Four new species of the Triplophysa stoliczkai-complex from China (Pisces: Cypriniformes: Balitoridae)

15 Pages·2001·3.1 MB·

by  Prokofiev, A M

© ZooloIgnisctSaitlt. uPteet,2e 0r0s1b urg,
Founr.s epwe ocfit ehTser iplosptohyslai czlwi-complex
fromC hi{nPai sCcyepsr:i niBfaolrimteosr:i dae)
A.MP.r okofiev
ProkAo.fi2Me0.v0F,1o .nu ersw p eoctfih eTresi p lospthoylsiac zkfraoiCm-h cionmap lex
(PisCcyepsr:i nBiafolrimteZosor:oi sydsateeR)mo:as ts1iic0r;:a(a 1,) : 193-207.
Founre swp ecoifte hgsee nTruisp lop(hT.qy qsuaa ecaeT.r huilaelamT.ea ,r i,
rossopereagnrT.di a nlaetxoarsnupdrmnpa ..efr) o mC hiarneda e scrAiltblhe eds.e
spebceiletoston h Tges. t o/iczkai-cdoimapglonetfoxh s.ciiAossrm epivlgsiei sxve edn .
ValidoifttT.ey n (uDiasT.y do )rs,o no(tKaetasa snT.lds e tre)n( uHrear zeibnsrs iteefliyn )
discussed.
A.MP.r okoLfiaebvo,ro afFt iosrahyne Fdsi sh-like PVaelrateeobnrtIaontlsetosig,­i cal
tutPer,ofso yuzunl1a.2y M3ao, s cIo7Iw6 4R7u,s sei-am;av inli:k [email protected]
Introduction Turd(a1k9o14v96 5,B4 e)rd.ge scarc ilboesde ly
relbautwtee ddl ils tNienmcatc hleaicluussn igri
Thgee nTruisp loRpehnyds1aa9h 3lo3,n, oe f fromK arak·Lo alkT eu.r daikn1o 9v4fi 6r rset­
thlea rigtneh sseut b famNielmya checiolmi­vniaseie,nd t rasvpaerciiaofibfNci e lmiatcyh eilus
prises vaablsoipudet cm io9eso0stfw , h ich satroel aincldzat ke(arii1 n 9 5d4i)s tinognuei shed
distriitnbh Hiuegt hAe sdi baatsiOicnn oesft . h em orN.e s toliczskpaeicfr-ioreemtsl hS aeyt re­d
widedliys trsipbeucotifete dhsg e e niusT.s DaryaRN.i vceorn,i pStuebrsuesqC.eh nitnleys,e
stol(iSctzekianid acIhtrnsae ncrgo,ev 1e8ra6su6 t)hd.oe rssc rfuirtbhenedert w a frxoam C hina
al artgeer rfriotmot rhuyep preera cohfe [sT· t anggu(lZahe1un9,s8 Ti2x s)i ;n gshanensis
AmudaSryyrad,a Crhyuaan ,Id n druisv e(rYsan &g X i1e9,8 T3 c)a;k aeCnas&oi Z sh u,
throNu Ignhda inaid n nbears oifKn osk ono1r9,8T 8g ;e rzeeCnas&oiZ sh u1,9 8a8no dt hers
QingahnaQdia idtaohm e waadetrosft he( sbeeel owwh)i]sc,eh e mb ect loo tsoTe
Hwanga-riChdho a ngjriiavniegNnr W sC hinsat ol(iZchz1uk9,a8Y i2a ;n& gX i1e9,8 C3a;&o
(Turd1a9k4o16v9,,5 B 2e;r 1g9,4 J9a;y araZmh,1u 9,8 Wu8 ;&W u1,9 8Z8h;a& oW an1g9,8 8;
198M1e;n on,Z1h9u8,71;9 89). WiDtihn1ig9n,9 t3h)i.s large
terrTi·s t tooryl,si hcozawkg sar iev aatr iabiTlhiaetyu trheovrit shceeod l leocftliooaoncf h es
anwda dse scarsian b uemdbo efnr o misnpael­t hZeo oloIgnisctSaitlt. uPbteuet(r,eZg rI saS nPd)
ci(eC so biutriasn osKceospsu1ls8e 7rC2,.; revetahle4enr dee sw p eccileosrs eellaybt uetd ,
e/egans1K8e7Nes4ms;al cehre,ti e/nDuuasiy s,n oitd enttoT.i sctaoll iTchzenkseaeswi p .e cies
187N.6 ddrs;d notaKteusss1 l8e7N.r9 s,;t enu(rT.ua sq uaecaT.e hriuallemT.aa rero,is ,s opere­
Herze1n8s8tN.8efe;idts nc,h enAk.No iakeo lsgrkiin,a taonrT.du a ml exasnpdnpr..aa) er e de­
190N.3 c;u eljµAennisk1ii9sn0 w,5h )i,wc ehr es criibnte hpder espeanpTteh rce.o lleocft ion
synonywmiitTzhs e tdo /biycB zekr(ag1i 9 49t)h.Ze o oloMguisceauMlmo ,s cUonwi versity
Ber(goc pi.at l.ss)uo g getshsteye ndo noyfmyT (ZMMwUa)es x amianswe edlO lu.ar n alysis
stolwiicNetzmhkaa ic helihlaRusesag e1a 9n0,5 s howtehda futrt hesrtd uyo fC hniese
anN.d c hopHroar1ia9 ,2 b2u,tt h etsweto a xan emachreeilltaitoTne ssdt oliicnsze kaeid ed,
seetmob e v al(iTcdh aenatgl 1 .9,6J 2ay;ar am,b ecasuosmieed entifiocflat oiaocanhsT.se s
19)8T.1h fier isntv estoitfgh Taes t tioolnisc sztkaoil­ikcnzokwiannli i teraarutenu creer·t ain.
comlpew xe rmea de Bebyr(1 g 9 3a1n)d Turkdoa(v1 95n2o)t tehdas to mCeh inese
194 A.M.P rokoNefiwes vp:e coifeTrsi plo•p hZyOsOaSYRSOTS.S IVCoA1l 0.
populoatfth isiopsne smc ayir eesp rdeissetnitan ncadtn tenraisooaprle nciolnsgyess lp actehde,
subspLeickieetwshis.esp ee,c ifrmoeTmn ssa ngp­osteenrcilotorhas ene tse Lriimpoosrr o.elr ess
pod escarnifidbg eudrb yeTc dh aenagtl (1. 9 62·fu, r rolwoewdle,iwr pi mtehn ltoab(les so metimes
p6.2fi5g;1. - a3sN) e machsetiolluaiscc tzukw­aeia kelxyp rebsuswtei dt)hl,oa uttle orbaels ;
albleyl toaonn ug n desscpreicobinete dhbs ea ­uppleiwrpi thloatueptrr aolgla toinaobnot vhee
siosft heciorl orpartoipoonra,tib irlo andsdt,eh ripraodifb r a rbUeplpmsea.rr ogfil no wlpeir
capsaunildne t emsotripngheyo .l o noctl otsome a rgoifln o wjearwP .r ocessus
ManCyh inaeustehc oornss iTd deorresdod ­entioforr mokineus ep lpj earaw b sDeonrts.a l
notaantTda s tenausdr ias tinct spfienoc riiseglsii ngp( hoZtshltutye,om r iidobroo red qy­u i
198129,8Ya 9n;g& Xi1e9,8 Z3h;a& o W ang,d isttsoan noatun ctda ubdaa(sel ex cTet petn uis),
198e8t,cO .ue)rx. a minattwsioyo nnty opofef sw it6h-( 9u su7a-lb8lr)ya nrcahyeAsdn. afi ln
T dorso(nZoItriaSbtP4.a,1 7h5o)l,o otfTy pew it5h-b 6r anrcahyPesed.c tfionnr oartle ach­
sten(uZrInaSo P7.,3 5a5n)6d 5s peciomfTe nisn vge nfitnro arli Vgeinntfi.rn pasll a frcoemd
stolfriocmzA kmauid aSryyrad,aC ryhaau,n d sligahnttletyrso il oirgp hotslttyedo ro irofisrna l
uppreera cohfHe wsa nga-nhCdoh angjianogr iAgniunrs.e mofrvoemad n fianlo ringoitn
rivbears iannfrsdo, mi nnbears oifQn asi dammo rteh baynt hdei stance between posterior bor­
anKdo kon(oZrI SnPo7:s2 5772,7 723,1 2d,e orsf e cnoantrdpoe o stbeorriodoefery r( e u su­
78559,7 1120,4 2126,3 7262,0 5Z6M;M U: alnloymt o rteh bayne ydei ameCtaeurfid)na.l
emargtiomn oadteer ateNloyt rfoaorcfkeesd .
no2s248,2541,3499,3644,3646,5520,5558)
adipcorseoesn ct asu pdeadlu Anicrl-eb.l adder
m�e���ili�iliefurmcr�o�a�
boncya pswuiltreho unwdeesldpl,a lcaetde ral
conspwehciiTlfis ect ,e nduirffiaent r hspe r es­
plaatnesdsh o(rts omeitnidmiespsto isntcetr)i or
encoefc haracyteelrliodswatirmiskach r kings
cornIenrtsew.si t4tio hnrm eo rceo i(el xsc Tept
ont hcea ubdaasmleo ,ra en teproisoiortft i hoen
hialsmpna.w.r i i2t c ho iSlesx)du.ia mlo rphism
dorfisnam,lo rpeo rietnsh s eu pratceommp­oral
pronoumnaclweeidstt; wh o orm orber ush-like
miss(u5vr ste.y pi3c)aa nlidlps yo ssaid bilsy­
padosft ubeirnpc rleeosr bital/suborbital region
tinct bsufoptrie tcrsei -eesv,a mluucmahot rieo n
one acshiod fhe e aidmn;o sstp ectiheepssa ed s
mateirnsie aeldA eldsT.ot ,e n(u6si pse cimens
alpsroe soenbn rto adpeencetrdoay rsa.l
fromK alai-PVaammaZirIr,S,n P o2 .0 71e8x,­
Inclusdpeedc( iveaslo iftd hitetyw s op ecies
aminieasdd i)s tsipnecrctai tethsh eaavr na riety
markweidt* wh a psr ovbeyZd h a1o9,8 T4):
ors ynoonfyT m s tolibcezckaiaudtisi ,eff ers
fromt hlea tittneh mreo reel onbgoadltyoe,n gearl exansdprn.a.T e;.a quaecasepnr..uT. ;l eae
breviDbian1rg9b,9a* 3 Tb; r evi(cHaeurdzae n­
ansdh allcoawuepdrea dlu nmcolraeen, t erior
ste1i8n8,T8 c )a;k aeCna&so Zi hsu 1,9 8T8;
posiotfdi oorfins n(ad lo rosrailg icnlaottsoee sr
snotuhtta onc aubdaasile nc, o nttrora esvte rcsheo p(rHaoi1r 9a3,T.4 c)o;n ip(tTeurrad akov,
condiotrei qouni,dp iosstioatfdni otor nis nT.a l 195T4 g)e;r zeCean&os Z ihsu1 ,9 8T.8 h;i al­
stoliaclziskntoa h sieh) a,op fbe r ushp-aldismk aers ipn ..T. ; i nter(mKeedsi1sa8l 7e*6rT),;
lepto(sHoemraz e1n8s8Tt8l e)hi;an (s,Ra eeg an,
inp reorbitaolfm/ asluaebnsoid rnm b oirtea l
1905T) m;i cr(oSptsae ciehnrnd1,8 66T);
porietnsh s eu pratceommpmoir(satslyu priec ally
5o r6 ,ra r3ev lsty.y pi3c,ra alrl4eyi l nTy obtusWiur& oW sut1,r9 a8T. 8 r;o ssoperegrina­
torsupnm.. T ;s tol{iSctzekiani1d 8a6c6h)n;er,
stoliczkai),
T tanggul(aZehn1us9i,8s 2T ) t;e nuicauda
Ts tolaincrdze kalisa pteec(dii ensc lauldli ng
(Stein1d8a6cT6h t)ne;en r(u,Di asy , T1876);
oft hodsees cbreilbaoenswde wfo)r,m itnhTge
texi(eZnh&saW ioas n 1g9,8 T8x )i;n gshanensis
stoliczdkiaffifre-orgmo r tohsueppre, c aineds
(Ya&nX gi 1e9,8 T3y )a;s in(eAnlsc1io8sc8 k9,) .
specgireosou fpt shg ee nTursi pliontp hhey sa
RemarSksp.e coifte hsci osm palremexo st
follocwoimngb inoatfci hoanr acters.
Bodeyl oncgyatlei ntdhri(icbcrkuae tll ,a ticvleolsye ltyto hrT.ee b loamtbeifdr onsT-complex.
bombif(rHoenrsz e1n8s8tt8eh)ioe,nn ,ld ye ­
comprieTnsa sleedx asnpndr..aw )ei, nt eha rly
scrisbpeedco ifte hlsea ttgerro duipff,e rs from
stradiogrphsrtao lfic laeu;pd eadlu mnocdleer ­
thset udcioemdpi lnte hxce o nvienxt erorbital,
atetlovy e erlyo nhgeaastdte r;o dnegplrye ssed.
smaelyleearsrc, hd eodr psraolpfi elcet,il niaptsed
Eyemso detrosa mtael l i(nTm miincurtoep s);
wiuthp pleiforpr mianfi gn gerp-rloijkeec tions
interroergbfliiaotatnan lmd o derbartoealdy.
latearbaotlvhlbeeya osfet hipraodifb r a rbels,
Nasflaalsp h obrtr,o aandrd o unPdoesdt.e rior
ZOOSYRSOTS.S VoIlCJ.AO A.M.P rokoNfieeswvp :e coifTre ips/ ophysa 195
mucmho reel onagnandta erdcr aouwpd eadlu nd­ep8t.(h28 .0t-i1m0ie.sns3t )a nldean(rgSdtL h) .
cle daenedpfo lrkye cda udfnai.Al l sTo Caudpaeld uonfcm loed edreaptttehh le,a tter
macromaa-ccuolm(apitlnecxls updeecTdi ecso:n ta1i8n.(e81d 9 .6t-i2m4ie.nSs0 La ) n 5d. 0
macromaYacnug1l,9a 9Tt0n a;c mdaLnaenns,.i (s5 .0t-i5mi.ent3s h) ce a updeadlu lnecnlget h.
Yan&gC he1n9,9T 5n ;a npanjiaZnhg&ue nsLiesn gotfch a updeadlu ncle much greater than
Cao1,9 8s8e)e tmobs ec lorseelltayott heTed maximduempo tfbh o dayng dr etathheaerna d
stoliczk<taJit-hdcoiouffmgfrephorr mt�s hx le a t­length, 37c.6o ( n3t.a9it-ni4em.ide5nS s) L .
teirnt hper esoefen lcoen gaten baasraMblae xli-mluimk oefw b ioddetyqh u taoml a ximum
flalpo,n pgeecrtfi onar naldli w pist hfuorurto wdse.p ths.t rHoednaegdpl ryec sosneibdcl,au ln,t ly
Tg rif.fitGhiiin therfr,o mt1 hI8en6 db8ua,ss irno,u nadnetde rmiuocwrhil dyte;hr ad ne eipt,s
alissvoe rcyl otsoTe s toliczkbauit-l ceonmcgpotlnhet x4a .i(7n4 e.d4t -i5mi.eSn0sL )S . n out
diffientr hspe r esoefsn lciedg ohrtts iadolgn e l encgotnht a2i.(n02e .d0 t-i2m.ie3nhs) e ad
thlea tshtio rfcd a updeadlu ncle. lenagntsdhl ighttlhyla egnnr getaht eorf postor­
Itnh dee scroifpntseipwoe nctsih foeers m,u lbai ptaaorlhft e aEdy.em so derdaotriespn,ao ls i­
NFi gsi vaeftnNe ars e(k1a9 9m6e)a,s uremteineotyndse;i ameter1 .(c14 o. n2t5at-ii1nm.ee3sd)
anrda tfoirho osl oatrygepi evfi ernas ntfod l ­iinn terdoirsbtiPatrnaoclpe o.art rigeoi nvisen n
lowbeydd a fotrap aratyippnea (rse)n theTsaeb1sl..e
Botahn tearnipdoo rs teriort on­ares close
Triploapqhuyaseac asepnr..u l eae getthhelera ,t tetrh aalnnat reogrneaierno e drn ­
(Fi16g-.)s cloispteo ss teNraisoflaralslp yh .or rto,u nded.
Mouitnhfe riipnoo rs i(tFii2og)ln.;i s ptsr ongly
HoloZtyIpSne.Po1 .2 4fe9m4a,9l 4me m,S Ll;o calfuirtyr: owLoewdle.iwr pi dwei,wt ehl l-developed,
Ba-[tBsQahuu R? i]v terri,b ouaft B(aryl= u eC hangjiang)
Riv[ee nrov,f. Yu sQhiun,ge ha3a.°3i N 9, °2E ]N,WC hinafl;am te nltoablUe pspm.ea rr ogfilno lwiseper p a­
coKlolz.&l:K o avz nAaukgo1uv9s,0t 0 . ratfreodmu ppmearr ogfilno jwaeLwro. w jeawr
ParatyZplenSsPo:1 . 2 439s 4pae,c i8m0e-mn1ms0 ,8 broardoluyni doneu dt lwiintehm,oe udtik eaenl .
SLs,a mleo caanlddia ttye . Nop rocedsesnutsio forkr emoeinlus p pjearw .
Diganos.iB sodtyh imcakx,i mwuimd tThh reeo fmp oadierrsla otnbegal ryb els. Length
subetqomu aaxli mduempC tahu.dp aeld uncoifefi( rrsotsp tacrioarnl t)1a .i(6n1 e.d4t -i1m.e6s)
lencgotnht a3i7.n6 e-dt4 .i5mi enss t andainrt dhle e nogftt hhse e c(omnadx ialnltdah rey ),
len(gStLhC) a.u dpaeld udnecplcteoh n tailnaet1dt. (e13 r- 1t.i2mi)ent s hl ee nogftt hhte h ird
5.0t-i5m.ie3ins t l se nSgntohsu.lt i glhotnlgye( rm andipbauMilaran.rd )ib baurlbeaexrlt tseo n d
thpaons topraborifhtt e aaAldn .t emrairoogrfi np ostmearriogofior nr btihtel,ie nrcg otnht·a ined
eysei tupaotsetdte orm iooucrto hr nIenrfr.a 2­.(235t. i3m)ie hnse leandg Stuhp.r asoernb­ital
orbciatnwaaillt haonugtufl leaxruB rreu.s h-sloicrkayen oafhl e a(dc scoo)n tinuous, confluent
padosnp reorbitarle/gsoiufomb nao lrebwsii titanhlfr aocrabn(iactliat olhl) ea; tc toenrfl uent
fineglryan ulDaotrefisdnoa. rl i gitnco ca luo­wsietsrhu pratceommpmoir(saeslsau tnrb)edo dy
dabla tsheta inop fs nowuitst,lh i gchotnlcyal vaet leir(nacelL ( LF)i3 g4,s) C .ib oe anroas n gu­
uppmearr gainendi gbhrta nrcahyeTsdio. pf laflre xuberlepo ows teroorvbeminattrarglai ln .
adprdeosrsfisenead xl t epnodsitntegvor e irtoirP­ reoperculcoa-n(mca aplnme d)ni odbnstu hlea r
caolaf n fianlo riTgiiopnpfs.e ctfionforsra mle dl ionfme o uNtuhm.b oefpr o rielnsa terosensory
bys ecaonntddh iorrod n tlhyib rrda nrcahyesds. y ste7m-,:c8 i1co1s -oe1 s34tc,, p 6mc, L 7L8 -
Porienls a ter7a8l- I8ln6it.ne ews itt5ih-n 6e 8 6L.atleircnaoelm plete.
coiClosl.o roandt oiroasnna ldla tseurrafalc eDso rfisnao lr igisnlaitgcehlstot lsocye a ru dal
ofb odvye drayr wki,t4 th o1 0p aslaed dolfe bsa tsheta toni opfs notuiotpfa; d predsosresda l
varisohuaospn e dorsum. fine xtenndeiatnrotgl h yve e rttihcraotluh geh
DescrDipUtii8,Ao. I n I5.P,. I 9,- 1V1I 6.,7- . midodfal nefia nlb asUep.pm earr gidno rosfa l
CI +8+S8pb+.r1I .2.Ve . r t. 41+= 3472--N34F83 +fi. n s ligchotnlcAyan vafieln.s hort-bwaistehd ,
42-(4137 -2128-()23+ 4)( 11)9 -(2108 -19)s.traliogwmheatrr gin. PmeocdteorraatLefilpys
Bodsyc alelesst,h iwccyikltm,iho n rdelroinecgxa,tl e,fon drni enatgrw loy- tpheicrtdos- ven­
orl eusnsi fodrempa tnhwd i dbtehfo droer satlrd ails tianln ecnegO tuhtp.ee rc tfionrr aayls
finb,u bte comsihnagl laonswdle irg chotrnl-byr oadteionpffie ndfo ,r mbeyds ecaonntddhd i r
•p restsoewcdaa rudpd eadlu mnacxliemb;uo md yoro ntlhyib rrda nrcahyVesed.n t firnsasil t uated
196 A.M.P rokoNefiwe svp:e cofiTersi p/ophyZsOaO SYSRTO.S SIVCoA1l 0.
[)CJ
6
Fig1s- Tr6i.p /opahqyusaae caesrpnu..1l :,le aateve irea2wl,v; e ntvriaeolwfm out3h,4; ,c epch alliaterosensory
syst(e3sm,u pratecmopmomriasplsa utrt4ee,l r ant;ve irea5wl,i) n;t es6ta,ii bnrle a;db doencrya psAublber.e viations
forl aterocsaennaasrlogesri yvi entn e xt.
froms ligahnttletyrso il oirgp hotslttyedo ro iro­rC omparTihsinosens w.p ecsitersi dkiifn­gly
safilno rigtihnefi.info,rsr mebdys ecoanndd ferisnt hceo lorfraotmai looltn h sepre coife s
thiorrod n sleyc bornadn rcahyVesed.n terxa­ltshg ee nIuatsl .s o frdoi.moff tehresr osfp ecies
tensdleidg bhetyloaynn dus so,m ettiotm heest hTes toliczka(ei x.ccTeot pmetpn lueixc auda,
anfianlo riCgaiunfid.nas ll ightlal yi tfotrTlkc eeh do,pa rnaTdci o nipitnte hcreoa n)c uapv­e
shotrhthaeenra d. pedro rfisnma alr gTihaned. d itdiiosntailn ctions
Anucsl otsaoen fi anlo ridgiisnt;ba entcwee aernae sfo lloTwhsne.e swp ecdiiefffrseo rmsT
thesmu.b etqoeu yadeli amIentteerls.ot nignt,ee nuiicnta humedo arr eo bcuasutpd eadlu ncle
wi5t-hc6 o i(lFsi5 g)A..ib rl adcdaeprsa usil ne ( deopfct ahu pdeadlu in6sc. li3eni tlse nignt h
Fi6gF..r peoer otafii obrnl adderS erxeudaTul tc eendu.i cgaruedamata)ex,ri mbuomdd ye pth
dimorpprheissmeman lwtei;sta ph a oifer l on(­8.0v-s17.0.i .7nT3 t enuiacnasdut draa)i ght
gabtreu shp-aldoisftk ueb eirnsc ulbeosr b(ivtcsao.ln /c ianTvt ee nuilcoawumedaar)r ogfi n
preorrbeigtiTaoulnb .e rcounlp aatdviseo rnyt haen afilnT .h nee swp ecdiieffsfre ormsT
finNeo.s upcahdo snt hber oadpeencetfidon r aclo nipitnte hgreea n stlloyp (innhogut m ped)
rays. dorpsraiollf oefs nosuitl,vp eerryi toneum
ColorBaotdviyeo drnay:rb kr,o wannitsehr (iborro­wtnobi lsahc iknTi c sohn ipstheorrat)e,r
lya,np da sstimona gr bwlieptdah bl aec kgroiunntde lsatringnueemr,bo efbr r anrcahiyentsdh e
postenrodi iosrtldiyans;rcp kto o trbs a rHse.a dd orfisn(a 8vl s 7.i nTc onipatnbeder tadt)ee ­r
dadrokr smaalrlbyol,nce hde aenkodsp ercuvlealrot puebde rpcauodlnast t hpeerd e orbital/sub­
reglioownpe;ar or hfte aadnb de lplaylF er.o4 m obrti a rle giionmn ae lsT.h em ucfhni er
toI l0a rpgasela ed doblrae rsvs a,r iiasnbh laep et,u bercountl habetr iuosenhs -ulbiokrpbaidtsa l
ond orsPuomo.r lyd-avrikmsiaisrbhklo iecn ­gosfm alaesws e alslt hfoer mo ft hpee ctfionr al
cuorna lfiln bsu,o tn dlaysr tkr oenta hkbe a se( apfoerxm beyds econvds-t.th hiirr(ddu- sfiuft­h
.o fu nsegmdeonrtfisenard la fuylsl -coloaulforlueyrd bt.rh a)n rcayhisenc d o mpasrpeedc ies)
Peritsoinlevbueemcry o,m diarnnkgee varer r tdei­stintghuneie swsh peescfr ioembs o tTh
brcaoll umn. stoliacnTzd ck haoip Tr cahio.p arlahsiao s
EtymolTohgysep: e ncaimefoesrm efrdo amq uas ligdhetelcpyae urpd eadlu (n4 c:l5e-i 5ni. t2s
(waitLneat ria nnc)da er(ubllieLunuate si rne)fe rlse nvgst5.h. 0-i5n'. tn3he esw p ecainetdsh )e
ttoh dei striitnbh Buelt Ruiieov nse yrs tem. upppeoror fte h per eoperculcoa-nmaaln dibular
.
ZOOSYRSOTSC.SA VoI l 1.0 A.M.P rokofiNeevsw:p ecoi/eTsr iplophysa 197
TablMee aIs.ur efomer1n rtsi ploapquhayescaa seprn.u(.l en4a )e =
Character Holotype Lim M±m cr
Ipne rceonfst taangdleae rndg th
HL 21.3 19.9-22.251 ±. 03. 53 1.07
BD 12.2 I0 .122-.5 11.6±0.72 1.25
CPD 5.3 4.2-5.3 4.±9 0 .24 0.48
CPL 26.6 22.2-26.62 4±. 08. 94 1.87
aD 51.1 51.0l- 53. 5.1±6 0 .50 0.99
aV 52.1 51.5-52.55 2±. 00. 29 1.50
aA 67.0 67.700-.0 68.±9 05. 67 1.33
PV 28.7 28.7-30.62 6±. 04. 62 1.07
VA 17.3 17.3-18.41 7±. 07. 34 0.59
IP 19.7 17.6-19.71 8±. 06. 44 0.87
IV 16.8 15.7-16.186 .±1 05. 25 0.51
IC 16.5 16.3-18.11 6±. 09. 41 0.82
IDb 8.5 8.5-13.91 1.±8 15. 28 2.56
hD 18.1 17.1-18.91 8±. 02. 42 0.83
!Ab 6.9 6.9-8.2 7.±3 08. 31 0.62
hA 14.9 11.4-16.135 .±3 05. 43 0.86
an-A 4.3 3.8-4.6 4.2±0.17 0.34
BW 11.7 I0 .2-11.71 1.0±0.75 1.06
Ipne rceontfha egalede ngth
ao 50.0 42.9-50.407 .±3 15. 69 3.39
20.0 18.6-20.109 .±2 08. 29 0.59
00
io 28.75 23.3-28.7255 ±. 12. 23 2.46
hbr 40.0 33.3-40.03 6±. 11. 43 2.86
an-A 20.0 16.7-23.31 9±. 18. 37 2.73
NotAeb.b revairasae tf ioolnlnso m w-nnsb:eo rfs amplLlems- ;ma ximvuma rioafct hiaorn.asMc - tmeearn;;
m s-tanedrarrodomef ra n;s s-tanddaervdi HLa-thieoan;l de nBgtDh- max;i mumb oydd ehpC;tP -Dc audal
pedlllldcelpeCt PhL;c- aupdeadlllll lcelneag Dt- phr;e ddoirsstaaalV-n pcree;v ednitrsataAla n-pcree;a nal
distPaV-n eccet;o -dviesnttVarA-ncavele; n trdoi-satnaIa-lPncp ee;c tfionlr eanlIg V-tv he;n fitnral le ngth;
h
I-Cl engctahu doa!flD- dfion;rsfi anbl a lseen hgthD-; d orfisnha eli !Agbht ;a- nfma bla lseeng htAh-a;an l
fm heiag-nh tA; d -'-istfraoncmae n tuosana ·lfi no riBgWi- nmax;i mubmo dwyi dato-h s;no ulte nogo-t h;
horizodnitaamelot fee yrie -o;int ·e rorwbiidhttba-hrlh; e iogfbh rta nacpheiratlur e.
198 A.MP.r okoNfeiswep ve:cof iTersi plophZyOsOaSYRSOTS.S VIoC1lA0.
lioents h lei onfie n fraocrabni(atvloas ntl. h e DiagnBoosditysh .ii ctmksa, x imwuimd th
lionfme o uittnhh nee swp ecTia eqsu)a.e csaueb­etqomu aaxli mduempC tahu.dp aeld uncle
rulsepna.me. a byed istinfrgoumTs b hreevdi l­encgotnht ained i4nS. L9C.-a 5u.d3a lt imes
barbbyta h leo nhgeeaardn b da rbfreolmTs ,p edudnecplcteoh n ta2i.n7e5td-i 3mi.en3is t s
tanggublyta hepenr seisosef5 n -c(6ev 4 s)i. n ­lenHgetahmd.o rbel utnhtia onnt hsepre ocfi es
tesctoiianlnetysd p ircoauln (dne"odst h ovtehlce­o mplex. Stnhoaounrat sl osnahgso rter
likaesn" o tbeyZd h u1,9 8p22.,2 4l)o wjearw ,p ostopraborifhtt e aaAldn .t emrairoogrfie ny e
fromT m icrboytp hlsea rgenra reryioenws­e,pr l apcoesdt teorm oiuotrh Icnofrranoerrb.i tal
terorsbmiatlmalalex,ri mbuomdd ye pthi n( c7a.n5wa ilt haonugtufl leaxruB rreu.s hp-aldisk e
SLi nTm icrfoorpmos f)t ,hp ee ctfion(r aaple oxn p reorbitarle/gsoiufmob anol firenbsei ltya l
formbeyfod u rbtrha nrcahiyeTn md i craonpds g)r anulDatoerdfis.noa rli cgliotnsco ea ru bdaasle
morfuer rolwieTpdhs ne.e swp ecidisie sst itnhcatno fst niopu ts,t rwauiiptgphmhe atrr agnidn
fromT i nterimnte hdipeao siotfti houenp pesre vberna nrcahyTesido.p fa p dr essefidn dorsal
enodft hper eoperculsoe-nmscaoanr­dyni obetux ltaernt dovi enrtgio cfaa nlfia nlo riTgiipns.
naolnt hlei onfem ou(tvhos n.t he olfi onfep ectfionrfosar lm ebdys ecoanntddh ird
infraocrabniiatnTla i ln teramnesddim aa)l lberran crhaeyPdso .ri elnsa tleir8na6el- I9n1t.e s­
numboefpr o rients hs eu pratceommpmoirsta­ilwn iett whco o iBlosdw.yi dtahr mkaerrb lings
su(r3ev s4.)F .r oTm t entuhinese ,sw p ecioerrse ticaunl1da0 t-bi1ro1on wscn riosshos nb ars
diffietnrh sme o rreo bcuasutpd eadlu mnocrleed ,o rsfruommn apteoc audal baanstee,r itohrr ee
postpeorsiiootrfti hdoeon r fisnoa rli agniidnn owfh icclho sspealLcyoe ndg.i taurdriannagleldy
thneu mboefpr o rietnsh s eu pratceommp­obrlaalc kiosndh o rdspourtmes s ent.
missTul raec.u sdniifffrgreoirmt s h nee swp e­DescriDIp It7IiA.,oI nI5.,P. ,I9 -1V0I7.,.
ciients h gee nehreaacldo nfigusrhaatopifeo C nI ,+ 8+S8p+h.Ir1 ..0 V .e r4t+.4 0=4+51N.F-
brushp-aldiisnsk ueb orrbeigtoiafmol an l es4,5( :2 207)( 4+)( 11)8( 17).
muclha rpgoesrt nearrileoosrn,fi g neasrn c da u­Bodsyc alceylleisnstd,hr imicockrao,elrl , e ss
daple dusnhcoltreht haeenra (dvl so.n tghearn u nifoirdnme patnwhdi dbtehfo droer fisnab,lu t
heiatdnh nee swp ecTihedesi )s.t ifrnocmTt iboencso msihnagla lnosdwl eirgc hotmlpyro ens sed
lhasaien ctlhudede ee cpaeurd al (p5eo.dr3u cnaculpdeea dlu mnacxliem;bu omdd ye p8t.(h97 .4)
letsismi ecnsa udal lpeenvdgsu5th.n. o c8mrl o er et imienSs L C.au daple dumnocdleer daeteepl,y
iTnl ha)st,aio eptf h aed prdeosrsfisenead xl t eniddt­e pctohn ta1i6n.(e21d 4t .i5m)ie nSs La nd
intgto h vee rttihcroathulemg ihd odfaln eba als e3 .(32 .t7i5m)ie ncs a updeadlu lnecnlget h.
(insotfbe eaimdno grp eo stienTrl ihoaras nade )L engotfch a updeadlu mnucclgher eattheanr
thpee ctfionsr haa(alp p efoe rxm beysd e cond-mtahxiirmddue mpo tbfho dayns dl igghrettatlheyar n ,
branrcahiyetnsdh n ee swp evcsit.eh si rdi-nfo ourre tqhut aotl,h h ee alde nigittcs ho ;n ta4i.n9e d
Tl hasTay ea)s.i ndeinfffrseoirmss ntehswep e­(5.t3i)mi enSs L M.a ximwuimd otfbh o dy
ciietnsh r(evtweso .b) r uspha-odlnsie kaseci hd es ubetqomua axli mduemp th. Hweiadde rm uch
ohfe iamnda lreosu,ghs eurro ftfah cepesa aedn sd thadne espt,r odnegplryec sosneibdcl,au ln,t ly
int hfoer mo ft hpee ct(oa rpaefoxlr sm ebydt her ounadnetde rwiiotsrhll oyp;i ncgo ndtoorusra l
fourbtrhan crhayei dnT y asinFeontrshd ieis s)­fr.o mt hien tertoosr nboitutitHap el.a l de ngth
tincotfTi oanqsu aecaseprn.u.fr l oemTa ec onta4i.(n94e .td8i )mi enSs L S.n osuhto rter
hialmsapnr..Ti ,r osseogrpienrastpon.ra. un md thaonra sl onagsp ostopraboriftht eaaildt ;s
Ta lexansdpnr.s.a e teeh d ei agannoddse essc rliepn­cgotnht a2i.(n62e .dt2 i5m)ie nts h hee ad
tioonfts h esspee cies. lenEgytemhos.d ersamtadelollryi,s pn ao ls ition,
witehyd ei ameter1 .c(7o1 n.tt5ia)mii enns e d
Triplopbbiyaslasm pan.r. i intersoprabciet.a l
(Fi7g-s1 2) Measurienm% e onfSt Lhs:e alde n2g0t.h2
(20m.a9x)i;mb uomdd ye p1t1h(. 122 d.e6p)t;h
HolotypZeI.Sn Po2 .6 2m4a5l,8e 9m, m S LB;a -tsohfuc aupdeadlu 6n.c(2l6 e.l 9e)n;og fct ahu dal
[BQau ?R]i vterri,b oauBf tl a(ury=e C hanRgjiivaenrg,)
[enovfY. u sQhiun,g eha3a.°3iN ,,9 °2EJN,W C hinap;e du2n0c(.l12e9 p.e0c)t;o -dviesnt3ta2rn.ac0le
colKlo.z&:lK oavz naAkuogv1u,9s 0t0 . (28v.e7n)t;r doi-sat2na0an(.lc28 e6 p .r4e)d;o rsal
ParatZyIpSnePo.2 . 6 24I5s ap,e c(ifemmeanl ed)i,s tanc(e5 6p5.r33e.)da4;in salt7 a3n(.c70e5 .9);
87m mS Ls,a mleo caanlddia ttye . prevednitsrt5aa2ln.( c35e 1 l.e2n)og;ffit nhs :
ZOOSYRSOTS.S VIoC1lA0. • A.M.P rokoNefiwes vp:e cofiTreisp /ophysa 199
--c
9�
cso
C;:::j
12
10
Fi7g-s1 T2r.i plhoipah/ysmspnaa.r. 7 i:l, a tveire8awv,l;e nvtiroeamfwlo u9t,1h 0;c, e phlaaltiecr ossyesntseomr y
(9s,u pratceommpmoirpsaasltu t1re0elrn, a ;tv eirea1wl1i), n; t e1s2ta,iib nrle a;bd doencrya psule.
pecrta1ol5 (.127 v.e2n)1t,2r (.a19l7 c.a2u)d,a tli miehnse alde nSgutphr.a soernbsicotaryan la l
15.(217 l.e2n)og;fdt ohr fisnab la s9e.( 09 .8o)f;h ea(cd s ou)n intercrounpftlweuidet,nh t
lenogfant ahfil nb a6s.e(7 6 .d9o)rfi;sn ha eli ghitn fraocrabni(atclait loh )le;a tjtoeirns iun­g
15(.177 a.n8fia)nlh; e igh(t1 41d.2i9.s)9t; a pnrcaet ecmpoomrmail(s essuatrnbe)do dlyat eral
anuso-rain3g.a (i9l4 n . m6a);x imbuomdw yi dtlhi (nceL (LF)i 9g,1s 0 C)i.w oi thaonugtu lar
10.(11 2M)e.a surienm% e onfht esal de ngtfhl:e xbuerlepo ows teroovrebnimttararalgl i n.
snoluetn 3g8t.h(9 4 4.e4yd)ei; a m1e6t.e7rU pppeorr oefp reoperculcoa-nmaaln dibular
(22i.n2t)e;r woirdbtiht( a32l37b ..r38a) n;c h(ci pamol)nt hleip naes stihnrgot uhmgeoh u ctohr ­
aperhteuir3ge6h .(t13 6)d.;1i stbaentcwee neenNr u.m boefpr o rienls a terossyesntseomr:y
anuasna dn fianlo ri1g9i(.n24 2 .2). cs7o,c i1o2e ,s3 t,c p6m,c L8L6 -L9a1t.e ral
Antearnpidoo srt nearrcielosotr so eg etthheel ri,cn oem plete.
latltaerrtg heaanrnt eroinoaern edn cliots eDso rfisnao lr isgliingc hltotlscoyea ru bdaasle
poste.Nr ai�oarll yflaanrpdo usnhMdooerudtt. ht htatonh teio pfs notuhtte;io pfa dpredsosre­d
inferiinpo ors i(tFii8og)nl.; i sptsr ofunrg­lsya filnd oensor te asclhi gthaotn lfiaynlo rigin.
rowLeodw.He pwr i dwei,pt rho nomuenncteadUl p pmearrg oidfno rfisnsa tlr aAingafihlnst h.o rt
lobUepsp.me arrg oifln o wleisrpe pafrroamt ebda sweidts,ht raliogwmheatrr gPienc.tfi onrsa l
uppmearr goifln o wjearwL .o wjearbw r oadcloyn t2a.i·(11n 7. et) di miepnse ctodviesnttarnacle .
rounidnoe udt lwiinmteeh,d ikaen.N eopl r.o cO­utpeerc tfionrr aalby sr oadteinopeffi d n;
essduesn ti(fo)krrem oeinlsu . p perjaw.fo rTmhebrdyse eec aonntddh ibrrda nrcahyVesed.n ­
paiorfms o derlaotnebgla yr. bT.eh lfiesr stt rfianlps l aacnetde trdoio orrfislnoay rl i thgeiinr;
(rospti,tr1ia.r(l7 l.} 2 5) sthiormttehesatr nh e t ifoprsm beyds ecootrnh dib rrda nrcahtyei;dp s
sec(omnadx iolnlteah,sre ey c)po aneidqr u taol o fv enfitnresan lsd l igihfrnto lnoytfa n uCsa.u ­
(ihno lotyopr1e .t)4i, m sehso (rtiepnra ratydpafieln)s ligfohrtklealyd i ,stt lheot rtheharen a d.
thtahnte h (imradn dipbauMilaran.rd )ib baurl­aArn ucsl otsaoen fi anlo ridgiisnt,ba entcwee en
beelxst teoon,erd n sdl igbhetylotynhp deo, s tet­hesmu betqoou rsal li ghtltyhe.aygnder i e­ater
rimoarr ogfio nr bainatdr ceo nta3i.(n62e .d6 a)m etIenrt.em sotdienrweati ett,whc o o i(lFsi g.
200 A.M.P rokoNfieeswvp :e cofiTersi plo•phZyOsOaSYRSOTS.S VIoC1lA0.
11A)i.br l adcdaeprsa usil nFe i1 g2.F. r peoer ­inT. t anggualnardeo nusniidonset dhs epre cies
tioofan i brl adder reddiumcoerdp.ho iftSs hemcex ou maplls emxa)ln,lu emrbo efgr i ll-rakers
expremsaslweeidsbt, rh u spha-odlfvsi e kfiryen e (alw1a0vy ss1. 0 -1s3h)o,rtv eern t(rwahlisc h
tubeirpncr leeosr bitarleg/Nisoous nbu.oc rheb xittteaonal dn fianlo riignci onm pasrpeedc ies)
paodsnt hber oadpe_ne.ecdt oral r.aa ynlsde. ds eseb opd Cyo.m pawriettdhhfo e rm dis­
Colo?nr:aBa tcikgnrdQc ulooiroapnta le.c, u·s.st ehmdeo, ss ti micloalrolsuypr eecidTis e s
browgnrieasybhv�.o a nwdh i�tb ieslSocwa'.t r­ossoperesgpnr..(i nsbaeetelo orbwuui)mtn;
tered,bmrarokwipnnrigess(s,ho � ndo oransda lc onttroia tT.s ,ht i alsmpan.r.l i a caknssyp ots
latseurrlifaolcfb e osda ynh de afdo rmami anrg­ ons idoefts hb eo daync da updeadlu Inancd l­e.
bloerrde tipcautltVeaertrnsye.h o(rtne oxtc eeddi­tiihtoa tnsh,d e o rbsaairlfrns o nottf hdeo rsal
inhga dlifs tbaentcwdeeo ercnso anlta onludar t fi­no riagswi enpl rlo noaustn hcoebsdee h iitn d
erlaildn aer)k -dborrobswaanprl rs e asleolnv te ra nadl dsios pulnauysbsul aalcd koiotsnsth h e
thaer freoam n apteoc aubdaaslNe u.m boefr dorssuarlfao cfte hb eo daync da updeadlu ncle.
ba1r0s- o1fw1 h,i scihax rs ei tuianfrt oendt T.ohfi alsmpan.r.a i l so frdoimffT. e rossros­
thdeo rosrailgT ihnra.en et edroirobsraa rls p eregrisnpna..it n to hraeub ms eonftc haeen ­
closspealcye dfr,o m3tt ho1o 1ws eesl pla cegdu lflaerx uorfte hi en fraorbital sensory canal,
(bayd isteaqnutcatoelh weiidrTwt ohp )a.r alalnetlem rairoogrfit nho er bsiitt uatetdo posterior
rowosfm inubtlaec dkoiitsnfrsh o notfd orsatlhv ee rtoiftc hamelo uctohr nmeurcfi,hn er
fina nodn mee diroawbn e hiiDtna.dr kmiasrhk t­ubercoutflh paert eioornb itaplai/dsnsu borbital
inpgrse osendn otrc saaulpd,ea clt,ao nvrdea nl­ malsemsa,l lero fnguimlbl(e-1vrr0s a1 .k2 e-rs
trfianlbs ,up tr onoaurnoecn eltdyh oosnte h e1 3), pceaduudnnaeclala resll oyan shg e a(dv s.
cauadnaodlnu nbrarnacoyhdfseo drfi snAa.nl a lm uclho nger tahnaidnns ohmmeeea ads)u,r e­
finp alPee.r itsoinlevuemr y. men(tssde ees croiftp hteitsowsenop s e cT.ies).
EtymolTohgyse.p ecniaemisesg ivaeftne rh ialsmpan.r.d i i ffefrrosmt hsey mpatTric
Swediicshht hyHoilaolRgmeiansrdtfo a rhh ils a quaecaseprn..ui lnte haaebe o mveen tioned
grecaotn tritbotu hsteyi sotne omfCat einctsr cahla raacnatdle isrntos h seh ovretnetrcr aaul­s,
Asifiasnh es. daple dunnecalaresll oyna gsh ea(dvl so.n ger
ComparTihsnioessnw p. e dciiffefresor mis t st hahne asdm)a,ln luemrbo efbr r andcohresda l
clorseelsatit tnih mveoe dse irnatteews it2ti hn rea (y7sv s8.)a ngdi ll-(r1va0sk 1.e2 rm)so, r e
coibllsu,sn ntoewurht i,ica shJs o anosgsr l ighntulmye lraotulesir pnaoelr( e8s6v -s79.81 - an8d6),
shotrtheparon s todribsit(tavamnsloc. reo e lr e ssi sno mmee asur(esmdeeeens tcsr oiftp hteisoens
oppocsointdeii ontt ihosenpr e coifte hgser oups,p ecies).
excfoerpT. tta ngguwliastenhno asulsito s an sg
postodribsittsaaenZlech eu1,,9 8p22.,2 3fi,g .Tr iploprhoysssao peresgprn.i. n atorum
1) i,tn h tahtte io pft haepd r esdsoerfidsnd a ole s( Fi1g3s- 18)
norte atchahen fianlo ri(gviesnx. t emnodroser
lebsesy otnhpdio si anntid)tn h ceo lorpaatti­Noenma chdeoirlsuosn oHtezaretuensis:1nt 8,8 380:
(partim).
teMrons.st p ecoifte hcseo mphlaevtxeh v ee n­
trfianlps l aocneo,drs ligphotslttye ort,ih oer Ho lotypZeI.Sn Po3 .7 1fe9m,a 8l7em ,mS LN;o rth­
vertoitfcha delo rfisnoa rli ogniilnT.ny;h ialmawersitGearnni snsu m,a mlolu nsttarienca omlNsl.;.M :.
sp.ann.sd,o mien divoifT.d sutaollsai nczdka Pirz ew1a8l8Js0uk.di g,fr ionmtg hP er zewraolustkei 's
in1 88t0ht,ey lpoec laliiietnts hmy eo untnaoirontfsh
T. aquaecaspen.r. tu hlveee aneta rrpaell asc ed
XinainnLdga nzhou.
sligahnttletyrot i hoer voeftr htdeio crfiasnla l
ParatyZpIeSrisPo.3 . 7 19sap,e c4i6 m4e-n8ms8m,. 5
oriTghinene .ws peciisuen si qaumeo nagl lSLs,a mleo caanlddia ttye .
known spT.e sctioelsi cozfki anti h-ecoDmipalgenxBo osditysh .ii ctmksa, x imwuimd th
posseosf2sv is4oo. nmr o r(ue s u5ao lrml oyr es)ubetqomu aaxli mduempC tahu.dp aeld uncle
coiolfis n teOsntlT.iyt n aen.g guhlaalsee nsssliesn cgotnht a3i.n9e5td-i 4m.ie6nSs L. C audal
th5ac no iolfisn tebsutttihn neee sw,p ecdiief­sp edudnecplcteoh n ta3i.n5e-td4i .mi9eni s t s
ferfrso mT. tanggujluadegnfrisonismZg, h u'lsenSgntohlu.ot nt gheparon s topraobrhfite t aadl.
(19d8e2s)c roiftp hlteai toitntne hsreh, o irtne­rA ntemrairogorife n ys ei tuoantt hevede rtical
tes(t2iv ns4e.c oiilnT.s t anggulatehnestihsr)om,uo guhct ohr Innefrra.o crabnioatfcla e l­
shaopfte h leo wjear(w w hii"scs hh ovelp-hlaislkeienc"ss oyrsywt ietmah n guflleaxrub ree-
ZOOSYRSOTS CSAIV ol. IA .0M.P rokofiNeevws: p ecofiTersi plophysa 201
O(J
18
Fi1g3s- T1r8i.p lroopshsyospae resgprn.i.1 n:3la ,at otvreiurem1aw 4lv;, e nvtireoawflm o u1t5h1,;6c ,e phalic
.l aterossyes(nt1ses5omu,ry p ratemporpaalt t1ce6lorn,am; tm veiirseas1wlu7)i r,;ne t e1s8ta,iib nrle a;db doenry
capsule.
lopwo sterboovreodnfeot rrr baBilrt u.s h-ilnSi LkM.ea ximwuimd otfbh o deyq utaool r
padosnp reorbitarle/gsoiufomb nao lrebssil tiagllhet tshlsmay an x imduempH tehas.dt rongly
rougghrlayn uDloartfisenaod lr. i cglionts oe rd eprebslsuencdto,ln yii ctwasil d,mt uhc ehx ­
caubdaatslhe ta inop fs nowuitst,th r auipgphectre etdhdsee pHtehal.de ncgotnht 5a. i(04n .e9d-
maragni6dn7- b ranrcahyTesido.p fa dpres5s.et4di) mi eSnsL S .n oluetnc gotnht 2a.i(3n2 e.d4 -
dorfisnea xlt ending boefay nofiannld 2 v.e5rt)ii htncei alamlede n sag ntsdhl ighttlhya ng reater
oriTgiipnos.fp ectfionrsa lfo brymt ehdi rlde nogfpt ohs topraborifhtt e aaEldy .em so der­
branrca.hPy eodri ens llaitne7er4 aI6.ln 8 t-east­teso m adlolri,spn ao ls ietyideoi na;mc eotne­r
tiwniet5 h-c6 o iBlosdc.yo lorsatiimoitnlo a tra i1n7.e 1 d(. 3-t1i.mi8eni)s n tersoprabciet.a l
thoatfT h ialsmpna..rb ,iuc tr ososnbd aorrs­P ropoarrtgeii ovineTansnb l2e.
subme codmies toinnplocyst t tedoro irofisrna l Antearnipdoo rs tnearrecislo orts oeg etthhee r,
� na nldo ngitaurdrianbnalglaelcddyko itsslha tltaertrgh neaa rn teorniaeon erdn cloisti ng
ond orsaubms ent. posteNraisfloaarlspl h yoa.rn rtdo unded. Mouth
DescriDpI tI6iI-o,A7n I ..I5 ,P. ,I9 .V I6,. infe(rFi1io4grl). i; sp tsr onglLyo fuwlreirrpo wed.
C1 +8+S8p+b.1r1 ..2 -Ve1r3t4..+ 37-38+w1i=it4nh2c -onsmpeincltuoaoblUue pssp m.ea rr ogfi n
43N.F- 42-(4138:-2 139-()23 5-+ (4 1)1) 8 -19lowleisrpe parfratoemd uppeorf lmojarwagewir.n
(17-18). Lowerb rjoaarwdo luyni dnoe udt lwiinteh,o ut
Bodsyc alelesst,h imccoykrol,erli e nsdsmr eidcikaealNen,ol p .r ocdeesnstuisofo rkr emoeinls
unifoirndm e pitnfrh o notfd orfisnab,lu b te ­uppjearTw h.r peaeio rbfsa rbsehloisrfen,tm ales
comisnhga llaonmwdoe rrce o mpraelsosneagdn mdo derlaotinemngl a ylR eo.s tratwlo ­barbels
caupdeadlu mnaxcilmebu;om d dye pctohn taitnheiasdrl doassnt ghm ea xiolnletharysel , a stltiegrh tly
8.(38 .0-t1i0mi.en6Ss )L C .a udpaeld uncslheot rhtmanean rd ibounleMarsan .d ibbuarlbaerl s
modeirdna etpeat nhtd h ickintdeses pcsto,hn ­extensdliinaggnh tteltrmyoii odordof l reib feni­ t
tai1n7e(.d14 6 .0t-i1mi9eSn.sL 7a )n4 d. (43 .5m-albeustt,op ostmearrgioiofnor r biinmt a le.
4.t9i)m iencs a updeadlu lnecnlget lha.t tTSehurep rasoernbsicotaryano lafh l e a(cd s uon)i nter­
is mgurcehta htmaeanrx imduempo tfbh o doyr rupctoendfl,wu ieitnnhtfr aocranba(cil it) toa;h l e
helaedn agntidhcs ;o nt3a. i9(n45 e. d0 -t4i.m6el)sa tter wciotsnhufl pureantcteo mmpmoirsaslu re
202 A.M Prokosfipeevco:i/ eTNesrw i plopZhOyOsSaYRSOTS.CS AVI o 1l0.
TablMee a2s.u rfoerTrme inptlso prhoysssoap eresgprni. (n n=a5 t)o rum
Character Holotype Lim M±m (J
Ipne rceonfst taagnlede arndg th
HL 20.1 18.0-20.31 9±. 03. 44 0.98
BD 12.1 9.4-12.51 1.2±0.70 1.56
CPD 5.75 5.12-56 . 5.±6 0 .20 0.44
CPL 25.3 21.9-25.233 .±105. 75 1.50
aD 52.9 51.4-54.15 2±. 08. 46 1.03
aV 54.0 53.7-55.95 4±. 08. 46 t.03
aA 70.1 70.1-72.97 1.1±0.90 1.56
PV 34.5 31.3-34.353 .±1 05. 67 1.34
VA 15.5 15.1-16.105 .±508. 15 0.34
IP 16.1 15.5-21.21 7.7±1.28 2.56
IV 155. 14.1-15.51 4±. 07. 30 0.60
IC 17.2 17.2-21.21 9±. 01. 79 1.76
IDb 10.35 9.9-10.91 0±. 04. 17 0.38
hD 17.8 16.8-18.81 7±. 06. 46 0.91
!Ab 7.5 5.74.-5 6.±6 0 .30 0.67
hA 14.4 14.4-15.61 4±. 09. 20 0.45
an-A 4.6 4.5-5.3 4.±8 0 .15 0.33
BW 10.9 9.4-12.41 0±. 08. 50 1.11
Inp erceontfha egalede ngth
ao 42.9 41.4-42.492 .±2 05. 31 0.63
17.1 15.4-20.91 7.9±0.89 2.00
00
io 28.6 27.3-33.32 9±. 12. 40 2.79
hbr 40.0 36.4-40.03 8±. 04. 81 1.61
an-A 22.9 22.9-27.62 5±. 10. 02 2.27
NotAeb.b revaisia Tnta ibIol.me