Table Of ContentFAO Fisheries Synopsis No. 125, Volume 10 FIR/S125 Vol. 10
FAO SPECIES CATALOGUE
VOL. 10. GADIFORM FISHES OF THE WORLD
(Order GADIFORMES)
AN ANNOTATED AND ILLUSTRATED CATALOGUE
OF CODS, HAKES, GRENADIERS AND OTHER
GADIFORM FISHES KNOWN TO DATE
FOOD AND AGRICULTURE ORGANIZATION OF THE UNITED NATIONS
FAO Fisheries Synopsis No. 125, Volume 10 FIR/S125 Vol. 10
FAO SPECIES CATALOGUE
VOL.10 GADIFORM FISHES OF THE WORLD
(Order Gadiformes)
An Annotated and Illustrated Catalogue of Cods, Hakes,
Grenadiers and other Gadiform Fishes Known to Date
prepared by
Daniel M. Cohen
larutaN yrotsiH muesuM fo soL selegnAytnuoC
Los Angeles, California, USA
(Taxonomy of Bregmacerotidae, Euclichthyidae, Gadidae,
Melanonidae, Moridae, Muraenolepididae)
Tadashi lnada
Tohoku National Fisheries Research Institute
ehonihcaH ,hcnarB ,emaS ehonihcaH ,130 napaJ
(Merlucciidae)
Tomio lwamoto
ainrofilaC ymedacA fo secneicS
naS ,ocsicnarF ,ainrofilaCASU
(Taxonomy of Macrouridae)
Nadia Scialabba
yrehsiF secruoseR dna tnemnorivnEnoisiviD
,OAF emoR
(Compilation of information on habitat and fisheries)
FOOD AND AGRICULTURE ORGANIZATION OF THE UNITED NATIONS
Rome, 1990
The designations employed and the presentation of
material in this publication do not imply the
expression of any opinion whatsoever on the part of
the Food and Agriculture Organization of the
United Nations concerning the legal status of any
country, territory, city or area or of its authorities, or
concerning the delimitation of its frontiers or
boundaries.
M-40
ISBN 92-5-102890-7
All rights reserved. No part of this publication may be reproduced, stored in a
retrieval system, or transmitted in any form or by any means, electronic,
mechanical, photocopying or otherwise, without the prior permission of the
copyright owner. Applications for such permission, with a statement of the
purpose and extent of the reproduction, should be addressed to the Director,
Publications Division, Food and Agriculture Organization of the United Nations,
Via delle Terme di Caracalla, 00100 Rome, Italy.
0
C FAO Rome 1990
iii
PREPARATION OF THIS DOCUMENT
Since the Gadiformes include some of the world’s most important commercial fish species, plans for the present
catalogue were conceived many years ago. However, because of the confused state of the higher systematics of this
complex and highly polymorphic group of fishes, the large number of poorly known species in some gadiform
families, and the wealth of information on habitat and fisheries available on others, the original draft manuscript
prepared in 1985 had to be substantially revised several times. Even in its present stage it does not reflect the latest
findings in both the fields of taxonomy and fisheries. Nevertheless, it represents an up-to-date inventory of the
species considered to be of present or potential interest to fisheries and is expected to become an important
reference source for fishery workers throughout the world.
The 3 ichthyologists who have prepared the basic manuscript of the catalogue are internationally recognized experts
in their respective fields and have been collaborating with the FAO Species Identification and Data Programme for
many years.
Technical Editors: W. Fischer, W. Schneider and L. Garibaldi, Fisheries Resources and Environment Division, FAO
Illustrators: P. Lastrico, D. de Merich and M. D’Antoni, FAO, Rome
Page composition: G. Sciarappa-Demuro, FAO, Rome
Cohen, D.M.; Inada.T.; Iwamoto, T.; Scialabba, N.
FAO species catalogue. Vol. 10. Gadiform fishes of the world (Order Gadiformes). An anno-
tated and illustrated catalogue of cods, hakes, grenadiers and other gadiform fishes known
to date.
FAO Fisheries Synopsis. No. 125, Vol. 10. Rome, FAO. 1990. 442 p.
ABSTRACT
This is the tenth issue in the FAO series of worldwide annotated and illustrated catalogues
(synopses) of major groups of organisms that enter marine fisheries. It includes all gadiform
fishes (order GADIFORMES - cods, hakes, grenadiers, moras, moray cods, pelagic cods, codlets
and eucla cods), comprising 8 families, 59 genera and more than 180 species, which contribute
more than a quarter of the world’s marine fish catch. Keys are given to the families, subfamilies
and genera, with a glossary of technical terms and measurements, and a review of major
literature sources within each FAO fishing area. Genera and species are diagnosed, with
drawings, scientific and vernacular names, information on habitat, biology, fisheries and
distribution (with map); reference is also made to subspecies and to any taxonomic or other
problems. The occurrence of the species within the FAO fishing areas is tabulated. All scientific
names still applied to gadiform species are given in the synonymies, and these as well as the
vernacular names are indexed; there is also a complete bibliography of all literature cited.
Distribution
Authors
FAO Fisheries Officers
Regional Fisheries Councils
and Commissions
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TABLE OF CONTENTS
Page
1. INTRODUCTION .................................................................................... 1
1.1 mesrofid ansGokram elRareneG .............................................................. 1
1.2 seirehsiF no noitamrofnI ....................................................................... 4
3.1 nalP fo eht citametsySugolataC e ............................................................... 6
4.1 Problems with Identification ................................................................... 7
1.5 Illustrated Glossary of Technical Terms and Measurements ...................................... 9
2. SYSTEMATIC CATALOGUE OF SPECIES .......................................................... 13
1.2 seilimaF ot yeK ................................................................................ 13
Code
2.2 YLIMAF BREGMACEROTIDAE ............ .......... .......... .......... . BREG ............. .. . 16
suneG Bregmaceros ..................................... BREG Breg .......... 16
Tentative list of species .......................................................... 16
Bregmaceros sp. ........................................... . BREG Breg 1 ......... 17
2.3 YLIMAF EUCLICHTHYIDAE ............................................. EUCL .................18
2.4 FAMILY GADIDAE .................................................... G A D I ............... 18
Key to genera ........................................................................... 20
suneG Arctogadus ...... ....... ...... ....... ...... ...... ....... ... G A D I A r c ............ 25
Key to species ................................................................... . 25
Arctogadus borisovi ....................................... G A D I A rc 1........... 25
Arctogadus glacialis ....................................... G A D I A r c 2 .......... 26
suneG Boreogadus ................................. .............. G A D I B o r ............ 27
Boreogadus saida ......................................... G A D l B o r 1 .......... 27
Genus Brosme ................................................... G A D I B r o ............ 29
Brosme brosme ............................................ G A D l B r o 1 .......... 29
Genus Ciliata .................................................... G A D l C il ............. 31
Key to species .................................................................... 31
Ciliata mustela ............................................ G A D I C i l 1 ........... 32
Ciliata septentrionalis ..................................... G A D I C i l 2 ........... 33
Genus Eleginus ................................................. GADI Ele ............ 33
Key to species .................................................................... 34
Eleginus gracilis ............................................ G A D I E le 1.......... 34
Eleginus navaga ........................................... G A D I E le 2........... 36
suneG Enchelyopus ............................................... G A D I E n ............. 38
Enchelyopus cimbrius ..................................... G A D I E n 1 .......... 38
Genus Gadiculus ................................................. G A D I G a d i ........... 39
Gadiculus argenteus ...................................... G A D I G a d i 1 ......... 39
v
Genus Gadus .................................................. G A D l G a d u . . . . . . . . . . . . .41
Key to species ................................................................. .. 41
Gadus macrocephalus................................... GADI Gadu1........... 42
Gadus morhua .......................................... GADl Gadu 2 ......... 44 ..................................
Gadus ogac ............................................. G A D l G a d u 3 .......... 47
Genus Gaidropsarus.. ............... ....................... ... G A D I G a i d r ........... 48
Tentative list of species ....................................................... . ......... 49
Gaidropsarus biscayensisl.....l....l.....l.....l....l.....l... . GA DI G aid r 3l ..l..l..l...l.. 49
Gaidropsarus macrophthalmusl..l...l..l..l...l..l..l...l..l...l.. . G A D l G a id r 4l l..l..l...l... 50
Gaidropsarus mediterraneus l...l..l..l...l..l...l..l..l...l..l..l... G A D I G a id r 1 l .l..l..l...... 50
Gaidropsarus vulgaris l...l..l..l...l..l..l...l..l..l...l..l..l...l.. .. G A D I G a i d r 2 ..... ..... 5 1
Genus Lotaa................................................... GADl Lota ............. 52
Lota lota ................................................ GADl Lota 1l ........... 53
Genus Melanogrammus ..................... ................... G A D I M e l a ........... .. 54
Melanogrammus aeglefinus ............................ G A D I M e la 1 ........... 54
suneG Merlangius .............................................. G A D I M e r la ............ .55
Merlangius merlangus .................................... G A D I M e r la 1 ............56
s uneG Microgadus .............................................G A D I M i cr o ............. 57
Key to specie s .................................................................... 57
Microgadus proximus..................................... GADI Micro 1.............58
Microgadus tomcod....................................... GADI Micro 2............58
s uneG Micromesistius ..........................................G A D I M i c r o m ........... 60
Key to specie s .....................................................................60
Micromesistius australis....................................G A D I M ic r o m 2 ..........60
Micromesistius poutassou ................................ G A D I M ic r o m 1 ..........61
s uneG Molva .................................................. GADl Molv .............. 63
Key to species .................................................................. 63
Molva dypterygia ...................................... GADI Molv 1 ......... 63
Molva molva ............................................ G ADl Molv2 ......... 64
Genus Phycis ................................................ GADl Phyc ............ 65
Key to specie s .................................................................. 66
.
Phycis blennoides ........................................G A D I P h y c 1 ........... 66
Phycis chesteri .......................................... GADl Phyc 3 ......... 67
.
Phycis phycis ................................................. GADI Phyc 2 ........ 68
s uneG Pollachius ............................................ GADl Pol ............ 69
Key to specie s .................................................................. 70
Pollachius pollachius...................................... G A D I P o l 1 ........... 70
Pollachius virens ....................................... GGAADDlI PPoll l 22 ............ 71
s uneG Raniceps ............................................. GADI Ran ............ 73
Raniceps raninus...................................... G A D I R a n 1 ............ 73
s uneG Theragra ...................................................................... 74
Key to specie s .................................................................. 74
Theragra chalcogramma....................................G A D I T h e r 1 ........... 75
Theragra finnmarchica .................................... G A D I T h e r 2 ........... 76
s uneG Trisopterus ..............................................G A D I T r is o ............. 77
Key to specie s .................................................................. 77
.
Trisopterus esmarkii .................................... G A D I T r is o 3 ........... 77
Trisopterus luscus ....................................... GADI Triso 2 . . . . . . . . 79
Trisopterus minutus ...................................... G A D I T r iso 1 . . . . . . . . . . . 80
vi
suneG Urophycis ............................................... GADI Uroph .............81
Key to species .................................................................. 81
Urophycis brasiliensis ................................. GADI Uroph 3 ........82
Urophycis chuss ....................................... G A D I U r o p h 4 ........83
Urophycis cirrata ....................................... GADI Uroph 5 ........85
Urophycis earlii ....................................... G A D I U r o p h 6 . ........85
Urophycis floridana .................................... GADI Urop h 1 .........86
Urophycis regia......................................... G A D I U r o p h 2 .. .......87
Urophycis tenuis....................................... G A D I U r o p h 7 ........ 88
2.5 FAMILY MACROURIDAE .......................................... MACROUR..............90
Key to Subfamilies ....................................................................91
Subfamily Bathygadinae .................................................................92
Key to genera ........................................................................93
Genus Bathygadus ............................................ MACROUR Bath........93
List of species .................................................................. 94
Bathygadus macrops.................................... M A C R O U R B at h 1 ......94
Bathygadus melanobranchus .......................... M A C R O U R B a t h 2 ......95
Genus Gadomus ............................................. MACROUR Gado ......96
List of species ..................................................................97
Gadomus arcuatus ........... ........................ M A C R O U R G a d o 1 ......97
Subfamily Macrourinae .................................................................. 98
Key to genera .....................................................................99
Genus Albatrossi a ............................................. M A C R O U R A l b ........110
Albatrossia pectoralis ................................... M A C R O U R A l b 1 .....110
Genus Coelorinchus ........................................... M A C R O U R C o e l.......111
Key to species .l.................................................................112
List of species .................................................................137
Coelorinchus acanthigerl................................. M A C R O U R C o e l 4 .....139
Coelorinchus aconcagua ................................. M A C R O U R C o e l 5 ....140
Coelorinchus anatirostris ................................ M A C R O U R C o e l 6 .....141
Coelorinchus argentatus................................. M A C R O U R C o e l 7 .....142
Coelorinchus argus . ..................................... M A C R O U R C o e l 8 .....144
Coelorinchus aspercephalus ............................. M A C R O U R C o e l 9 .....145
Coelorinchus australis................................... M A C R O U R C o e l 1 0 ....146
Coelorinchus biclinozonalis............................... M A C R O U R C o e l 1 1 ....147
Coelorinchus braueri .................................... M A C R O U R C o e l 1 2 ....149
Coelorinchus canus....................................... M A C R O U R C o e l 1 3 ....150
Coelorinchus caribbaeus................................. M A C R O U R C o e l 1 4 ....151
Coelorinchus chilensis .................................. M A C R O U R C o e l 1 5 ....152
Coelorinchus coelorhincus............................... M A C R O U R C o e l 1 .....154
Coelorinchus denticulatus ............................... M A C R O U R C o e l 1 6 ....155
Coelorinchus fasciatus .................................. M A C R O U R C o e l 2 ....156
Coelorinchus formosanus................................ M A C R O U R C o e l 1 7 ....158
Coelorinchus hexfasciatus ............................... M A C R O U R C o e l 1 8 ....159
Coelorinchus innotabilis.................................. M A C R O U R C o e l 1 9 ....161
Coelorinchus japonicus.................................. M A C R O U R C o e l 2 0.....162
Coelorinchus kaiyomaru ................................. M A C R O U R C o e l 2 1 ....163
Coelorinchus kamoharai ................................. M A C R O U R C o e l 2 2 ....164
Coelorinchus karrerae ................................... M A C R O U R C o e l 2 3 ....166
Coelorinchus kishinouyei ................................ M A C R O U R C o e l 2 4 ....167
Coelorinchus labiatus.................................... M A C R O U R C o e l 2 5 ....168
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Coelorinchus longicephalus .............................. MACROUR Coel 26 ... 169
Coelorinchus macrochir...................................MACROUR Coel 27.... 171
Coelorinchus marinii .................................... MACROUR Coel 3..... 172
Coelorinchus matamua ................................... MACROUR Coel 28.... 173
Coelorinchus multispinulosus ............................ MACROUR Coel 29.... 174
Coelorinchus occa ...................................... MACROUR Coel 30.... 176
Coelorinchus oliverianus ................................. MACROUR Coel 31.... 177
Coelorinchus parallelus ..................................MACROUR Coel 32.... 178
Coelorinchus productus ................................. MACROUR Coel 33.... 179
Coelorinchus scaphopsis ................................ MACROUR Coel 34 .... 181
Coelorinchus ventrilux ................................... MACROUR Coel 35.... 182
Genus Coryphaenoides .......................................... MACROUR Cory.......183
Key to species ..................................................................183
List of species ................................................................. 201
Coryphaenoides acrolepis ................................ MACROUR Cory 1 .....202
Coryphaenoides anguliceps ............................. . MACROUR Cory 2......203
Coryphaenoides ariommus ............................... MACROUR Cory 3 .....204
Coryphaenoides armatus ................................. MACROUR Cory 4......205
Coryphaenoides carminifer ............................... MACROUR Cory 5 ....206
Coryphaenoides cinereus ................................ MACROUR Cory 6 ......207
Coryphaenoides delsolari ................................ MACROUR Cory 7 .....209
Coryphaenoides guentheri ............................... MACROUR Cory 8 .... 210
Coryphaenoides longifilis ................................ MACROUR Cory 9 .... 211
Coryphaenoides macrocephalus .......................... MACROUR Cory 10 .... 212
Coryphaenoides marginatus ............................. MACROUR Cory 11 ... 214
Coryphaenoides mexicanus .............................. MACROUR Cory 12.....215
Coryphaenoides nasutus ................................. MACROUR Cory 13....216
Coryphaenoides rupestris ................................MACROUR Cory 14.... 217
Coryphaenoides serrulatus ............................... MACROUR Cory 15.....219
Coryphaenoides subserrulatus ........................... MACROUR Cory 16.....220
Coryphaenoides zaniophorus .............................MACROUR Cory 17 ....221
Genus Cynomacrurus ........................................... MACROUR Cyno ...... 223
Cynomacrurus piriei ..................................... MACROUR Cyno 1 ... . 223
Genus Hymenocephalus ........................................ MACROUR Hym ....... 224
Key to species ...................................................................225
List of species ............................................ 230
Hymenocephalus italicus ................................. .......
Genus Lepidorhynchus .........................................MACROUR Lep ........231
Lepidorhynchus denticulatus ............................. MACROUR Lep 1 ...... 232
Genus Macrourus .............................................. M A C R O U R M a c r ... .. ...233
Key to species ................................................................... 234
List of species ................................................................... 234
Macrourus berglax ...................................... M A C R O U R M a c r 1 ......235
Macrourus carinatus ..................................... M A C R O U R M a c r 2. ..... 236
Macrourus holotrachys ................................... M A C R O U R M a c r 3 ..... 237
Macrourus whitsoni ....................... .............M A C R O U R M a c r 4 ......238
Genus Malacocephalus ......................................... M A C R O U R M a l ........ 240
Key to species ................................................................... 240
List of species ................................................................... 241
Malacocephalus laevis .....................................M A C R O U R M a l 1 .......242
Malacocephalus occidentalis ..............................M A C R O U R M a l 2 .......243
Genus Mataeocephalus ......................................... M A C R O U R M a t ........ 244
Key to species ................................................................... 244
List of species ................................................................... 245
Mataeocephalus acipenserinus ............................M A C R O U R M a t 1 .... ...246
Mataeocephalus tenuicauda ...............................M A C R O U R M a t 2 .......247
viii
Genus Nezumia .................................................. M A C R O U R N e z ........ 248
yeKt o Nezumia groups ............................................................... 248
yeK o tseicepso f Nezumia by groups ................................................... 249
tsiL foseiceps ........................................................................ 263
Nezumia aequalis ..........................................M A C R O U R N e z 1 ....... 264
Nezumia atlantica...........................................M A C R O U R N e z 2 ....... 265
Nezumia bairdi..............................................M A C R O U R N e z 3 ....... 266
Nezumia brevibarbata ......................................M A C R O U R N e z 4 ....... 267
Nezumia condylura .........................................M A C R O U R N e z 5 ....... 269
Nezumia convergens........................................M A C R O U R N e z 6 ....... 270
Nezumia duodecim ......................................... M A C R O U R N e z 7 ....... 271
Nezumia latirostrata ....................................... M A C R O U R N e z 8 ....... 272
Nezumia liolepis.............................................M A C R O U R N e z 9 ....... 274
Nezumia loricata ........................................... M A C R O U R N e z 1 0 ..... 275
Nezumia micronychodon....................................M A C R O U R N e z 1 1 ...... 276
Nezumia milleri.............................................M A C R O U R N e z 1 2 ...... 278
Nezumia orbitalis...........................................M A C R O U R N e z 1 3 ...... 279
Nezumia propinqua .........................................M A C R O U R N e z 1 4 ...... 280
Nezumia proxima............................................ M A C R O U R N e z 1 5 ...... 281
Nezumia pudens.............................................M A C R O U R N e z 1 6 ...... 282
Nezumia pulchella..........................................M A C R O U R N e z 1 7 ...... 283
Nezumia sclerorhynchus....................................M A C R O U R N e z 1 8 ...... 284
Nezumia stelgidolepis.......................................M A C R O U R N e z 1 9 ...... 286
Genus Sphagemacrurus ......................................... MACROUR Spha ...... 287
Key to species ...................................................................... 287
tsiL foseiceps ...................................................................... 288
Sphagemacrurus grenadae ................................ MACROUR Spha 1 ..... 289
suneG Trachonurus ............................................... MACROUR Tra ........ 290
evitatneT tsil foseiceps .............................................................. 290
Trachonurus sulcatus ...................................... MACROUR Tra 1 ...... 290
suneG Ventrifossa ............................................... MACROUR Vent ...... 292
Key to adults of genus ............................................................. 292
List of species ...................................................................... 296
Ventrifossa atherodon ................................... MACROUR Vent 1 ..... 296
Ventrifossa ctenomelas .................................. MACROUR Vent 2 ...... 297
Ventrifossa divergens ................................... MACROUR Vent 3 ...... 299
Ventrifossa garmani ..................................... MACROUR Vent 4 ...... 300
Ventrifossa macropogon ................................ MACROUR Vent 5 ...... 301
Ventrifossa macroptera .................................. MACROUR Vent 6 ...... 303
Ventrifossa misakia ...................................... MACROUR Vent 7 ..... 304
Ventrifossa mucocephalus ............................... MACROUR Vent 8 ..... 305
Ventrifossa nasuta ....................................... MACROUR Vent 9 ...... 306
Ventrifossa nigrodorsalis .............................. MACROUR Vent 10 .... 307
Ventrifossa nigromaculata .............................. MACROUR Vent 11 ..... 309
Ventrifossa petersoni .................................... MACROUR Vent 12 ..... 310
Subfamily Macrouroidinae .................................................................. 311
Key to gener a ......................................................................... 312
Subfamily Trachyrhyncinae ................................................................... 312
Key to genera ............................................................................. 313
List of specie s ............................................................................313
suneG Trachyrincus .............................................. MACROUR Tra ......... 313
Key to species ......................................................................314
Trachyrincus helolepis .................................... MACROUR Tra 2 . ...... 315
Trachyrincus scabrus ..................................... MACROUR Tra 1 ........ 316
Trachyrincus villegai ..................................... MACROUR Tra 3 ........317
ix
2.6 FAMILY MELANONIDAE ................................................ M E L A N . .. .. .. ... .. .. ..318
Genus Melanonus .............................................. MELAN Melan ........ 318
2.7 FAMIL Y MERLUCCIIDAE .............................................. M E R L U ... .... ... ... .. 319
Key to genera ........................................................................... 320
Genus Lyconus ..................................................M..E.R.L..U. .L.y.c.o........... 322
List of species .................................................................... 322
Genus Macroronus ..............................................M E R L U M a c r u .. . . . . . . . .322
Key to species .................................................................... 323
List of species ..................................................................... 323
Macruronus capensis ...................................... M E R L U M a c r u 1 . . . . . . . 324
Macruronus magellanicus ................................. M E R L U M a c r u 2 . . . . . . 325
Macruronus novaezelandiae ............................. M E RL U M a cr u 3. . . . . . . 326
Genus Merluccius............................................... M E R L U M e r lu ......... 327
Key to species .................................................................... 328
List of species ..................................................................... 330
Merluccius albidus........................................ M E R L U M e rl u 2 ........ 330
Merluccius angustimanus ................................. M E R L U M e r l u 5 ....... 331
Merluccius australis ...................................... M. E R L U M e rl u 6 ........ 332
Merluccius bilineari s ...................................... M E R L U M e rl u 7 ........ 334
Merluccius capensis ...................................... M E R L U M e rl u 8 ........ 335
Merluccius gayi........................................... M E R L U M e rl u 9 ........ 336
Merluccius hubbsi ........................................ M E R L U M e rl u 1 0 ....... 337
Merluccius merluccius .................................... M E R L U M e r l u 1 ........ 339
Merluccius paradoxus..................................... M E R L U M e rl u 1 1 ....... 340
Merluccius polli ............................................ M E R L U M e rl u 4 ........ 341
Merluccius productus..................................... M E R L U M e rl u 1 2 ....... 342
Merluccius senegalensis .................................. M E R L U M e r l u 3 ........ 343
Genus Steindachneria ........................................... M E R L U S te i n . . . . . . . . . . . 345
Steindachneria argentea ................................... M E R L U S te i n 1 . . . . . . . . . 345
2.8 FAMILY MORIDAE ...................................................... M O R . . . . . . . . . . . . . . .. . . 346
Key to genera .......................................................................... 347
Tentative list of genera ................................................................... 351
Genus Antimora ................................................. M O R A n t . . . .. . . . . . . . . .352
Key to species ..................................................................... 352
Antimora microlepis ....................................... M O R A n t 2 . . . . . . . . . . . . .353
Antimora rostrata ..........................................M O R A n t 1 . . . . . . . . . . . . .354
Genus Auchenoceros ............................................ M O R A u c h . . . . . . . . . . . . . 355
Auchenoceros punctatus .................................. M O R A u c h 1 . . . . . . . . . 355
Genus Austrophycis ............................................ MOR Aust ............ 356
Austrophycis marginata .................................. MOR Aust 1 .......... 356
Genus Eeyorius ................................................. M O R E e y . . . . . . . . . . . . . 357
Eeyorius hutchinsi ........................................ M O R E e y 1 . . . . . . . . . . . . 357
Genus Gadella ...................................................M O R G a d e l . . . . . . . . . . . 358
List of species ..................................................................... 358
Gadella imberbis .......................................... M O R G a d e l 1 . . . . . . . . . . . 358
Gadella maraldi ............................................M O R G a d e l 2 . . . . . . . . . . 359
