-
Distribution: Eastern Australia,
mens of A. maxmouldsi have ever been recor-
Chile, western North America,.
ded from north-eastern Queensland, whereas
Species in Australia: Prow- A. ignicollis is known only from a unique fema-
chauliodes biconicus KlMMINS, P. kirramae THEI- le from Tasmania.
SCHINGER, P. eungella THEISCHINGER.
Distributions: Several subspecies
and local forms of P. biconicus are known from Genus Stenosiaiis TILLYARD
New South Wales and south-eastern Queens-
Stenosiaiis TlLLYARD 1919: 823.
land. P. eungella is only known from the Eun-
Type species: Stenosidis australiensis TlLLYARD
gella area, whereas P. lärramae has been found 1919.
only north from Paluma.
Larvae: Diagnostic characters not
Larvae studied: Queensland: Broken R., Eungella
available.
N.P.; Bunya Mts; Big Falls, Bunya Mts; Cheetah Ck,
Evelyn Tableland; The Crater, SW of Malanda; Cun- Adults: Forks of Rs, MA, MP and CuA
ninghams Gap; 16 km on Davies Ck Rd, E of Mareeba;
very different from each other in length. Ster-
Flaggy Ck, Mistake Mts, via Laidley; Kirrama S.F.; Litt-
le Crystal Ck, Mt Spec; Beatrice R., Palmerston N.P. num 8 of female wide and very short, not inci-
New South Wales: Clyde Mt; Fitzroy Falls; Mt Keira; sed posteromedially.
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Species: Stenosiaiis australiensis TlL- Spiracles of 8th abdominal segment each
LYARD, S. holbwayi THEISCHINGER. raised on a definite stalk (Figs 30, 36,42);
lateral abdominal processes clothed with
Distributions: S. australiensis is
short hair (Figs 25, 31, 37)
known from numerous localities of coastal
Subgenus Riekochauliodes, 5
eastern Australia (Cape York-Victoria), S. hoi-
lowayi only from a few specimens from north- 5 Spiracles of 8th abdominal segment each
on moderately long stalk (Fig. 30); stalks
eastern Queensland.
hardly half as long as distance between
them Archichauliodes (Riekochauli-
odes) guttiferus group
Key to the supraspecific taxa of
Australian Megaloptera (larvae) " Spiracles of 8th abdominal segment each
on long stalk (Figs 36,42); stalks almost as
1 Seven pairs of non-articulated, lateral long as distance between them, or longer
abdominal processes (Fig. 48); apex of 6
abdomen drawn out into a long terminal .. ,, , . . . „ . . . . . •,
c
6 Stalked spiracles of 8th abdominal seg-
subanal filament Family Sialidae , ... _,, , . ,
ment not disxtinctly l arger iLthan spiracles
- Eight pairs of articulated lateral abdomi- on preceding segments (Fig. 36)
nal processes (Figs 1, 11, 25, 43); apical Archichauliodes (Riekochauliodes)
segment of abdomen with a pair of pro- deceptor group
legs; functional spiracles on abdominal . | pj | s of 8th abdominal seg-
Staked S race
segment 8, sometimes each drawn out on distinctly larger than spiracles on
ment
a stalk Family Corydalidae, 2 preceding segments (Fig. 42)
. „ , .... . , , . Archichauliodes (Riekochauliodes)
iU iU
2 Both mandibles with 3 teeth on inner
• / i /i owe AA\ u J A polypastus group
margin (rarely 4-3)(Fig. 44); head and
pronotum densely clothed with setae;
apical 2 segments of antennae combined
much shorter than segment 2 (Fig.47)
Genus Protochauliodes Discussion
- Left mandible with 4, right mandible _ , , . „ , ,
. , , , , . . . .. . , , , Lorydalidae: boutn-w estern AuAstralian
with 3 teeth (in addition to apical tooth)
. ... - ,. ... , larval and adult material of what was hitherto
on inner margin (Figs 2, 12, 26); surface
of head and pronotum almost free of considered t0 be Archichauliodes WEELE, is qui-
setae; apical 2 segments of antennae te distinct from eastern A^tralian material;
combined about as long as segment 2 Apochauliodes THEISCHINGER is therefore and
(Figs 5 15 29) reconsidering the diagnostic characters of the
Archichauliodes group, 3 adults (THEISCHINGER 1983 and this paper)
given generic rank. What remains of Australi-
3 Second segment of labial palp almost as ,
gn Archichauliodes can> on the basis of larva
long as first (Fig. 4); apex of labium not , , , , , ,. ., , . ,
3 and adult characters, be divided into 2 groups
produced, truncate; known only from . , , , . _
,. one of which ( AA. anagaurus group of THEI-
western Australia „ „ , „ ,
_ . . .. . SCHINGER 1983) appears markedly closer to
Genus Apochauliodes rr '
the type species of Archichauliodes from New
- Second segment of labial palp much Ion- Zealand and to Archichauliodes from South
ger than first (Figs 14, 28); apex of labium America, than to the other group of Australi-
produced, convex, slightly lobed; from cies. The Archichauliodes anagaurus
an spe
eastern Australia group is therefore assigned to Archichauliodes s.
Genus Archichauliodes, 4 . _ R ko hauliodes,
sa whereas a new subgenuS) ie C
4 Spiracles of 8th abdominal segment each is established for the remaining Australian
only slightly raised (Fig. 16); lateral abdo- species of Archichauliodes. At present larvae of
minal processes clothed with long hair two of the four species groups distinguished by
(Fig. 11) THEISCHINGER (1983) of what is now Archich-
Subgenus Archichauliodes auliodes (Riekochauliodes) cannot be separated.
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Thus only three species groups of Archichauli- References
odes (Riekochauliodes) are recognised in this
paper. Clearly diagnostic larval characters wit- KIMMINS D.E. (1954): A new genus and some new spe-
cies of the Chauliodini (Megaloptera), with
hin the species groups of Archichauliodes were
notes on certain previously described species. —
not found. The larvae of all species of Proto- Bull. Brit. Mus. (nat. Hist.) Ent. 3: 417-444.
chauliodes WEELE are very similar; at present
RIEK E.F. (1954): The Australian Megaloptera or
specific identifications of larvae on morpholo- alder-flies. — Aust. J. Zool. 2: 131-142.
gical basis are not possible. RIEK E.F. (1970): Megaloptera (Alderflies). — pp. 465-
471 in CSIRO (sponsor): The Insects of Australia.
Sialidae: The available larvae from Aus-
A textbook for students and research workers.
tralia appear very homogeneous. Probably Melbourne: Melbourne Univ. Press.
they represent only a single species. Larval
THEISCHINGER G. (1983): The adults of the Australian
diagnoses of the sialid genera Auscroskdis and Megaloptera. — Aquat. Ins. 5: 77-98.
Stenosialis cannot be given. THEISCHINGER G. (1991): Megaloptera (Alderflies, Dob-
sonflies). — pp. 516-520 in CSIRO (sponsor): The
Insects of Australia. A textbook for students and
research workers. Second edition. Melbourne:
Melbourne Univ. Press.
Acknowledgements TILLYARD R.J. (1919): Australian Megaloptera or alder-
flies, with descriptions of new genera and spe-
cies. — Proc. Linn. Soc. N.S.W. 43(1918): 819-
Dr E.F. Riek (Canberra) recently returned
825.
illustrations of adults and larvae of Megalop-
WALKER F. (1853): List of the Specimens of Neurop-
tera to CSIRO Entomology (Curator of aqua-
terous Insects in the Collection of the British
tic insects Dr P.S. Cranston). These illustrati- Museum Part 2 (Sialidae-Neuropterides). Lon-
ons were done by Mrs S. Monteith (Brisbane) don: British Museum: pp. 193-476.
more than two decades ago. 1 wish to thank WEELE H.W. van der (1909): New genera and species
of Megaloptera LATR. — Notes Leyden Mus. 30:
Mrs S. Monteith, Dr E.F. Riek and Dr P.S.
249-264.
Cranston for giving me the opportunity to use
WILLIAMS W.D. (1980): Australian Freshwater Life. The
some of those illustrations.
Invertebrates of Australian Inland Waters.
Second edition: 321 pp. Melbourne: Macmillan.
Zusammenfassung
Die verfügbare Information über die Lar-
ven der australischen Megalopteren wird
zusammenfassend unter dem Gesichtspunkt
der Beurteilung der Systematik einiger
supraspezifischer Taxa diskutiert. Apochauli-
odes THEISCHINCER, ursprünglich als Subgenus
von Archichauliodes WEELE beschrieben, wird
in den Genus-Rang erhoben. Ein neues Sub-
genus, Riekochauiiodes, wird für den Großteil
der australischen Spezies von Archichauliodes
errichtet. Nur drei der vier Spezies-Gruppen
von Archichauliodes (Riekochauliodes), die auf
Grund imaginaler Merkmale abgegrenzt wer-
den können, lassen sich auch auf der Basis lar-
valer Merkmale unterscheiden. Es wird ein
Bestimmungsschlüssel der Larven der australi-
schen Megalopteren (soweit möglich) präsen- Anschrift des Verfassers:
tiert, für die meisten Spezies werden detaillier-
Günther THEISCHINGER
te Angaben zur Verbreitung gegeben, womit
2a Hammersley Rd, Grays Point
eine Basis für weitergehende Studien vorliegt. N.S.W., Australia 2232
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Figs 1-6: Apochauliodes cervulus (THEISCHINGER), larva: 1 - habitus, dorsal aspect; 2 - head, dorsal aspect; 3 - head, ventral aspect; 4 - labi-
um and palps; 5 - antenna; 6 - segment 8, dorsal aspect.
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Figs 7-10: Apochauliodes cervulus (THEISCHINGER), male: 7 - wings; 8, 9 - anal claspers: 8 - dorsal aspect; 9 - lateral aspect; 10 - aedeagus,
ventral aspect.
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11 12
Figs 11-16: Archichauliodes (Archichauliodes) anagaurus RIEK, larva: 11 - habitus, dorsal aspect; 12 - head, dorsal aspect; 13 - head, ven-
tral aspect; 14 - labium and palps; 15 - antenna; 16 - segment 8, dorsal aspect.
94
