Table Of ContentSSN 0286-9810
Vol.52 No 2-1 Dec. l5, 1997
見最學評論
THEENTOMOLOGICALREVIEW OFJAPAN
日 本 甲 蟲 學 會
THE JAPAN COLEOPTEROLOGICAL SOCIETY
OSAKA
TheJapanColeopterologicalSociety
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President: MASA0 HAYASHI
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HIROSHIKON0, YOSHIHIK0KUROSAWA, TAKEHIK0NAKANE,SHUN-ICHIUEN0
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会 長 林 匡夫
運営委員 ・林 靖彦, 水野弘造, 野村英世, 伊藤建夫
木村史明, 初宿成彦
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酒井雅博, 林 靖彦, 伊藤 昇
評議員 河野 洋, 黒澤良彦, 中根猛彦, 上野俊一, 大川親夫
久松定成, 木元新作
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En†.Rev.Japan,52(2):57-60,Dec.15,1997
ThreeNewCetoniidBeetles(Coleoptera,Scarabaeidae,Cetoniinae)
from Southeast Asia
By KAORUSAKAI
9-]8,Higashiyaguchi2-chemo,0ta-ku,Tokyo,146Japan
Abstract Twonewspeciesandanewsubspeciesofcetoniidbeetlesaredescribedunderthe
namesofRhombo,-hlna(It/1onlbo,・/1lna)castaneasp nov fromnorthernVietnam,Theodosia
1) ( uheodosia 川, yasf1lfa s nov from Borneoand efero1'r/una ( otel-01'1-1/加cl)フ/01''lee11.fls
sumatranasubsp nov from W.Sumatra.
Inthispaper,theauthor isgoingtodescribetwonewspeciesandanewsubspeciesofthe
cetoniidbeetles from SoutheastAsia.
Beforegoingfurther,IwishtoexpressmysinceregratitudetoMr.K.IwASE,Tokyoforhis
kindandcontinuoussupportinmanyways.MythanksarealsoduetoMessrs.M.FUJIOKA,N.
KATsURA,K.KUME,T.MIYAsHITAfortheirkindofferofthematerialstothisstudy.
Rhomborhlna (Rhomborhina) castanea spnov. (Figs.1-3)
Somewhatnarrow;shining, darkreddishbrown, femoraandtibiaebright reddishbrown,
antennae,tarsiblakishbrown;hairsof inner facesofmiddleandhindtibiaeandmarginsofpy-
Fjgs.1_3.Rholnborhina(Rile川bet-/11na)(・1.・ltalea、pno、,..1.habitusinmale(holotype);').habitusinfemale
3.malegenitaliaindorsalview(、cafe:3mm).
58 KAORU SAKAI
gidiumblackishbrown.
Headrelativelysmall;clypeusdenselypunctate,quadrate,about1.1timesaswideaslong,
weaklyarcuateatsides,straightandreflexedatanteriormargin,roundedatanteriorangles.
Pronotum transversely trapezoidal, about 155 timesaswideas long, gently convex,
sparselyandminutelypunctate, lateralbordersslightlyangulatebehindthemiddle,eachgroove
alongthebordersendingbeforeposteriorangle.
Scutellumtriangular,nearly1.3timesaslongaswideandalmostimpunctate.
Elytraabout145 timesaslongaswide,widestbehindshoulders,marginatealonglateral
borders,finelyselTatealongposteriorborders,suturalintervalssharplycarinateinposteriorhalf
ofinnermargins,slightlybutshalplyprominentatapices;surfacealmostpolished,transversely
rugosenearapices,withsomerowsofminutepuncturesnearbases.
Pygidiumtransverselytriangular,about2.3timesaswideaslong,slightlyconvex:surface
finelyrugose,withlongsetaealongborders.
Fronttibiaslender,withtwoouterteethincludingapicalone,secondtoothobtuseinmale,
andwidersecondtoothsharpinfemale.0uter faceofmiddletibiasimpleinmale, frequently
unispinousatthemiddleinfemale.Inner faceofhindtibiaclothedwithlongsetaeinbasal2/3.
Stomalprocesslonger thanwide,parallel-sidedinbasalhalf,semielliptical inapicalhalf;
sutureofmeso-andmetasternumalmostdisappearedattheprocess.Third to fifthabdominal
sternitesslightlydepressedinmale,notdepressedinfemale.
Malegenitaliaresemblesthoseofotherspeciesinthegenus(Fig 3).
Bodylength:31.2~34.4mm;width:14.6~16.8mm.
Typeseries.Holotype: Mt.Piaoac,CaoBang,northernVietnam,25.VI.1996,M.ITOleg.,pre-
,
servedbytheNationalScienceMuseum(Nat.Hist),Tokyo.Paratypes:samelocalityastheholotype,1早,
3.VIL1995,K.KuMEleg;2早早,3.VIII.1995;1 ,5.V1.1996,M. IT01eg;1 25.VI.1996,M. ITO
,
leg;2早 ,27.VI.1996,M.ITOleg;1 ,1早.25.vm.1996.
Distribution. Northern Vietnam.
ThisnewspeciesresemblesRhombo,-hina(」R11omborhina)JeanneliRUTER from Yunnan,
ChinaandThailandinthebodyshapeandthestructureofsurface,but iseasilydistinguished
fromthelatterbythedifferentcoloration(almostblackinthelatter),thenarrowerbodyandthe
hindtibiamoresparselysetose.
Hete1-orr11ma (Heter・orrhina) borneensis
sumatrana subsp nov. (Figs 4-5)
Differentfromthenominotypicalsubspecies
fromBorneointhefollowingpoints:bodynar-
rower, glossyblackish brown, withsidesof
pronotum,scutellum, themedialmarkingof ely-
tra,mesostemalprocess,metasternumand ventral
Figs.4-5. ore,-o1-,-1/lna( cto・,o,-,/l加a) o,,1eensls
stl'natl-a'tasubsp nov;4.habitusinmale(holotype).
5.habitusin female.
ThreeNewCetoniidBeetlesfromSEAsia 59
faceof femorapalereddishpurple, whereasinthenominotypica1one,bodyglossyblack,with
brilliant greenmarkingsonpronotumandelytra;scutellumlarger; elytramorestrongly and
moresharplypointedatthesuturalapices.
Bodylength:19.0~21.lmm;width:9.6~10.2rmn.
Typeseries.Holotype: (i1',Padang,W. Sumatra, Indonesia, V.1991,preservedin theKanagawa
Prefectua1Museum,0dawara.Paratypes:]早,samedateastheho1otype11 ,1早,samelocalityasthe
holotype, VIL 1991; 1 HarauValley,W.Sumatra, Indonesia,111.1995;1 W. Sumatra、Indonesia,
, ,
IV.1981.
Distribution. W. Sumatra
8pz . eodosia ( eodosza) mzyas 加ﾍ s nov (Fig s6, &10
Male.Elongate-oval;headandpronotumyellowishcupreousgreen,apicesandvent1-a1
facesof cephalicand thoracichornsdeepred, elytrayellowbrownandwithgl'eenish luster,
ventralsurfaceandpygidiumbrightcupreousbrowntocupreousgreen,withyellowishtintand
closelyclothedwithpalegrayhairs.
Clypeusstronglytransverse,about3timesaswideaslong,denselyandfinelypunctate,
angulatebutnotsharplytoothedatanteriorangles;cephalichornstout,arcuatelycurved in lat-
eralview,sharpatapex,sometimeswithatuberclebeforethemiddleofdorsalface.
Pronotumsemi-global,aboutaslongaswideexceptfor thehorn,narl-owlythickenedin
posteriorhalfalonglateralborders;surfaceimpunctateondisc,shagreenedatsidesandthe
horn;thoracichemslender,somewhatshorter thancephalicone,horizontallyproducedanteri-
ad,bifurcateatapex,tallerthanwideincrosssection,withalargetubercleatthemiddleofven-
tral face.
Elytratransverse,about12-1.3timesaslongaswide,widestbehindshouldersandgrad-
uallynarrowedposteriad,marginateatsides,truncateatapicalpart,butslightlypointedatthe
10
Fjgs 6,8&10 Theodosia(Theodosia)11iyashitaispnov 16.habitus(holotype).8.headandthoraxinlater-
alvjew;10.malegenitaliaindorsalview(scale:2mrn).Figs7,9.T.(T)chewiOcHI :7.habitus;9.head
andthoraxinlateralview.
60 KAORU SAKAI
middle;surfacealmostwhollycoriaceous,sparselypunctatenearonlybase,
Fronttibiaveryslender,glabrous,withanoutertoothbehindapicalone;middleandhind
tibiaedenselyclothedwithlonghairsoninnerfaces.
MalegenitaliaasshowninFig.10.
Female. Unknown‘
Body length: 30.2~39.5mm(incl hem), 24.0~30.9mm(excl horn); width: 14.2~15.8
m m.
Typeseries.Holotype: Tawau.Sabah.Borneo,E.Malaysia,21. IV.1996.preservedintheNa-
,
tionalScienceMuseum(Nat.Hist),Tokyo.Paratypes:5 ,samedateastheholotype; l ,samelocali-
tyastheholotype,28.V.1977.
Distribution.Borneo.
ThisnewspeciesiscloselyalliedtoTheodosia(171eodosla)oho、t,iOcHI(Figs 7,9)which
wasrecentlydescribedfromBorneo,butiseasilydistinguishedfromthelatterbytheclypeus
nottoothedatanterioranglesandthoracichemnotmuchshorterthanthecephaliconewhereas
inthelatter,theclypeusissharplypointedatanterioranglesandthethoracichornisdistinctly
shorterthanthecephalichem.
References
MIKSIC、R.,1967.RevisionderGattungRhomborr1-lirlaHOPE. Elite,lt. Ab/1.Mus.Tie,k.D・tesde,t 35:267-
335.
l - l977.Mo'1ograp/lie er Cefominae ferpafaa,-・t1sc/1e, M o,・lenra/lscfle17 egie,1, II. l-400.
Sarajevo.
OCHI,T.,1993.AnewscarabaeidspeciesofthegenusI71eodostafromBorneo.Akitu、newseries,136:1_
4.
PAULIAN.R.,1960.CoIeopteresScarabeidesdoL'Indochine(RutelinesetCetonines).A,1,1. Soc. e,zt.
Fl-ance,129:137-223.
RUTER,G..1965.ContributionaletudedosCetoniinaeasiatiques.Bull.Soc.ent.F,・ance,70:194_206.
SAKAI,K.、l994.AnewsubgenusandanewspeciesofthegenusTheodosta.Gekkan-Mus/1i,281:10_12.
(ReceivedJuly15.1997:AcceptedJuly29,1997)
En†.Rev.Japan,52(2);61-71, Dec.15, l997
Revisionof theGenusSepedophilusGIsTEL(Coleoptera:Staphylinidae:
Tachyporinae) fromJapan:SpeciesGroupsofSglabratusandS ex:iguus
SHUN-lCHIR0 NAOMI
NaturalHistoryMuseumandInstitute,Chiba
955-2Aoba-cho,Chuo-ku,Chiba, 260Japan
and
MUNETOSHI MARUYAMA
DepartmentofBiology,FacultyofScience,TohoUniversity
2-2-1Miyama,Funabashi-shi,Chiba, 274Japan
Abstract TheJapanesespeciesofglab1-a11is-ande?1g1ltis-groupsofthegenusSepedophiius
GlsTELarerevised. Thethreenewspeciesandanewsubspeciesaredescribedasfollows:
, Sepedo/フ/litisgfa/,,-aflls,Sgfa ,arils oｽl1lall,a11ls e.、-1giMl s,andSe/egan「1ss加111s. Keysto
themembersofeachspeciesgroupareprovided. Theaedeagiandtheapicalabdominalseg-
mentsofmaleandfemaleareillustratedindetailforcomparison.
Duringthecourseof thestudyontheJapaneseTachyporinaewefoundaseriesofsma11-
sjzedsepedophitus-species,whoseelyu-ahavepairedmarkings.Thesespeciesremarkably
resembleS armatus(SHARP, l888),buttheyarereadilydistinguishablefromthelatterbythe
absenceofbristlesalongthemarginalareasofelytra,etc.UptothepresentsuchSepedophilus-
specjeshavenotbeenknownfromJapanatall.Bythecloseexaminationwecametoaconclu-
sionthattheyareclassifiedinto3speciesandasubspecies(Sglabratus,S.9labratusokinawa-
exjguusandSelegartli、.,sl,nus).AlthoughtheyhavesuchcommoncharactersastheSmall
njs,s
body,theelytra1markingsinpair,thehindtibiaewithoutsetae,thechaetotaxyon3rdto6thto「一
gjtes0_1_l_l,themale9thtergitecontinuous,andtheinternalstructureofmedianlobewitha
H_shapedor1,ectangularsclerite,SglabratusisdefinitelyseparablefromtheothertwoSpecies
bythestructuresofthemidandhindtarsomeresandthefemale8thter9itementionedinthe
djagnosjs. Thus,s glabratusandtheothertwospeciesshouldbeclassifiedintothedifferent
species-groups.
InthjspaperthespeciesgroupofS glab,-tltusisestablishedforS glab'atusandS.9.oki-
ss r nawanl's,andalsothespeciesgroupofSeーklgMl i fo e_fgut'sandSe/egan「IsS'm,tS.T ee
newspecjesandanewsubspeciesaredescribedandillustratedindetail;thekeystotheJapan-
esespecjesandsubspeciesarealsoprovided.Forthepartsofbodymeasuredandtheabb「eVia-
tjonsusedjnthetext andalso、for thetaxonomiccharactersincludingchaetOtaXy,SeeNAOMI
1)TaxonomicstudyonthesubfamilyTachyporinaefromJapan,3
62 SHUN-1C1-llR0 NAOMland Mt「NETOSHI MARUYAMA
andMARUYAMA(1997). Inthefiguresthesensorysetaeareshownbythearrows1thelongand
shortbristlesareillustrated、butthepubescenceisomitted.
SpeciesGroupofSglabratus
Diagnosis: Bodysmall;antennaebrownwithbasalsegmentspaler; elytramaculated,
withoutbristles;elytra1epipleurasubacute;midandhindlegseachwith3rdtarsomerebearinga
ventralflapwhichextendstotheapexof4thtarsomere,chaetotaxyin3rdto6thtergites0-1-1-
1,male8thtergitewith4longbristles;male8thsternitewith3longbristles;male9thtergite
continuous,with5longand1shortbristles;female8thtergitewithinnerlobelongerthanouter
lobe.
Onespeciesandonesubspecies,Sglab1・atusandSglabratztsokinawanis,belongtothis
species-groupinJapan.
KeytoSpeciesandSubspeciesofSglabratus-group
1(2)Aedeaguswithmedianlobelarger morestronglybulbousatbase,withnaITowerapex; internalstruc_
turewithadistinctrectangularsclerite,thesetousareabroader;paramereslessstronglyattenuateal
apices S glabratusglabratus sp andsubsp nov
2(1)Aedeaguswithmedianlobesmaller,lessstronglybulbousatbase,withbroaderapex;inlema]struc_
turewithrectangularscleriteincompleteandpartiallyamalgamatedwiththenarrowersetousarea;
Pa「an、eresmorestronglyattenuateatapices Sglab・,atusokirlawanjssubsp nov.
SepedophilusglabratusNAOMI et MARUYAMAsp nov
(Figs.1;2.A-E,G;5.A,D)
Maleandfemale.Forepart:1.52-1.62mminlength(Fig.1),weaklyshining,withrelative
P「oPOrtiOns: HL:7;HW: 13;PL:15;PW:24;SL:21;EW:24. Headbrown, with reddish
b「ownClypeofronta1area;pl'onotumbrown,withbroadmarginalareayellowishbrown;elytra
b「owntoreddishbrownoryellowishbrown,withsuturalareasometimesjnfuscale andalso
Withapairofmedic-lateralmarkings, themarkingsdistinct,oftenindistinctoralmostvanjsh_
in9;abdomenbrowntoreddishbrown:antennaebrown,withbasalsegmentspaleyellowish
brown;mouthpartsandlegsyellowishbrowntoreddishbrown.
Headt「ansverse,withdistinct frontoclypea1andmidcrania1sutures;eyesalittleshorter
thanthe2ndand3rdantennalsegmentscombined,weaklyconvex;pubescencesparse,occur_
「in9almostConcentrically,distinctlylongerandmuchsparserthanthatonpronotum;surface
Withfain、t fin9erprint-likemicrosculpture:antennaerelativelyslender,3rdto11thsegments
beCOmin99「aduaIlybroaderapically,10thalittlebroaderthanlong,lithslightlyasymmetrjca1,
obtuselypointedatapex,withALP:8:5:5:4:4:4:4:4:45:5:7
P「onOtummoderatelytostronglyconvexdorsally,broad,basalmarginveryweaklyarcu_
atelyema「9inateatSides,posterolateralcornersalmostrounded,notprotrudjngposterjorly;
PubescenceShort,verydense,almost fumingposteriorly;microsculptureextremelyfjne,run_
StudiesonJapaneseTachyporinae,3 63
ning transversely.Mesosternumwithamediankeel
low, runningalmost in full length,mesosterna1
processacutelypointed;mesocoxa1cavitiesnarrowly
separated;metasternumatmedianpartabout2/5 the
lengthoflateralpart
Forelegswithtibiaebearing3thinsetae,inter-
nallywithalineof thinsetaealongapicalmargin,
ventrallywithanoblique(ortransverse)lineofsetae
nearapex,externallywithayellowlongctenidium
whichoccupies the full lengthoftibiaandbecomes
graduallylargerapically;1stto3rdtarsomereseach
moderatelydilated,4thmuchsmaller thanandabout
halfasbroadas3rd.Midlegswithfemorabearing1
longand2or3shortapicoventra1setae;tibiaewith2
setae, f longapicalspur,andfimbriatesetaeatapical
margin;3rdtarsomerewithaventral flapextending
totheapexof4thtarsomere.Hindlegswithfemora
bearing1longapicoventra1seta;tibiaewithoutsetae,
f longapicalspur,andfimbriatesetaeatapicalmar-
gin;3rdtarsomerewithaventral flapsimilar tothat
Fig.1. Sepedophihtsglab,'atussp nov ofmidleginstructure,withTLP:10:5:4:2:6.
Scale:0.4mm. Elytrawell convexabove,weaklynarrowed
posteriorly,posterolateralcornersrounded,conjoint
hindmarginsshallowlyemarginate;epipleuronsubacute,withoutermarginalittlehi9herthan
innermarginonlyinposteriorpart;pubescencesimilartothatonpronotum;micrOSCulPtu「e
veryfineandminute,reticulate.
Abdomenstronglynarrowedposteriorly;pubescenceverydense,distinctlylonger than
thatonelytra;chaetotaxyon3rdto6thtergites:0-1-1-l;7thtergitewithoutb「Si tleS.
Male. Ejghthtergite(Fig2A)entire,with4longbristles;8thstemite(Fig 2B)Witha
largev_shapedemargjnatjon,3longbristles;9thtergite(Fig5A)narrow,completelyContinu-
ous,apical lobesmoderatelyseparated,with5longand1shortbristles;9thstemite(Fi9.2D)
symmetrical,narrowedapically,with2shortbristles;10thtergite(Fi9.5A)na「「oW,With1
shortthjnbrjstle.Aedeagus(Fig2E)withmedianlobemoderatelybulbousatbase,g「adua11y
narTowjngtopointedapex;internalstructurewithabasaltubeshort,almostSt「al9htandb「oad-
enedposterjorly,withascleritebehindbasalorifice,arectangularscleritejustbehindthebasal
tube,andtheareadenselyclothedwithmoderatelylongsetaeinadditiontopairedIon9itudina1
bands;parameresslender,extendingposteriorlyfarbeyondtheapexofmedianlobe,eachVe「y
thinandextremelyattenuateatapicalpart.
Female.Ejghthtergite(Fig2C)withinnerlobeshortbutalittlelongerthancute「lobe,
medjanemargjnatjondeep,notextendinganteriorlybeyondthebaseofnarTOWlate「alema「9ina-
tjon,wjth8longbristlesinadditionto2sensorysetae;8thsternite(Fig2G)ent「ie,With31on9
andl shortbrjstlesjnadditionto6sensorysetae;9thtergite(Fig 5D)widelySepa「ated;9th
stemjte(Fjg 5D)wjthhemistemitessmall,coxitesandstylielongate;10thto「gite(Fi9・5D)
almosttriangular,ciliateapically.
SHUN_lcHRI 0NAOMIandMuNETOSHIMARUYAMA
64
