Table Of ContentArachnol.Mitt.31:40-45 Nürnbergjuni 2006
Further faunistic notes on Cozyptila and Xysticusfrom Turkey
(Araneae,Thomisidae)
DmitriV.Logunov & Hakan Demir
Abstract:NinerecentlydescribedorpoorlyknownspeciesofthethomisidgeneraCozyptilaLehtinen&Marusik,
2005andXysticusC.L.Koch,1835arereportedfromTurkey.Fivespecies,Cozyptilablackwalli(Simon,1875),C.thaleri
Marusik&Kovblyuk,2005,XysticusbacurianensisMcheidze,1971,/.thessalicoidesWunderlich,1995andX.xerodermus
Strand,1913,arenewrecordsfortheTurkishspiderfauna.Twospec\es,X.bacurianensisar\diX.xerodermusareillus-
trated and a distribution map is providedfortheformer.Afewadditional recordsaregiven forGreeceand
theCaucasian countries,ofwhichX.bacurianensisis newforAzerbaijan.
Keywords:Caucasus,crabspiders,faunistics,MinorAsia,newsynonymy.
&
The thomisid fauna ofAsia Minor is notyet fully genera Cozyptila Lehtinen Marusik, 2005 and
known.According to the most recent checklistby Xysticus C.L. Koch, 1835, ofwhich five are new
TOPQU et al. (2005), the Turkish thomisid fauna records fortheTurkish spiderfauna.The newdata
comprises56validspecies,mostofwhichareknown allowus to significantlyclarifythe distributions of
from a single or just a few localities. Significant the species in question.
contributions to our knowledge of the Turkish Specimens for this studywere borrowed from,
Thomisidaewere made by KAROL (1966a, 1966b, or are distributed among the following museums:
1966c, 1966d, 1968),whorecorded 18 species(five HECO=HopeEntomologicalCollection,Oxford,
newtoscience),mostlyfromthevicinitiesofAnka- UK(Mr Hogan);IBPN= InstituteforBiological
J.
ra. Unfortunately, four ofher newspecies are now ProblemsoftheNorthFEB RAS,Magadan,Russia
regarded as invalid.ThreeXysticusspecies (X. sislii (DrY.M.Marusik);LNMC= LiverpoolMuseum,
Karol, 1966; X. turcicus Karol, 1966; and X.pelini NationalMuseums Liverpool, Liverpool,UK (Mr
Karol, 1968) proved to be junior synonyms ofX. G. Night); MBCG= Museo Civico di Scienze
thessalicusSimon,1916(seeWUNDERLICH 1995:p. Naturali “Enrico Gaffi”, Bergamo, Italy (Dr P.
X MMUM=
752) andone jezequellKarol, 1966)was shown Pantini); Manchester Museum, Uni-
(
UK
to be synonymous with X. gymnocephalus Strand, versity ofManchester, Manchester, (Dr D.V.
NHMW=
1915 (ASSI 1986: p. 45). Furthermore, having ex- Logunov); Naturhistorisches Museum,
NUAM=
amined the original description and a rathergood Wien, Austria (Dr Gruber); Arach-
J.
figure ofOzyptilaankarensisKarol, 1966 described nology Museum ofthe Nigde University, Nigde,
fromasingle 8 (seeKAROL 1966c:fig. 1,cf.UTO- Turkey (Dr A. Top£u); SMFM= Naturmuseum
CHKIN 1960: fig. 12 and ROBERTS 1995: p. 166), und Forschungsinstitut Senckenberg, Frankfurt
weareoftheopinionthatthisspeciesislikelytobe am Main, Germany (DrP.Jäger).
ajuniorsynonymofO.praticola(C.L.Koch,1837), Completereferencelistsforeachspeciesarenot
acommon Palaearctic species.Afinaldecisionhas provided, as theycanbe obtainedfromPLATNICK
beenpostponeduntilthe holotype ofO. ankarensis (2006). Here, under the heading 'Identification',
canbe found and re-examined. wereferonlyto one ortwoworks thatwe consider
The main aim ofthis short paper is to provide mostuseful forthe identification ofthe species.
newfaunistic datafornine thomisid species ofthe
Cozyptilablackwalli(Simon, 1875)
Identification:MARUSIKetal. (2005).
Dmitri V. LOGUNOV:The Manchester Museum,The University Materialexamined: Turkey: Id39 (NHMW), Kizil-
ofManchester,Oxford Road,ManchesterMl39PL,UK.E-Mail: m
[email protected] cahaman(40°28’N,32°37'E),1000-1200 a.s.l,thicket
&
Hakan DEMIR: Department of Biology, Faculty of Educati- offruit trees,27.-28.05.1967, leg.J. Gruber, F. Ressl
on and Art,Gazi University, 06500 Ankara,Turkey. E-Mail: A. Radda.
[email protected]
CozyptilaandXysticusfromTurkey 41
According to MARUSIK et al. (2005: map 1), this SHARENKO 1979). Our findings from Azerbaijan
species is distributed in Europe.The localityofC. and Turkey are new records for these countries,
blackwalliinTurkeyisitseasternmostrecordandis andtheysignificantlyextendourknowledgeofthe
a newrecord fortheTurkish spiderfauna. distribution ofthis rare species (Fig. 7).
AmongstthePalaearcticspecies,A.bacurianensis
&
CozyptilathaleriMarusik Kovblyuk, 2005 stands alone due to theunique conformationofits
Identification:MARUSIKetal. (2005). copulatory organs in both sexes (Figs 1-6), espe-
Materialexamined: Turkey: 39 (NHMW), ca 12 km ciallyofthe embolus (Fig. 3).The mostcloselyre-
SSWofAdapazan (40°45'N, 30°23'E), ca 100 m a.s.l., latedspeciesseemtobeX. embrikiKolosväry, 1935
oak forest with Carpinus, Tilia, Ligustru&m, Crataegus, known from East Europe (KOLOSVÄRY 1935) to
e1tc9.,(N14H.M06W.)19,67c,al1e5g.kJm. NGrEuboefrA,baF.ntRseeses,lAbanAt.MRtasd.,dac;a East Kazakhstan (MARUSIK & LOGUNOV 1995:
900ma.s.l., Quercus-Fagus-Abiesforest,25.05.1967,leg. fig. 40) and X. gymnocephalus Strand, 1915 from
J.Gruber,F.Ressl&A.Radda;2$ (NHMW),Tekketal, Minor Asia and the Levant (LEVY 1985: figs
SWAk^ehir(38°21'N,31°24'E),1100-1200ma.s.l.,pine 162-164, ASSI 1986). Males ofX. bacurianensis
forestwithQuercuscoccifera,Corylus etc.,11.06.1967,leg. differ from those ofboth related species in having
,
J. Gruber,F. Ressl&.A. Radda. a massive embolus, which is twice as long as in
ThisspecieswasrecentlydescribedfromtheCrimea the aforementionedspecies;itis stronglyextended
(Ukraine) byMARUSIKetal. (2005: map 1).Thus, laterally and rests on the finger-shaped tutaculum
theoccurrencesofC.thaleriinTurkeyrepresentthe (Figs 1,3).The 9 ofX. bacurianensishas a strong
southernmost records ofits distribution, in addi- transverse membranous duct ofthe spermathecae
tion to being a new species record for theTurkish (Fig. 6), which is absent from this structure in X.
spiderfauna. gymnocephalus', the 9 ofX. embrikiremains poorly
It is also worth noting that together with two known.
aforementioned species (blackwalli and thaleri) all
three described Cozyptila species are now known Xysticuscristatus(Clerck, 1757)
from Turkey. The third species, C. guseinovorum Identification:AZARKINA&cLOGUNOV(2001),JANT-
Marusik&cKovblyuk, 2005,was reliablyrecorded SCHER(2001).
from Hendek-Gumusova (41°21'N, 41°27'E) and Material examined: Turkey: 193juv.9 (HECO)m,
Trabzon, Zigana (40°36'52"N, 39°21'47"E), 2200
Yamanlar Dagi Mt. (ca 38°33'N, 27°8'E) (see &
a.s.l.,understones onherbs shortgrass (herbhillside
MARUSIK et al. 2005). The three other records andtop),22.08.1958,leg. G.Lampell.
byMARUSIK et al. (2005) were made from single This is a common Euro-Siberian temperate spe-
immaturefemales and are therefore nottakeninto cies (Azarkina&Logunov2001,Jantscher
account here.
2001) previouslyrecordedfromTurkey(TOP£Uet
al. 2005).
XysticusbacurianensisMcheidze, 1971
Identification: OVTSHARENKO (1979), present work
XysticuskochiThorell, 1872
(Figs 1-6).
Material examined: Turkey: 8612$ljuv.6 (HECO), Identification: LEVY(1979, 1985).
Trabzon,Zigana(40°36'52"N,39°21'47"E),2200ma.s.l., Materialexamined:Turkey:19 (NUAM),£ankiriProv.,
under stones on herbs &,shortgrass (herb hillside and Korgun Distr., Kiyisin (40°49'60"N, 33°37'0"E), under
top),22.-24.08.1958,leg. G.Lampell. tree bark, 28.07.2005, leg. H. Demir; 19 (NUAM),
Comparativematerial:Georgia:3<529 (IBPN),Lago- Konya Prov., Yunak Distr., Be^ikli (39°32'24"N,
dekhi Reserve (41°49'N, 46°16'E), 2800-2900 m a.s.l., 34°22'15"E), under stones, 13.05.2005, leg. H. Demir;
2.03.1982,leg.Y.M.Marusik.Azerbaijan:19 (ZMUM), 19 (LNMC), Yozgat (40°01'N, 34°37'E), Bogazkale,
Gusary Distr., foothills ofBazardyuzyu (= Bazardüzü) Bashkent Motel, grassy hillside, Populus copse&with
ME.tsG.us(e4i1n°o1v3.'N, 47°51'E), 3000 m a.s.l., 9.08.2001, leg. RFealntuonnc;ul1u9s d(oLmNiMnaCn)t,, I1s4p.a0r5t.a1,9c9a4,4lekg.mSE. JoufddEgridCi.r
(37o52'10nN, 30°50'57"E), 8.07.1992, leg. S.Judd; 16
ThisisaninhabitantofthealpinebeltoftheCauca- (LNMC),Bolu,ca21 km E ofAk£akoca (41°06'27"N,
sianMts.,whichhasbeenknowntodatefromonlya 31°15'54"E),sparsemixedweedsonroadslope,wetfield
sainndgltehleocNaliWtyipnarGteoorfgtihae(CMaCuHcaEsIuDsZMEa1j9o7r1,(1O9V9T7-) w&itCh.JFuenlctuosn;ad1j6ace(nLtNtMoCs)tr,eacma,71k8.m05E.1o9f94K,üclüekg.geSz.bJeuldid,
42 D.Logunov&H.Demir
Adana (38°09,03"N,36°07'27"E),roadbankwithgrass (LNMC),ca12kmSWofSarica,Kayseri(38°08'59"N,
and herbs, and adjacent wet areas, 13.05.1994, leg. S. 35°22'44"E), herb-rich orchard with deep A’ horizon
Judd & C. Felton; 19 (LNMC), Konya, Sakyatan ofdecomposing litter, 11.05.1994, leg. S. Judd & C.
(37°50'25"N, 32°47'6"E), 10.07.1992, leg. S.Judd; 29 Felton; 29 (LNMC), ca 4 km S ofAhmetyeri, Sinop
Figs1-6:ThecopulatoryorgansofXysticusbacurianensisMcheiöze,1971 fromTurkey(Trabzon:Zigana):1,malepalp,ventral
view;2,ditto,retrolateralview;3,embolus,dorsalview;4,tibialapophysis,lateralview;5,epigyne;6,spermathecae.Scale
bar0.1 mm.
Fig.7:DistributionofXysticusbacurian-
ensisMcheidze,1971.Onedot
mayrepresentmultipleadjacent
localities.
35°
CozyptilaandXysticusfromTurkey 43
(41°49'38"N, 35°05'24"E),freshwaterswamp at side of XysticusthessalicoidesWunderlich, 1995
river,10.07.1995,leg.S.Judd8cC.Felton;Id (LNMC), Identification:WUNDERLICH (1995).
Zonguldak, Amasra (41°44'42"N, 32°23'12"E), waste Material examined: Turkey: Id (MMUM), Antalya
groundbyflats,18.05.1994,leg.S.Judd6cC.Felton;Id Prov., Kalkan (36°15'N, 29°24'E), from village and
(LNMC), Nev^ehir, Nev§ehir-Urgup, 11.07.1992, leg. local garrigue, 04.2003,leg. L. Cook; 29 (LNMC), ca
C.Felton; 19 (LNMC),Nev^ehir,ca5 kmEofUrgup 5 km E ofBallidag Gecidi, Kastamonu (41°37'52"N,
m
(38°37'53"N, 34°54'47"E), 12.07.1992, leg. C. Felton; 33°23T2"E), 1267 a.s.l.,pinelitter,Seneciodominant
1$ (LNMC), Kayseri (38°43'56"N,35°297ME),31 km with Malus 17.05.1994, leg. S.Judd 8cC. Felton; 59
southofDuvalli rockyoutcropwithEuphorbiasp. and (NHMW),,ca15kmNEofLakeAbant,nearBolu(Bolu:
, m
mixed scrub, 15.06.1993, leg. S.Judd 8cC. Felton; 19 40°44'22"N,31°36'42"E),AbantMts.,900-1200 a.s.l.,
(HECO),Trabzon,Hamsikoy(40°41T4MN,39°287"E), Quercus-Fagus-Abies forest, 26.05.1967, leg. J. Gruber,
1250 m a.s.l.,beatenfromblueflowerpatchonhillside, F.Ressl8cA.Radda;29 (HECO),Trabzon,Hamsikoy
m
23.08.1958,leg.G.Lampell. (40°4T14"N,39°287"E),1250 a.s.l.,underneedleand
Comparative material: Italy: 19 (MBCG), Emilia- leaflitter,26.08.1958,leg. G. Lampell.
Romagna, Piacenza, Bobbio (44°46'18"N, 9°23T1"E), Comparativematerial: Greece: Id (LNMC), Lesbos,
1.05.1957,leg.A.Valle. Vatera(39T2'13"N,26°10'53"E),Aphroditehotelwall,
X. kocht is a common Euro-Mediterranean-Cen- 20.04.1997,leg.C.Felton; Id (LNMC),Lesbos, Skala
tralAsianspecies,previouslyrecordedfromTurkey Kallonis(39°12'25"N,26T3'13"E),beachandalongriver,
23.04.1997,leg. S.Judd.
(ToP£U et al. 2005).
This species was described from Greece (Crete,
XysticusninniifusciventrisCrome, 1965 SantorinandGreekmainland)(seeWUNDERLICH
Identification:UTOCHKIN8cSAVELYEVA(1995). 1995).Thus,this is the firstrecord outside Greece
Materialexamined:Turkey: 19 (NUAM), Sivas Prov., and a newrecord fortheTurkish spiderfauna; the
Yildizeli Distr., §eren (40°58'N, 31°25'E), under tree recordsfromLesbosclarifythespecies’distribution
bark, 22.07.2005, leg. H. Demir; ldl9 (NUAM), in Greece.
Cankin Prov., ^abanözü Distr., Hüyük (37°577"N,
31°35'45"E), under tree bark, 29.07.2005, leg. H. De- XysticusthessalicusSimon, 1916
mir; AS (LNMC), Kayseri,Tahtali Daglari, ca 8 km E Identification: WUNDERLICH (1995), AZARKINA 8c
ofBakirdagi (38T3'0"N, 35°47'60"E), 13.07.1992,leg. Logunov
(2001).
C.Felton;4d19 (LNMC),Konya,Kizilören(41°16'N, Material examined: Turkey: Id19 (NUAM), Ko-
33°TE),9.07.1992,leg.C.Felton. nya Prov., Gencek (41°57'N, 34°17'E), under stones,
This is a Euro-Central Asian species, distributed 15.05.2005,leg.H. Demir; Id (NUAM), KonyaProv.,
eastward as far as western Mongolia (UTOCHKIN Doganhisar (38°8'N, 31°40'E), near Degi^tigin, under
8c SAVELYEVA 1995: map 1). This species has stones, 14.05.2005, leg. H. Demir; 19 (LNMC), fyel,
previously been recorded from Turkey (TOP£U ca 8.8 km N ofAnamur (36°08'N, 32°51'E), hillside
with scattered, mixed,lowscrub andherbs Matricaria
Ietraanl.(2O00N5O: s8ucbMXA.RnTinEnNiiS) a2n00d4)re;cebnottlhy aclosuontfrrioems c(lLovNeMr,Ct)hi,stYloezagnadtT(h4y0m°u0s1)',437".N0,5.3149°9347,'2le1g".EC).,(.FBeolgtaoznk;al1ed,,
represent the southernmost limits ofthe species’ Bashkent Motel 14.05.1994, leg. S.Judd 8cC. Felton;
distribution. 29 (LNMC),Silifke(36°22'40"N,33°56'4"E),Mersin,
CROME (1965) thoroughly examined the Eu- sanddunesystemwithscatteredscrubupto400mfrom
ropean and Asian material ofX. ninnii Thorell, sea, 10.06.1993,leg. S.Judd8cC.Felton.
1872 and described a new subspecies (X n.fusci- Comparativematerial: Greece: 19 (LNMC), Lesbos,
v(ceant4r3i°s3),2o'fN,w4h1i°c4h7'tEh)e,ttyhpeeNolorctahlitCyaulcieassuins,TeRubsesridaa. aSknadlaalKoanlglorniivesr,(3293.T024'.2159M9N7,,2le6g°.1S3.TJ3u"dEd);,2b9ack(LoNfMbeCa)c,h
TheTurkishspecimensexamineddefinitelybelong Lesbos, Vatera (39°01273"N, 26°10'53"E), Aphrodite
to this subspecies,which accordingtoUTOCHKIN hotelwall,20.04.1997,leg. C.Felton.
8cSAVELYEVA (1995) probablydeserves separate ThisisanEast-Mediterraneanspeciesknownfrom
species status. The European records ofX. ninni Greece, the Balkans (Croatia), Israel and Turkey
(see in PLATNICK 2006) need to be re-examined (WUNDERLICH 1995).The newrecords clarifyits
in orderto clarifythe taxonomic status and distri- distribution inTurkeyand Greece.
bution ofboth taxa. This problem is beyond the
scope ofthe current study.
:
44 D.Logunov&H.Demir
XysticusxerodermusStrand, 1913 with a wider one (Fig. 10).The latter is identical
Identification: LEVY(1976, 1985). tothefigureprovidedbyLEVY(1985: fig. 156).X.
Material examined: Turkey: 1$ (NUAM), Konya xerodermus belongs to the sabulosus species group
Prov., Seydifehir (37°25'N, 31°51'E), Rizebeli Pass, and seems to be closest to X. kempeleni Thorell,
understones, 15.05.2005,leg.H.Demir; 1$ (LNMC), 1872 (sensu LOGUNOV in press), but the female’s
Yozgat(40°01'N,34°37'E),Bogazkale,BashkentMotel,
grassyhillside,PopuluscopsewithRanunculusdominant, epigyneissmallerandtheinseminationductsshor-
14.05.1994, leg. S. Judd & C. Felton; 19 (LNMC), ter and do not form a loop (Fig. 9; see also LEVY
Yozgat(40°01'43"N,34°37'21"E),Bogazkale,Bashkent 1985:fig. 157).Inanycase,femalesinthesabulosus
&
Motel, 14.05.1994,leg. S.Judd C.Felton. grouparehardtodistinguishfromoneanother,and
X.xerodermusisapoorlyknownEast-Mediterrane- thereforemalesarerequiredtoconfirmorrejectour
anspecies,todatereportedonlyfromIsrael(LEVY identification.
1985).Thus, it forms a new species record for the Acknowledgements
Turkish spiderfauna.
We have identified this species on the basis of Wcoellewaigsuhest:oJeüxrpgreesnsGmryubdeerep(gNraHtiMtWud)e,toJtahmeesfolHloogwianng
thepublicationsofLEVY(1976,1985)only.Atleast (HECO), PeterJäger (SMFM),Yuri Marusik (IBPN),
twotypesoffemaleshavebeenfound,viz.onewith Guy Night (LNMC), Timo Pajunen (ZMHU), and
arelativelynarrowepigynal fossa (Fig. 8), another
Figs8-10:The $ copulatoryorgansofXysticusxerodermusStrand,1913fromTurkey(8-9,RizebeliPass;10,Yozgat):8,10,
epigyne;9,spermathecae.Scalebar0.1 mm.
CozyptHaandXysticusfromTurkey 45
Paolo Pantini (MBCG), for allowing us assess to the LOGUNOVD.V. (inpress): Notes onXysticuskempeleni
collectionsoftheirmuseums.DavePenney(Manchester, Thorell 1872 and two closely related spider species
UK) kindlyeditedthe Englishofthedraft. (Araneae,Thomisidae).-ActaArachnol.
Marusik Y.M., P. Lehtinen 8c M.M. Kovblyuk
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