Table Of Content)
THE NAUTILUS 127(4):160-161, 2013 Page 160
Research Note
Intaglicollonia, a new name for the Eocene interpreted originally as an unusual form of discor-
gastropod Nehalemia Hickman, 1974 dant (non-collabral) secretory activity by the mantle
(Hickman, 1974). Discussion with Jonathan Todd (The
(Gastropoda: Coloniidae)
Natural History Museum, London) raises the interesting
Intaglicollonia is proposed as a replacement name for pPoaslsmibeirliatyndthPaltewtehse, 1li9n9e3s; Taroedd“pasnedudPoaeltmcehri,ng2s0”02)(segnesnu-
the monotypic Eocene gastropod genus Nehalemia
erated as an active secretory response to the presence of
Hickman, 1974, family Colloniidae Cossmann, 1916.
The type species, Nehalemia hieroghjphica Hickman, a thin, tubular commensal cnidarian. In this instance the
shallowly-incised lines would represent systematic and
1974, was based on 11 specimens from a siltstone unit at
continuous growth response by the snail to deformation
the top of the Cowlitz Formation in Washington County,
northwestern Oregon (early late Eocene). The siltstone of the periostracal sheet at the growing shell margin.
Similaractivedeformation responses havebeen reported
is immediately below the uneonformable contact with
the basal memberof the overlying latest Eocene to earli- in fossil bivalves (Todd, 1993).
est Oligocene Keasey Formation. The genus name is CanItnrtaagilniecaollJoenfifare,ys,Ph1a8n8e3roalreepikdnaownDaloin,ly f1r9o0m7 deaenpd-
preoccupied byNehalemia Blake, 1973, an asteroid eelh-
noderm from the middle member of the Keasey Forma- water faunas and sedimentary facies (>200 m) and are
tion in Columbia County, Oregon. The type localities for dtihseticnoglulionsihiendefgreonmutsheHoshmaallloowp-owmaaterCa“rdpweanrtfert,urb1a8n6s4’ boyf
both species are on the Nehalem River, approximately
25 km apart and with a vertical stratigraphic separation their larger size and lack of nodulose spiral ribs sepa-
rated by deep grooves. Shells ofthe Mediterranean type
of approximately 450 m. Although the taxa are in sepa-
species, Homalopoma sanguineum (Linnaeus, 1758) are
rate phyla, the geographic and stratigraphic proximities 6-7 mm in diameter and red or pink in color, in contrast
of the type localities support the decision to avoid an
to the unpigmented shells of the larger-shelled species
orthographicallysimilar replacement name.
retIanitnasgltihcoellsopneicaifiiscmoenpiotthyepticoraingidnailtslytyppreosppoesceidesunnadmeer in Tthheedreeeppl-awcaetmeerntgennearmae.Intaglicollonia is chosen to call
attention to the finely-incised intagliare markings on
Nehalemia by Hickman (1974). Although the genus is (
monotypic, it is the basal member of a putative evolu- ptohseitsihoelnlofsutrhfeacgeenasusweilnlCaoslleomnpihiadaseizCionsgsmthaennt,ax1o9n1o6m,ica
tionarylineage in which intraspecificsculptural polymor-
basal gastropod family group with a concentration of
phism has been studied, and named patterns of incised
morphological diversity in the Cenozoie fossil record
lines have an ontogeneticorderofappearance in individ-
(Hickman and McLean, 1990). Publicationof the replace-
uals (Noda and Ogasawara, 1976). Further study ofvari-
ment name is prerequisite to a forthcoming reinterpreta-
ation in theonsetoftimingofthe incisedpatternshas led
tion ofthe taxonomy, paleoeeology, and unusual features
to a model of heterochronic evolution in the lineage
(Majima and Murata, 1992). The genus also has figured that Intagliocollonia shares with fossil and living species
of Cantrainea and Phanerolepida.
in the identification ofthe late Eocene origins of north
Pacific deep-water communities (Hickman, 1984), in
the characterization of biotic response to the Eocene-
ACKNOWLEDGMENTS
Oligocene climatic transition (Hickman, 2003), and in
t(hHeicrkemcaonns,tr2u0c0t3i;onAmoafnpoa,tt2e0r0n5s).oftrans-Paeifie migration rIeavimewgerratfeofrulthteoirRsiucghgaersdtiSoqnusifroersiamnpdroavnemaennotnyomfotuhse
Blake’s (1973) genus is also monotypic and of consid- manuscript. I thank Jon Todd for introducing me to
erable interest as a component ofthe famous Mist fauna pseudoetchings and criteria for recognizing periostracal
in northwest Oregon in the upper part of tire middle deformation structures. Thanks also go to D. Smith
member of the Keasey Formation. The Mist locality is for assistance in preparing the Figures. This is con-
noteworthy as a Tertiary erinoid Lagerstiitte (Moore and tribution 2047 of the University of California Museum
Vokes, 1953) as well as for its well-preserved asteroids, of Paleontology.
ophiuroids, and echinoids (Burns and Mooi, 2003). The
carbonate layers at the Mist Locality are also noteworthy
for providingpetrographic and isotopicevidence of meth- LITERATURE CITED
anogenesis, possiblycontemporaneouswith thefaunaand
a potential cold seep environment (Burns et al., 2005). Amano, K. 2005. Migration and adaptation ol Late Cenozoie
New illustrations of the holotype of Intaglicollonia cold-water molluscs in the North Pacific. In: A.M.T
hieroglyphica are presented (Figures 1, 2) along with a Elewa (ed.) Migration of Organisms. Springer, Berlin
detail ofthe fine engraved lines (Figure 3), which were pp. 128-150.
C.S. Hickman, 2013 Page 161
Figures 1-3. Holotype (USNM 647339) of Intaglicollonia hieroglijphica Hickman, 1979; height = II mm, diameter = 16 mm
1. Apical and 2. Ahapertural view, 3. Detail of incised sculptureon bodywhorl.
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