Table Of ContentEEllyyttrraa,,TTookkyyoo,,NNeewwSSeerriieess,,11((22))::226633(cid:2)(cid:2)227711
December31,2011
Four Species of the Genus Leptoglypus from Japan
(Coleoptera, Bothrideridae)
Jun-ichi AOKI
3(cid:2)8(cid:2)12Nishi-Azabu,Minato-ku,Tokyo,106(cid:2)0031Japan
Abstract InadditiontotwoknownspeciesofthegenusLeptoglyphus,twomorenewspecies
werefoundfromJapan.OneofthemisdescribedhereunderthenameofLeptoglyphustanakaisp.
nov.,whichwasformerlyidentifiedasL.orientalisS=6GE.Alargenumberofspecimensoftrue
L.orientaliswerecollectedinthesouthernpartofJapan.Theothernewspeciesisdescribedhere
asL.kubotaisp.nov.
Two species of the genus Leptoglyphus have hitherto been known from Japan: Leptoglyphus
vittatusS=6GEandL.orientalisS=6GE. TheformerisveryrarespeciesonlyknownfromKyushu
and its adjacent islands, while the latter is often found in central part of Japan. However, my
identification of the latter species was found to be a mistake and it should be described as a new
species. On the other hands, number of specimens of true L. orientalis were collected from
Shimo-KoshikiIslandandYakuIsland.AnotherunknownspecieswasalsofoundinYakuIsland.
Thus, a total of four species of Leptoglyphus were ascertained from Japan and are described
below.
Leptoglyphus vittatus S=6GE
(Figs.1(cid:2)6)
LeptoglyphusvittatusS=6GE,1885,75;S6H6?>,1997,114,fig.4;AD@>,2009a,4,fig.6;2009b,106,Fig.
Body length: 3.2(cid:2)3.5mm.
Color: Castaneous or dark reddish brown.
Head. Eyes large, separated by 1.5times diameter of single eye. Antennae (Fig.2) 9-
segmented;relativewidths of segments: I(cid:2)II(cid:2)III(cid:3)IV(cid:3)V(cid:3)VI(cid:3)VII(cid:4)VIII(cid:4)IX; terminal seg-
ment IX forming distinct club divided by transverse line into two parts, with small constriction.
Pronotumnearlyaslongaswide,itsL/W0.97;lateralmarginswithweakangularityalittle
in front of middle (Figs. 1 and 3); anterior corners distinctly projecting anteriorly; posterior
cornersnearly right angled. Prosternal disc densely covered with oval foveolae, each with short
decumbent seta.
Elytranearlytwiceaslongasbroad,distinctlywiderthanpronotum(about1.2timeswide),
lateral margins gently swollen; elytral carina II ending shortly before reaching apical border of
elytron; III and IV jointed together apically and jointed again more apically with V (Fig.4);
carinae II and III bearing long setae (Fig.4). Even intervals with double rows of punctures.
Ventralside.Prosternalprocesswithdilatedapex.Labialmentum(Fig.6)alittlewiderthan
long,withconcaveanterior and posterior margins, bearing 4 long and 8 short setae in asymmet-
ricalposition;segmentIIoflabialpalpwith2longand4shortsetae;segmentIIIwith1longand
2 short setae. Prementum with angulate anterior margin, densely beset with thick setae. Male
genitaliapeculiarinshape,withapairofshorthorn-likeprocessesonapexandapairofsharpand
264 Jun-ichiAD@>
Figs.1(cid:2)6. Leptoglyphus vittatus S=6GE. (cid:3)(cid:3)1, Habitus; 2, antenna (right side); 3, outline of pronotum;
4, apical part of elytron (right side); 5, apical partofmalegenitalia;labiaum;6,labium.(Scalebars:1
mmforFig.1;0.5mmforFig.3;0.3mmforFig.4;0.1mmforFigs.2,5and6).
curved spines laterally (Fig.5).
Legs. Tibiae,segmentsIandIIdenselybearing setae on ventral side; segment III narrower
than I and II, bearing only a few setae.
Material examined. 1ex., Teuchi, Shimo-Koshiki Island, Kagoshima Prefecture, West
Japan, 17(cid:2)V(cid:2)1994, T. U:CD leg.; 1ex., Ohko Rindo, Yakushima Island, Kagoshima Prefecture,
Southwestern Japan, 27(cid:2)XI(cid:2)2002, Y. KJ7DI6 leg.
FourLeptogyphusSpeciesfromJapan 265
Leptoglyphus tanakai sp. nov.
(Figs.7(cid:2)12)
Leptoglyphusorientalis:AD@>&H>G6CD,2008,1,figs.1(cid:2)3;AD@>,2009b,104,fig.
Body length: 1.85(cid:2)2.50 (av.2.18)mm.
Color: Body castaneous; pronotal margins and edges of elytral carinae black.
Head. Anteriorclypealmargin straight; clypeus and frons with sharply pointed erect setae;
vertexwithdecumbent setae. Eye separated by 1.9 times diameter of eye. Antennae (Fig.8) 9-
segmented;relativewidths of segments: I(cid:2)II(cid:2)III(cid:3)IV(cid:3)V(cid:4)VI(cid:3)VII(cid:4)VIII(cid:4)IX; terminal seg-
ment IX forming distinct club divided by transverse line into two portions, larger basal portion
with a few setae and smaller apical portion densely covered with long and short setae and
constricted laterally.
Figs.7(cid:2)12. Leptoglyphus tanakai sp. nov. (cid:3)(cid:3)7, Habitus; 8, antenna (right side); 9, outline of prono-
tum; 10, apical aprt of elytron (right side); 11, malegenitalia;12,labium.(Scalebars:0.5mmforFig.
7;0.3mmforFigs.9and10;0.1mmforFigs.8,11and12).
266 Jun-ichiAD@>
Pronotumratherbroad(Fig.9),slightlywiderthanlong,itsL/W0.88(cid:2)1.00(av.0.97),with
nearly straight anterior margin and weakly convex lateral margins gently and smoothly curved,
widestinfrontofmiddle,showingveryinconspicuousangularity;anteriorcornersneverproject-
ing anteriorly, posterior corners nearly right angled. Pronotal disc densely covered with oval
foveolae each with long decumbent seta, almost twice as long as foveola.
Elytranearlytwiceaslongasbroad,slightlyswollenatmiddleoralittleposteriorly,itsL/W
1.89(cid:2)2.14(av.2.48);elytronwithsixcarinae;carina II strongest and carina IV weakest; carinae
II and III ending shortly before reaching apical border (III shorter than II); carinae IV and V
joinedtogetherapically(Fig.10)andreachingapexofelytron;cainaIIonlyslightlymoreraised
apicallythanotherportion,bearingminutesetae. Evenintervalswithdoublerowsofpunctures,
each with minute seta.
Ventral side. Prosternal process with dilated apex, constricted lateral margins and anterior
margin distinctly concave medially. Middle coxal cavities separated by 1/3 of cavity diameter;
surface of mesosternum and metasternum except on smooth median part covered with oval
faoveolae,eachwithfineseta. Ventritesshowingsurfacestructuresimilartothatofmetasternum;
intercoxal process of ventrite I very broad and trapezoid, wider than half width of ventrite;
ventriteItwiceasboardasventriteV.Labialmentumbecomingalittlewiderposteriorly;second
and third segments of labial palp each with two setae (Fig.12). Male genitalia (Fig.11)
lanceolate, with two pairs of setae at apex.
Legs. Femora with a pair of lobe-shaped appendages on ventroapical end. Tarsi parallel-
sided, with truncated rectangular apex.
Holotype:Mt.Wakakusa,NaraCity,NaraPrefecture,CentralJapan,15(cid:2)III(cid:2)2009,T.S6>ID
leg.;7paratypes(including1(cid:2)and1(cid:3)),thesamedataasholotype;1paratype,Mt.Wakakusa,
Nara City, Nara Prefecture, Central Japan, 13(cid:2)III(cid:2)2005, T. S6>ID leg.; 4 paratypes (including
1(cid:3)), Sasayakinokomichi, Kasugano-chˆo, Nara City, Nara Prefecture, Central Japan, 13(cid:2)III(cid:2)
2005,T.S6>IDleg.;2paratypes,NaraPark,Takahata-chˆo,NaraCity,NaraPrefecture,Central
Japan, 19(cid:2)II(cid:2)2005, T. S6>ID leg.; 1 paratype, Near Kasuga-Taisha, Takahata-chˆo, Nara City,
Nara Prefecture, Central Japan, 17(cid:2)I(cid:2)2009, T. S6>ID leg.; 1 paratype, Asebi, Nara Park, Nara
City,NaraPrefecture,CentralJapan,28(cid:2)I(cid:2)2006,[email protected](NSMT-I-C200200)
and3paratypes(NSMT-I-C200201(cid:2)200203)aredepositedinthecollectionofNationalMuseum
of Nature and Science, Tokyo.
Remarks. AD@> and H>G6CD (2008) and AD@> (2009) made an error in identifying this
species from Central Japan as Leptoglyphus orientalis GGDJK:AA:. Another species surely
consideredastrueL.orientaliswascollectedlaterinthesouthwesternpartofJapananddescribed
below. Thus, the species from Central Japan was described above as a new species.
Leptoglyphus orientalis GGDJK:AA:
(Figs.13(cid:2)18)
LeptoglyphusorientalisGGDJK:AA:,1906,117.
Body length: 1.70(cid:2)1.95 (av. 1.83)mm.
Color: Body castaneous; pronotal margins and apical parts of elytral carinae dark-colored.
Head. Anterior clypeal margin straight; eyes separated by twice diameter of eye. Antennal
clubdividedintotwoparts,apicalportionweaklyconstrictedbasally(Fig.14);Otherfeaturesas
in the proceeding species, L. tanakai.
FourLeptogyphusSpeciesfromJapan 267
Figs.13(cid:2)18. Leptoglyphus orientalis GGDJK:AA:.(cid:3)(cid:3)13, Habitus; 14, antenna (right side); 15, outine of
pronotum; 16, apical part of elytron (right side); 17, male genitalia; 18, labium. (Scale bars: 0.5mm
forFig.13;0.3mmforFigs.15and16;0.1mmforFigs.14,17and18).
Pronotum rather broad (Fig.15), slightly wider than long, its L/W 0.90(cid:2)0.98 (av. 0.95),
withslightlyconvexanteriormarginandweaklyconvexlateralmargins,showingveryinconspicu-
ous angularity at level a little in front of middle point; anterior corners only feebly projecting,
posterior corners nearly right angled. Prosternal disc densely covered with oval foveolae, each
with shorter decumbent seta shorter than twice length of foveola.
Elytra nearly twice as long as broad, mostly parallel-sided, its L/W 1.96(cid:2)2.09 (av. 2.02);
elytron with six carinae; carina II strongest and carina IV weakest; carinae II and III ending
shortly before reaching apical border of elytron; apical part of carina II strongly raised and
thickened, bearing rather long setae; carina III also weakly thickened apically, with rather long
setae;carinaeIVandVjoinedtogetherapicallyjustbeforereachingapicalborder(Fig.16). Even
intervalswithdoublerowsofpunctures;minutesetaearrangedinrowsalongcarinaeandrowsof
punctures.
Ventral side. Prosternal process with dilated apex, constricted lateral margins and straight
268 Jun-ichiAD@>
anterior margin only slightly concave medially. Middle coxal cavities close to each other,
separatedby1/5ofcavitydiameter;surfaceofmesosternumandmetasternumexceptonsmooth
medianpartsparselycovered with irregular-shaped oval foveolae, each with fine seta. Intercoxal
process of ventrite broad and trapezoid, far wider than ventrite V; ventrite I wider than twice
breadth of ventrite V. Labial mentum with straight lateral margins a little diverging posteriorly;
second segment of labial palp strongly large and swollen, with three long and three short setae;
third segment with two short setae. Male genitalia lanceolate, with two pairs of thick setae.
Legs. Similar to those of the preceding species.
Materialexamined:3exs.,Mt.Toimo-dake,YakushimaIsland,SouthwesternJapan,13(cid:2)IV(cid:2)
2010,Y.KJ7DI6leg.;10exs.,thesameplace,26(cid:2)V(cid:2)2010;3exs.,thesameplace,29(cid:2)VI(cid:2)2010,Y.
KJ7DI6 leg.; 1ex., Ikoino-mori, Kusukawa, Yakushima Island, Southwestern Japan, 11(cid:2)VI(cid:2)
2010, Y. KJ7DI6 leg.; 67 exs., Sesenoura, Shimo-Koshikijima Island, Kagoshima Prefecture,
Southwestern Japan, 4(cid:2)X(cid:2)2009, J. AD@> leg.; 3exs., Kawanagano, Satsuma-Sendai, Kagoshima
Prefecture, Southwestern Japan, 9(cid:2)V(cid:2)2008, Y. H>G6CD leg.
Distribution. Sumatra, Java and Japan (new record).
Remarks. Two Japanese species of Leptoglyphus, the central Japanese species and the
southwestern Japanese species, both are very similar to L. orientalis. However, the latter species
recently found from Japan is more close to orientalis, because it is more in accordance with
originaldescriptionoforientalisinthefollowingtwopoints:(1)someerectsetaeinapicalpartof
elytralcarinae(inparteapicalicarinarumaliquispiliseretis)and(2)bodylength1.5(cid:2)2mm. The
knowndistributionoforientalisisintropicalarea,SumatraandJava,anditismorereasonableto
identifythesouthwesternJapanesespeciescollectedinsubtropicalareaasorientalisthantodoso
thecentralJapanesespeciescollectedinwarm-temperateareainJapan,whichisdescribedhereas
a new species.
Leptoglyphus kubotai sp. nov.
(Figs.19(cid:2)24)
Body length: 2.10(cid:2)2.20 (av.2.16)mm.
Color:Bodyandlegscastaneous;pronortalmargin,basalandapicalpartsofelytraandapical
portion of femora dark-colored.
Head. Eyes separated by 2.0(cid:2)2.5(cid:2) diameter of eye. Antennal club divided into two parts,
apical portion constricted basally (Fig.20); ratio in width of antennal segments: I(cid:3)II(cid:3)III(cid:4)IV
(cid:4)V(cid:4)VI(cid:4)VII(cid:5) VIII(cid:5)IX. Other features as in L. tanakai.
Pronotum rather narrow, a little longer than wide (Fig.21), its L/W 1.06(cid:2)1.17 (av. 1.15),
with weakly convex anterior margin; pronotum widest at middle part, showing feeble angularity
on each side; anterior corners never projecting, but cut obliquely, posterior corners inconspicu-
ouslyangled. Pronotaldisccoveredwithelongateovalfoveolae,eachwith short fine seta nearly
as long as foveola.
Elytramorethantwiceaslongasbroad,ratiooflengthtowidth2.16(cid:2)2.33(av.2.25);elytron
with six carinae; carina II strongest and carina IV weakest; carina II almost reaching apical
borderandmoreorlessraisedandthickenedapically;carinaIIIjoinedtocarinaIVapicallyand
then again joined to V to reach apical border of elytron; apical 1/3 of carinae II, III and IV
bearing long erect setae (Fig.22). Even intervals with double rows of large punctures.
Ventralside. Prosternal process with delated apex, constricted lateral margins and anterior
FourLeptogyphusSpeciesfromJapan 269
Figs.19(cid:2)24. Leptoglyphus kubotai sp. nov. (cid:3)(cid:3)19, Habitus; 20, antenna (right side); 21, outline of
pronotum; 22, apical part of elytron (right side); 23, male genitalia; 24, labium. (Scale bars: 0.5mm
forFig.19;0.3mmforFigs.21and22;0.1mmforFigs.20,23and24).
marginweaklyconcavedmedially. Middlecoxalcavitiesclosetogether,separatedby1/5(cid:2)1/8of
cavitydiameter;surfaceofmesosternumandmetasternumexceptonsmoothmedianpartsparsely
coveredwithveryelongateovalfoveolae,eachwithshortfineseta. Intercoxalprocessofventrite
Ibroadandtrapezoid,farwiderthanhalfofventritewidth;vantriteIlessthantwice breadth of
ventriteV. Labialmentumwithparallellateralmargins;secondandthirdsegmentsoflabialpalp
withtwoandthreesetae,respectively. Malegenitaliapeculiarinshape,apexwithapairofshort
horns rounded at tip and a pair of sharp and curved spines laterally (Fig.23).
Holotype(sexundetermined):Mt.Toimo-dake,YakushimaIsland,SouthwesternJapan.29(cid:2)
VI(cid:2)2010, Y. KJ7DI6 leg.; 2 paratypes (including 1(cid:2)), Mt. Toimo-dake, Yakshima Island,
SouthwesternJapan,21(cid:2)III(cid:2)2010,Y.KJ7DI6leg.;1paratype((cid:3)),thesameplace.13(cid:2)IV(cid:2)2010,
270 Jun-ichiAD@>
Y. KJ7DI6 leg.; 2 paratypes ( including 1 (cid:2)), the same place, 26(cid:2)V(cid:2)2010, Y. KJ7DI6 leg.; 3
paratypes (including 1(cid:2) and 2(cid:3)), the same place, 29(cid:2)VI(cid:2)2010, Y. KJ7DI6 leg.; 1 paratype,
Ikoino-Ie, Kusugawa, Yakushima Island, Southwestern Japan, 11(cid:2)VI(cid:2)2010, Y. KJ7DI6 leg.; 1
paratype, Issou, Yakushima Island, Southwestern Japan, 4(cid:2)XII(cid:2)2011, Y. KJ7DI6 leg.; 1 para-
tyepe, Hinoyama Rindo, Miyakejima Island, Izu Islands, Central Japan, 3(cid:2)23(cid:2)VIII(cid:2)2005, H.
M6@>=6G6leg.Holotype(NSMT-I-C200204)and2paratypes(NSMT-I-C200205,200206)are
deposited in the collection of Natioal Museum of Nature and Science, Tokyo.
Key to Four Japanese Species of the Genus Leptoglyphus
1(2) Elytral carinae II strongly elevated apically, provided with shot setae; lateral margins of
pronotumsmooth.Southwestern Japan (Kyushu, Shimo-Koshiki Island and Yakushima
Island).(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3) L. orientalis GGDJK:AA:
1(2) Elytral carinae II not or weakly elevated apically.
3(4) Apical half of elytral carinae II with short setae; lateral margins of pronotum nearly
smooth,onlyslightlyangulate. CentralJapan.(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3) L. tanakai sp. nov.
4(3) Apical half of elytral carinae II with long setae; lateral margins of pronotum more or less
distinctly angulate.
5(6) Pronotum as long as wide or a little wider than long, with anterior corners distinctly
produced; elytra twice as long as broad (L/W(cid:4)2.00). Southwestern Japan (Kyushu,
Shimo-Koshiki Island and Yakushima Island).(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3) L. vittatus S=6GE
6(5) Pronotum longer than wide, with anterior corners obliquely cut, without anterior projec-
tions; elytra longer than twice the breadth (L/W(cid:4)2.24(cid:2)2.25). Body length 2.10(cid:2)2.16
mm. SouthwesternJapan(YakushimaIsland)andCentralJapan(MiyakeIsland).(cid:3)(cid:3)
(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3)(cid:3) L. kubotai sp. nov.
Acknowledgement
ThenewspeciesLeptoglyphustanakaiwascollectedbyMr.IsamuT6C6@6(NaraCity)for
thefirsttimeinNaraCityandlatercollectedalsobyMr.TakumiS6>ID(AmagasakiCity). They
kindly o#ered a large number of specimens for my study. Mr. Yoshinori KJ7DI6 (Yakushima
Island) made thorough investigation in wide areas of Yakushima Island and succeeded in
collecting a large number of colydiid beetles including Leptoglyphus orientalis and L. kubotai.
Hence the names of the two species are dedicated to Mr. I. T6C6@6 and Mr. T. KJ7DI6. Mr.
YukihikoH>G6CDgavemespecimensofL.orientaliscollectedinKyushu. Iexpressmyheartfelt
thanks to them.
(cid:2) (cid:3)
(cid:5)(cid:6)(cid:7)(cid:8)(cid:9) (cid:10)(cid:11)(cid:12)(cid:13)(cid:14)(cid:15)(cid:16)(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)(cid:22)(cid:23)4(cid:24) (cid:25)(cid:26)(cid:27)(cid:28)(cid:29)(cid:27)(cid:30)(cid:20)(cid:31) !(cid:16)(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)"#$ (cid:3)(cid:3) (cid:10)(cid:11)%(cid:12)
&’(cid:13)(cid:14)(cid:15)(cid:16)(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)(cid:22)Leptoglyphus%((cid:13)(cid:14)(cid:15)(cid:16)(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)L. vittatus S=6GE)*+ (cid:19),(cid:18)(cid:16)(cid:17)(cid:18)(cid:19)
(cid:20)(cid:21)L. orientalis GGDJK:AA:(cid:23)2(cid:24)-./0123-4 56(cid:10)(cid:11)7/8(cid:23)orientalis9:;<0’(cid:24) (cid:25)(cid:13),
(cid:13)(cid:13)(cid:14)(cid:15)(cid:16)(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)4 =># -?@<04 ABCDEF9&’EG(cid:10)(cid:11)%HI&’(cid:19),(cid:18)(cid:16)(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)(
orientalisJ(KL4=(cid:24)9&MNO(cid:23)JP’Q9-RST4(cid:19),(cid:18)(cid:13)(cid:14)(cid:15)(cid:16)(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)Leptoglyphustanakai
sp. nov.9T1UVT3$ W34 56(cid:10)(cid:11)7/(</%X(cid:24)(cid:23)=(cid:24)-YZ<04 (cid:16)(cid:17)(cid:13)(cid:14)(cid:15)(cid:16)(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)
FourLeptogyphusSpeciesfromJapan 271
Leptoglyphuskubotaisp.nov.(cid:2)(cid:3)(cid:4)(cid:5)(cid:6)(cid:7)(cid:5)(cid:8)(cid:9) (cid:10)(cid:11)(cid:12) (cid:13)(cid:14)(cid:15)(cid:16)4(cid:17)(cid:18)(cid:19)(cid:20)(cid:21)(cid:22)(cid:23)(cid:2)(cid:15)(cid:24)(cid:25)(cid:12) (cid:23)(cid:26)(cid:27)4(cid:17)(cid:28)
(cid:29)(cid:30)(cid:31)(cid:6)(cid:7)(cid:21)(cid:22)(cid:2)(cid:2) (cid:15)(cid:12) !"#(cid:28)(cid:30)(cid:5)(cid:8)(cid:9)
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