Dendrotiidae (Crustacea: Isopoda) of the southeastern Australian continental slope

38 Pages·1998·10.9 MB·English

MemoirsoftheMuseum ofVictoria57(1); 1-38(1998)
DENDROTIIDAE (CRUSTACEA: ISOPODA) OF THE SOUTHEASTERN
AUSTRALIAN CONTINENTAL SLOPE
B. F. Cohen
DepartmentofCrustacea,MuseumofVictoria, 71 VictoriaCrescent,Abbotsford, Victoria 3067,Australia
Presentaddress: Marine andFreshwaterResources Institute, PO Box 114,Queenscliff, Victoria 3225,Australia
([email protected])
Abstract
Cohen, B.F., 1998. Dendrotiidae (Crustacea: Isopoda) ofthe southeastern Australian conti-
nental slope.MemoirsoftheMuseumofVictoria 57: 1-38.
Fournew speciesofAcanthomunnaandfournewspeciesofDendrotion are described from
material collected from the southeastern Australian continental slope. A. proteus Beddard,
1886 is refigured. The new species ofAcanthomunna represent the first records from Aus-
tralianwatersandthenewspeciesofDendrotion representthefirstrecordsfromwatersofthe
SouthernHemisphere. Akeytothe generaofDendrotiidaeispresentedalongwithkeystoall
describedspeciesofAcanthomunna,Dendromunna andDendrotion.Thispaperhighlightsthe
rich dendrotiid fauna ofthe Australasian region with 10 ofthe 21 described species found
between the KermadecTrenchinthe South Pacificandtheeast coastofAustralia. A tableof
habitat anddistributional data foralldescribed speciesofDendrotiidae ispresented.
Introduction Tenofthe21 speciesofDendrotiidaenowknown
are found between the Kermadec Trench in the
The crustacean fauna of the southeastern South Pacific and the east coast of Australia;
Australian continental shelf is species-rich eight are found on the southeastern Australian
(Barnard, 1991; Cohen and Poore, 1994; Poore continental slope.
and Wilson, 1993; Coleman et al., 1997) but Most ofmaterial onwhich this study is based
before the survey by Poore and collegues little has come from the southeastern Australian slope
was known ofthe small crustacean fauna ofthe study, 1986 and 1988 (station prefix 'SLOPE')
continental slope. Poore etal (1994) founda rich carried out by the Museum ofVictoria (NMV),
isopod fauna on the southeastern continental Melbourne (see Poore et al., 1994). Other ma-
slope, more diverse than that found on compar- terialisfromtheBass Strait Surveycarriedoutby
able slopes in the Atlantic, Arctic and Antarctic, the Museum ofVictoria and the Victorian Insti-
Dendrotiid isopods are one ofthe families con- tuteofMarine Science(WilsonandPoore, 1987),
tnreiwbutsipnegciteostohfisdednidvreorstiitiyd.iTshoipsodpsaipsebradseesdcroinbitnhge aanndd ftrhoem tNhaetuAruasltraHliisatnorMyusMeuusmeu(AmM),(BSMyNdHn)e,y
'SLOPE' collection housed mostly at the London. mm
Museum ofVictoria (Poore et al., 1994). The scale barinthe figures is 1 and refers
Dendrotiid isopods are confinedtothe shelf todrawingsofwholeanimals in dorsalviewonly.
astnuddydee1p3sesap(ec1i3e0s^48in85tmh,reeTabgleene1r).aPrhiaordtobteheins aFingdure2;labPeIls7,arepeasrefooploldosws:1A-7l;, AP2L,1-a5n,tenmnaale1
Adecsacnrtibheodm.unInnatahrise dpeaspcerirbefdo,urthenefiwrstsrpeeccoiredss ooff pmalnedoipboldes; M1P-,5;maxriMlDli,ped1;MMDX,1,riMgXht2,amnadxillleafet
the genus from Australian waters and increasing 1 and 2, and U, uropods. All illustrations are of
tfaes1hirrp8veene8ec6nti,HhueteesomfmbfriioenorsfisrmntpDehorN.eefenecrddAwoeercrsodacZatsnreniitaodobhlfneaotdnmhaidisursnpenceignsrcaeeailrnaesesusofpesirwgdoofutetrrrhseeleocuddrms.-inbtwuFheBiomeded.ubdeSredoTfraunhrrteeodhowmy,-f tg(nihSoesettnoTrephhatrooehoadlaonosr,tefpoyferAlpc1ueei9csf8tttuih8rcne)aallneeicsypkasihantosnohspkgeteeaathcnsnreiogrfoooiawnc.firlsoytefoheAseaslfttloaaurnntreedetadwhr.teoehiefsripreneeiorcuteiuhplenehasrstoinatvyrhei,esen
described species world-wide from five to nine. apposition.
—
1
B. F. C'OMEN
Table Distributional data for species ofDendrotiidae
Species Locality Depth (m)
Acanlhomunnaproleus Beddard, 1886 E ofNew Zealand, South Pacific 1281 and201
A. spinipes(Vanhoffen, 1914) OffAntarctic Peninsula 385
A. hystrix(Hansen, 1916) Iceland 1505
A. beddardi Menzies, 1962 South Atlantic 4885
A. tannerenis Schultz, 1966 TannerCanyon, North Pacific 813
A. bettongia sp. nov. Tasman Sea offVictoriaand Tasmania 695-1264
A. lagorchestes sp. nov. Tasman Sea from NSW to Bass Strait 130-429
A. macropus sp. nov. Tasman Sea offNSW and Victoria 400^29
A. potOfOUS sp. nov. Tasman Sea offPoint Hicks, Victoria 1840
Dendromunnu spinipes Menzies, 1962 South Atlantic 1816
D. mirableWolff, 1962 Kermadec Trench, South Pacific 5230-5340
D. compsa Lincoln & Boxshall, 1983 Rockall Trough, NE Atlantic 1271-2925
Dendrolion spinosum Sars, 872 Hardanger Fjord, Norway 281
1
D. parudoxum Hansen, 1916 Iceland 1600
D. hanseni Menzies, 1956 OffJamaica, Caribbean 1360
D. setosum Lincoln & Boxshall, 1983 Rockall Trough, NE Atlantic 160 and 2076
D. elegans Lincoln & Boxshall, 1983 Rockall Trough, NE Atlantic 1600 and2200
1
D. onychogalea sp. nov. Tasman Sea offPoint Hicks, Victoria 200-400
D. peradorcus sp. nov. Tasman Sea offVictoriaand Tasmania 500-2900
D. pelrogale sp. nov. Tasman Sea offNSW and Victoria 996 and 1277
D. thylogale sp. nov. Tasman Sea from NSW to Tasmania 720-1840
Dendrotiidae Vanhoffen Remarks: Wilson (1976) removed Munella
Dcndrotionidae Vanhoffen, 1914: 569. from Dendrotiidae into his newly erected
Wolff, 1962: 64.—Lincoln and Boxshall, 1983: family, Haplomunnidae. As Wilson (1976)
argued, the three remaining dendrotiid genera
298.
Munnini (part).—Hansen, 1916: 33. form a strong family, all united by the pos-
Dendrotidae Menzies, 1962a: 28. session of large uropods inserted on the dorso-
Dendrotioniidae Menzies, 1962b: 167. lateral surface ofthepleotelson. Many specimens
Dendrotiidae Bowman and Abele, 1982: 19. loose these massive uropods when they are
captured and brought to the surface. No con-
Diagnosis. Antennae long and slender. Pereon fusion with species of the genus Munella
bearing spines or long setae dorsally. Mouthparts should arise becausea large pairofsocketsonthe
generally not modified except maxilliped palp pleotelson indicate where dendrotiid uropods
articles narrow. Pereopod 1 prehensile, shorter attach; species of Munella lack these large
than 2 7. Pereopods 2-7 ambulatory, long and sockets.
slender; pereopod7absent in some species. Coxal Some confusion exists in the literature con-
plates maybe visible dorsally, extended in some cerning the correct spelling of the family name
species into long lateral projections. Uropods Dendrotiidae. Bowman and Abele (1982) intro-
large, biramous and inserted on dorsolateral sur- duced the correct spelling (Dendrotiidae) without
face ofpleotelson. Uropod insertion point marked explanation. Dendrolion is probably a conjunc-
by large socket on the pleotelson. Pleotelson ex- tion ofdendro- (branching) and the Greek word
tending beyond insertion point ofuropods. Anus -otion (little ear). As family names are made of
opening into brachial chamber. Plcopods gener- the Latinised genitive root, oti- in this case, the
ally typical ofAscllota except male pleopod 2 in correct family name is Dendrotiidae (G.D.F.
some specieswith extremely longpenial filament. Wilson, pers. comm.).
DENDR0T1IDISOPODS FROM THE AUSTRALIANCONTINENTAL SLOPH
Key to genera ofDendrotiidae
Cephalon with prominent lateral process supporting antennae; antenna
I
basal articleelongate(morethan 5 times as long as broad) Dendrotwn
Cephalon without prominent lateral process supporting antennae; antenna
1
basal articlenot elongate (less than 2 times as long as broad) 2
2. Eyespresent, pereon at most bearingmany small spines, percopod 7 present
in adults Acanthomunna
Eyesabsent, pereon bearing few large dorsolaterallydirected spine-like pro-
cesses,eachbearingapicalclusterofspines, pereopod 7absent inadults
Dendromunna
Acanthomunna Beddard Male pleopod subrectangular. Male pleopod
I
Acanthomunna Beddard, 1886a: 102. Beddard, 2 peduncle tapered; exopod bilobed. Some
1886b: 47.—Menzies, 1962b: 174.—Wolff, species, with extremely longappendix masculina.
1962: 65.—Schultz, 1966: 6.—Lincoln and Uropods large and robust with numerous spini-
Boxshall, 1983: 309. form setae, inserted posterolaterally on dorsal
Moromunna Vanhoffen, 1914: 569. surface; rami subequal, peduncle obvious.
Pseudomunna Hansen, 1916: 47. Remarks. Priortothis study only 12 specimensof
Tvpe species. Acanthomunna proteus Beddard, Acanthomunna had been collected, three of the
1886. five species known from only one specimen: A.
hystrix (Hansen, 1916); A. beddardi Menzies,
Diagnosis. Eyes present, located on small lateral 1962; A. tannerenis Schultz, 1966. A. proteus
protuberances. Cephalon broader than long, nar- Beddard, 1886 is known from two specimens and
rower than pereon. Antennae extremely long and A. spinipes(Vanhoffen, 1914)knownfrom seven.
slender. Antenna 1 basal article stout, not more The four new species represent the first substan-
than twice as long as wide; third basal article tial collection ofindividualsofAcanthomunna.A.
longer than first or second. Antenna 2 as long as beilongia sp. nov. is represented by over 200
animal and twice as long as antenna 1; peduncle specimens and A. lagorchestes sp. nov. by over
articles 1-3 short, articles 1 and 2 with spiniform 100 specimens.
setae; peduncle articles 4 and 5 extremely long Wilson (1976) suggested a phylogeny of the
and slender. Pereon oval; 1.5 to 2 times as broad Dendrotiidaeanda closely related family, Haplo-
as long at broadest point; all pereonites and pere- munnidae. He argued thatAcanthomunna, occur-
opods free and present. Mouthparts typical of ingin shallowwaters, istheleastderivedgenusof
Asellota, left mandible not stronger than right. these two families because they have retained
Maxillipedal epipod broad and flat, distally their eyes. The other genera have moved into
tapered; endite internal margin reflexed; palp deeperwaters and lost theireyes. A. lagorchestes
much narrower than endite, of 5 articles. Pereo- sp. nov. isthe only dendrotiid recorded from less
pod 1 prehensile, reflexed between carpus and than 150 m (130-429 m. Table I) although most
propodus, shorter than other pereopods. Pereo- individuals were caught around 400 m depth. A.
pods2-7ambulatory; longandslender, becoming macropus sp. nov. was also caught around 400 m
successively more elongate; basis, ischium and (400-^(29 m). A fifth species of Acanthomunna
merus compact, carpus and propodus greatly was captured from 200 m (stn SLOPE 21, NMV
elongate; ischium to propodus articles with spini- J18589) but was too badly damaged to be
form setae along anterior and posterior margins. described.
Key to species ofAcanthomunna
Pereonites 3 and4 with spiniform setae on middorsal surface 2
Pereonites 3 and 4 devoid of setae on middorsal surface (may have small
spinesnearlateral marginsofdorsal surface) 5
2. Cephalon with spiniform setae 3
Cephalon devoid ofspiniform setae 4
3. Spiniform setae branched A. proteus Beddard. 1886
Spiniform setaenot branched A. beddardi Menzies, 1962
COHEN
B. F.
4. Pereonite 1 with a single row of4-5 large spiniform setae
A.potoroussp. nov.
Pereonite 1 with 2rowsofmorethan5 large spiniform setae
A. tannerensis Schultz, 1966
5. Pereonite 1 with morethan 8dorsal spines A. hystrix(Hansen, 1916)
Pereonite 1 with4 orfewerdorsal spines 6
6. Pereonite2devoidofspiniformsetae A. bettongiasp. nov.
Pereonite 2 with spiniform setae 7
7. Pereonite 1 with4spines A, spinipes(Vanhoffen. 1914)
Pereonite 1 with 3 spines A. lagorchestes sp. nov.
Pereonite 1 with 2 spines A. macropus sp. nov.
Acanthomunna bettongia sp. nov. pereonites 5-7 compressed, successively becom-
ing more posteriorly projected. Pereonites 2—4
Figures 1-3
with short spine-like extensions of posterior
Material examined. Holotype. Tasmania. 48 km ENE lateral margins. Lateral margins of pereonite
ofCape Tourville (42°00.25'S. 148°43.55'E), 1264 m, 7 hidden dorsally by pleotelson; coxae visible
gravel with lumps of sandy mud aggregate, WHOI dorsahy on other pereonites, some with small
eOpcitbe1n9t8h8i,csstlnedS,LGO.PC.EB.81P,oNorMeVetJal36o9n84RV(feFmraalnek)l.in, 30 tapered projections. Lateral margins of pereo-
Paratypes. Type locality,NMVJ36985 (1 male). nites and coxae with numerous fine scales as
Victoria. SofPointHicks(38°16.40'S, 149°27.60'E), found on anterior margin of cephalon. Pereo-
800 m, coarse shell, biogenic seds, WHOI epibenthic nite 1 with transverse dorsal ridge supporting
sled, M.F. Gomon et al on RV Franklin, 23 Jul 1986, 2 diverging anterolateral spines and a bifid
stn SLOPE 34, NMVJ36983 (1 female). anteriorly directed middorsal spine; remain-
Other material. Tasmania. Off Freycinet Peninsula, ing pereonites smooth, devoid of obvious
WHOI epibenthic sled. M.F. Gomon el al. on RV spination.
Franklin,27Jul 1986(42°2.20'S, I48°38.70'E), 800m,
c(4o2a°r0s.e20s'hSe,ll1y48s°a3n7d.,70s'tEn),SL72O0PmE,4c5o,arNseMsVhelJl1y8s5a7n2d,(s4t)n; triPalnegounlairt,es1.f2usteidmetsolpolnegoetreltsohna.n wPliedoet;elwsiotnhspuabi-r
SLOPE 46, NMV J18573 (I). Type locality: NMV of large anteriorly directed spines on anterior
J18574(63), NMVJ18575 (64). third ofdorsal surface anda sulcusrunningalong
Eastern Bass Strait,naturalists' dredge,G.C.B. Poore lateral marginto posterolateral bosses which sup-
on HMAS Kimbla, Mar 1979: 55 km NE of Babel port uropods; ventrolateral margins with 3 short
sItsnlaBndSS(3394°,38N.2M'SV, 1J4188°5497.62'(E1)0),,6N95MmV,Jr1o8c5k7-7sa(n1d0-)m;u8d7, mspiinnuitfeorsmpisneitfaoe;rmpossetteareiolratmeraarlglyinanwditah carepnauilratoef
k14m8°5E2.N4E'E),of84N1ormt,hmuPodidnty, saFnldi,ndestrns BIS.S(3397°,28N.2M'SV, muraorpgoidnal bboestswese.en setae, extending beyond
J18Vi5c7t8or(i1a.).S ofPoint Hicks, WHOI epibenthic sledon Left mandibular incisor process 3-dentate,
RV Franklin, 23 Jul 1986: 38°21.90'S, 149°20.00'E, lacinia mobilis 2-3-dentate; right mandibular
1000 m, G.C.B. Poore et al., stn SLOPE 32, NMV incisor process 4-5-dentate. Left mandi-
J18568 (18); 38°19.60'S, 149°24.30'E, 930 m, rock, bular spine row with 9 armed spines; right
rubble, clay, sand, biogenic sediment, M.F. Gomon et mandibular spine row with 9 armed spines. Left
al., stn SLOPE 33, NMV .118569 (16); 38°16.40'S, mandibular molar broad, flat, positioned against
NM1.4MF9V°.J27G1.o860m5'o7En.18et00al.m,,stcnoaSrsLeOPshEell3,4,biNogMenVicJs1e8d5i7m0en(t8s), lmaacnidniiabulmaobrilimso;lawrithbro5adspiannidforcmonsceatvaee.; Rwiigthht
(7). numerous spiniform setae. Mandibular palp of
Description. Total length of holotype 2.94 mm. 3 elongate articles, typical. Maxilla 1 outer lobe
Cephalon subrectangular, twice as long as wide; bearing 1 1 stout spines, 7 denticulate; inner
anterior margin with middorsal notch between lobe with numerous setae. Maxilla 2 with stout
base ofantenna, covered with many tine scales. setae on all lobes; outer lobe with 2 simple and
Antenna 1 flagellum with 10 articles with 9 aes- 2 denticulate setae; middle lobe with I simple
thetascs located on all flagellum articles except and2 denticulate setae and innerlobe with 5 den-
basal three. Antenna 2 flagellum with numerous ticulate setae. Outerand middle lobes with a row
articles ofsubequal sizes. of simple setae on inner lateral margins. Maxil-
Pereon oval, widest at pereonite 3. Perco- liped with 3 coupling hooks; endite distal margin
nites 1-4 separated by narrow dorsal sulcus; with 7 denticulate setae.
DENDROTIID ISOPODS FROMTHEAUSTRALIAN CONTINENTAL SLOPE
Figure I. Acanthomunna bettongia. Holotype NMV J36984; A2 ofparatype NMVJ36983.
COHEN
B. F.
Figure 2. Acanthomunna bettongia. Holotype NMVJ36984; PL1-5 ofparatype NMV J36985.
DENDROTIID ISOPODS FROM THEAUSTRALIAN CONTINENTAL SLOPE
NMV
Figure 3. Acanthomunna bettongia. Holotype J36984.
COHEN
B. F.
mm
Pereopod 1 propodus with row of5 stout setae Description. Total length of holotype 2.20
on posterior margin; carpus with 2 rows ofstout Cephalon quadrate,as longas wide; anteriormar-
setae; other articles also with spiniforni setae. gin straight, produced between base ofantenna.
Pereopods 2 7 typical, merus and carpus with Antenna 1 flagellum with 5 articles with 3 aes-
large stout setae and dactylus with 7-8 spiniforni thetascs located on 3 distal articles. Antenna 2
setae posteriorly. flagellum with 18 articles.
Male pleopod 1 with rounded distal margin Pereon oval, widest at pereonite 3. Pereonites
bearing 7-8 long setae on each side. Male pleo- 1^ with narrow dorsal sulcus separating pere-
pod 2 proximal lobe small, knob shaped; distal onites; pereonites 5-7 compressed, successively
lobesetose;appendix masculina(endopod) small, becoming more posteriorly projected. Lateral
not reachingendofpeduncle. Pleopod 3 endopod margins of pereonite 7 hidden dorsally by pleo-
reaching beyond first article of exopod, with 3 telson; coxae visible dorsally on other pereonites,
longcompoundsetaedistally; exopodwith 1 long all with small lateral projections. Pereonite 1 with
setae. Pleopod4 endopod longerthan exopodand middorsal ridge supporting 3 small spines; pere-
2-3 times as broad. Pleopod 5 with no rami, onite 2 with middorsal ridge with six small pro-
lateral and distal margins crenulate. Uropod jections which appear to be the attachments for
typical; exopodas longas peduncle, endopodone setae; remaining pereonites smooth, devoid of
third longerthan exopod. obvious dorsal spination or setae. All pereonites
Distribution. Tasman Sea off Victoria and except pereonite 7 with 1 or 2 pairs of small
Tasmania. 695-1264 m depth. projections near lateral margins.
Pleonite 1 visible, pleonites 2-5 fused to pleo-
Remarks. Aeanthoimmna bettongia is most read- telson. Pleotelson subtriangular ventrally with
ily identified by the bifid, anteriorly directed pronounced posterolateral bosses protruding dor-
middorsal spine on pereonite 1 and the pair of sally; 1.1 times longer than wide; distal margin
anteriorly directed large spines on anterior third rounded; devoid ofany spination orsetae.
of dorsal surface of the pleotelson. A. bettongia Left mandibular incisor process 6-7-dentate,
appearstobeacommonspeciesontheuppercon- Iacinia mobilis 1-2-dentate; right mandibular
tinental slopeofsoutheastern Australia. Over200 incisor process 4-dentate. Left mandibular spine
specimenswerecollected fromthe SLOPE survey. row with 3 spines, simple; right mandibular
spine row with 5 spines, most armed. Left
Aeanthoimmna lagorchestes sp. nov. mandibular molar broad and flat, split into two.
Figures4-6 Right mandibular molar broad and flat. Mandi-
Material examined. Holotype. S of Point Hicks bular palp represented by single setae. Maxilla 1
(m3u8d',17.m7a0'nSy, 1s4p9onDg1e1s.,30'EW)I,I4O0I0 mc,pibceonatrhsiecsasnlde,d,graMv.eFl., olauttee;rilnonbeer wliotbhe 9wivtihsinbluemestroouutsspsientease,. 3Madxeinltliacu2-
Gomon etal. on RVFranklin,24 Jul 1986. stn SLOPE with stout setae on inner lobe; outer and middle
40. NMV.136980(female). lobes with a row ofsimple setae on inner lateral
Paratypes. Victoria. Type locality, NMV J36982 margins. Maxilliped with 2 coupling hooks;
(1 male). enditedistal margin with 6 denticulate setae.
eO1pc5itO1Nb°tee0h1nw89et.8rh08Si6,co'muEsastt)tlnhe,erdSi.Wa4LalG2lO..9eCPs.E.NBme.,45w64P,omkoNuSrmdoMeudVtEeythoaJtfc3.Wo6Naao9lorn8esw1seRr.Va(1shOF(efrf3leal4fm.n°ak5lNl5eWi.o)n7Hw..9rO'a2SI.2, bosopntiPnhpeiormfseaotorrepgmrioiondsrse;t1mapoae.trrhgoeipPrnoe;darruetcsoiapcrwlopeidsutsshalr2wsi-oot7wshuotpsfyptp4ooiurctstatlio,snuegttmaaseeefrteuoawsne
WHOI epibenthic sled. G.C.B. Poore et al. on RV and carpus with large stout setae and dactylus
Franklin: 34°59.52'S, 151°5.94'E, 204 m, coarse shell, with 5-6 spiniform setae posteriorly.
14 Jul 1986. stn SLOPE 1. NMV J18579 (1): Male pleopod 1 with rounded distal margin
34°55.79*S, 151°08.06'E, 429 m. muddy coarse shell, bearing 14 long setae on each side distal to
22 Oct 1988. stn SLOPE 56, NMV J18581 (37), NMV oblique groove. Male pleopod 2 proximal lobe as
J18582(38). asmall roundedknob, distal lobe setose; appendix
Tasmania. Eastern Bass Strait, 100km NEofNorth masculina (endopod) not long, extending just
sPBaoSniSndt..1R7F.0lS.i.nNdWeMirlVss.o1.n11(o83n85°85R32(V.26)'T.Sa,ng1a4r8o°a2,5.2'15E)N.o1v3019m8.1.fsitnne bhaelyfonwdaypeadluonncgles.ecPolnedoparotdicl3e oefndeoxpoopdod,rewaicthhin3g
Victoria. Type locality, NMV .118580 (20). 50 km S longcompound setaedistally; exopodwith 1 long
ofMallacoota (38°06.2'S, I49°45.5'E). 188 m. WHOI setae. Pleopods 4-5 similar to A. bettongia but
esptinbSe0n5thi8c4s3l0e.d.NRM.SV. WJi1l8s5o8n4o(nI).RVSoela, 14Oct ll)84. wmiatrhgincsoncsamvoeotdh.depUrreospsoiodntycpeinctarla;llye;ndpolpeoodpodone5
DENDR0T1ID ISOPODS FROM THE AUSTRALIAN CONTINENTAL SLOPE
Flgure 4. Acanthomunna lagorchestes. Holotypc NMV J36980; U, PI, Al and A2 ofparatypc NMV
J36981.
COHEN
B. F.
Figure 5. Acanthomunna lagorchestes. P2-6 of paratype NMV J36981; PL1-5 of paratype NMV
J36982.