Table Of ContentBonner zoologische Beiträge Band 53 (2004) Heft 1/2 Seiten 1-11 Bonn, Juni 2005
Clas M. Naumann (26.06.1939 - 15.02.2004)- in memoriam
After several years of suffering from cancer, Professor wieg. Certainly through these contacts, Clas Naumann's
Clas Michael Naumann zu Königsbrück died on Sun passion for butterflies and moths developed in those
day, 151h of February, 2004. He was a füll professor of years. He even wrote as bis first publication a short re
Systematic Zoology at the Rheinische Friedrich port on the Lepidoptera found in the sunoundings of
Wilhelms-Universität and director of the Zoologisches Maria Taferl in Austria, where he had spent a summer
Forschungsinstitut und Museum Alexander Koenig, holiday with his parents (in 1958, no. 1). In 1959, he
Bonn. passed the final secondary-school examinations. After
three trimesters of "studium generale" at Leibniz Kol
leg, Tübingen, he studied chemistry from 1960 to 1962,
and biology from 1962 to 1966 at the Eberhard-Karls
Universität in Tübingen. Here, his entomological inter
ests focussed on Bumet Moths, especially of the genus
Zygaena.
During his time in Tübingen, he undertook his first en
tomological trip through a country of the N ear East, and
subsequently published bis first species description: Zy
gaena problematica Naumann, 1966, from southem
Turkey (no. 2). He also made first experiences as a
businessman since he founded a trade of entomology
supply. In his later days, he used to tel! about this era of
bis life with great delight.
In 1967, he switched to the Rheinische Friedrich Wil
helms-Universität in Bonn. Here, he received his doc
Fig. 1. Professor Clas Michael Naumann zu Königsbrück, toral degree in 1970 after doing bis doctoral dissertation
June 12, 1995 (Photo M. Schmitt) at the Zoologisches Forschungsinstitut und Museum
Alexander Koenig on the systematics and phylogeny of
Clas Michael Naumann zu Königsbrück was bom the the holarctic Sesiidae - Insecta, Lepidoptera, under the
26th of June 1939, in Dresden as the only child of Dr. supervision of Günther Niethammer (published in 1971,
Eberhard Naumann zu Königsbrück and Freda-Irene, no. 6). Subsequently, he took up an opportunity to work
nee Hannemann. He spent the first years of his life at for three years as a university docent at Kabul Univer
the family estate (Königsbrück, near Dresden) from sity (Afghanistan). Besides his teaching, he built up a
where he could hear and see the bombing of Dresden zoological collection and a small university museum of
(13th/14th of February 1945), an impression he pre zoology. The three years he sperrt in Afghanistan were
served weil, as he recounted some fifty years later. He most influential upon his further life and scientific ca
fled from the approaching Red Army with his family in reer. He took more than just a fancy for the nature, cul
Spring 1945, another experience which left a lasting ture, history and people of that country. He leamed the
1. mpress1. 0n. language(s) ofthis country and did extensive field work,
even in quite remote parts of the country, such as Nuris
His family found a provisional home in the village of
tan and Registan, the provinces ofBadakhshan and Pak
Kirchohsen near Hameln, where Clas Naumann entered
tia, as well the Wakhan Valley in the Afghan Pamirs
primary school. From 1949, he went to high school in
(Naumann 1974, nos. 11-14). His strong affection for
Wilhelmshaven for two years, and finally in Braun
Afghanistan and the whole region of the Middle East
schweig, where his father, who had a doctoral degree in
lasted for the rest of his life.
agriculture, found a position with the Bundesforschungs
anstalt für Landwirtschaft (Federal Research Institute of He retumed to Bonn in 1973 and received a two-year
Agriculture). From his father, who was an ardent beetle post as an assistant at the Institute for Applied Zoology
co!lector, he was guided into natural history. Already (Professor Dr. Werner J. Kloft) in Bonn, followed by
during bis high school years, he made contact with local three years at the Ludwig-Maximilians-Universität in
entomologists, especially lepidopterists, e.g. Fritz Hart- München (Zoological Institute, Professor Dr. Jürgen Ja-
2 Bonner zoologische Beiträge 53 (2004)
cobs ), where he received his habilitation in 1977. His the state ministries of Bavaria and Nordrhein-Westfalia,
Habilitationsschrift (professorial dissertation) was on he managed to convince the govemment of the latter
the phylogeny and zoogeographical relations of the Zy state to guarantee a new building, which would entice
gaenini, published also in English in 1977 (no. 21). The him to remain in Bonn. He was intensively involved in
same year he was appointed a professor at Bielefeld the detailed planning and invested a good deal of hope
University, where he chaired the Department of Mor and energy in this enterprise. However, the construction
phology and Systematics of Animals until 1989. He ofthis new building began only the day after he died.
continued working on morphology, taxonomy, phylog
As a director, Clas Naumann was always concemed
eny and zoogeography of the Zygaenidae, but extended
about the welfare of the museum's staff members, the
his field of work into chemical ecology and sensory
countless colleagues who worked at the museum on
physiology. A quickly growing number of enthusiastic
grants, and of the students doing their theses at the mu
diploma and doctoral students soon formed a vivid and
seum. And while he mostly did not show much of his
prolific team of investigators and authors.
emotions, he could be as merry as a lark when he man
On July 1, 1989, he was appointed a füll professor at the aged to do something good to somebody, especially
Rheinische Friedrich Wilhelms-Universität in Bonn, and when it came as a surprise. Possibly, these were the
at the same time director of the Zoologisches For most rewarding moments in his daily work.
schungsinstitut und Museum Alexander Koenig. Under
Clas Naumann was a dedicated academic teacher who
his directorship, a corporate plan was elaborated that
also stimulated younger scientists to give university
provided the institution with an intemal structure, i.e.
courses. In many cases, he announced these courses
departments (vertebrates, entomology, exhibition and
only formally, while in fact they were given by his doc
public relations) and sections. He initiated monthly sci
toral students. Thus, he not only provided these younger
entists' meetings, weekly informal meetings of all staff
colleagues the opportunity of gaining experience in
members, and he made the regular meeting of the heads
teaching, but also considerably enhanced the range of
of the three departrnents an effectively working and
courses for students of zoology at the Bonn University.
well-accepted board of directors. His main goals were to
In addition, he supervised numerous diploma- and doc
increase the scientific output and to improve the interna
toral theses (see !ist below), both at Bielefeld and Bonn.
tional standing of Museum Koenig, and to enhance its
attractiveness to the public. In all of these activities and Systematists in Central Europe, zoologists and espe
throughout the whole of his decisions as Director, he cially museum entomologists, owe Clas Naumann a
strove for a consensus among the parties involved. And great deal for his never ending efforts for a better lobby
it caused literally anguish to him when he did not suc for our field of science within Germany's political and
ceed in reaching consensus. general scientific frame. He was one of the promoters of
the Gesellschaft für biologische Systematik (Society for
Clas Naumann clearly saw that the museum's infra Biological Systematics) founded December 12, 1997,
structure had to be improved drastically in order to and organised its first annual meeting at Museum
reach his goals. So, he worked inconceivably hard to Koenig in 1998. He strove for more co-operation be
acquire additional fünds, be it for additional technical tween the German museums with vast holdings of scien
equipment such as a scanning electron microscope or a tific collections, was one of the founders of the DNFS
bone-degreasing device, or for a total renovation of the (Direktorenkonferenz Naturwissenschaftlicher For
museum's basement, or for the redesign of its public schungssammlungen in Deutschland), and he served for
exhibition. Under his guidance, an entirely new concept several years as a referee for the Deutsche Forschungs
for the exhibition was elaborated. With a considerable gemeinschaft (German Research Foundation). Through
amount of "soft" money, the reorganisation of the exhibi his tireless lobbying, his own scientifi.c papers and
tion was achieved, at least partially, before his untimely through the numerous diploma and doctoral theses he
death. However, most tragically, he was already too feeble stimulated and supervised, Clas Naumann contributed
to attend the opening ceremony for the first redesigned immensely to the revitalisation and the expansion of
portion of the exhibition, on the 9th of October 2003. taxonomy and systematics in Germany.
One of his most notable merits for Museum Koenig was Among Clas Naumann's super-regional scientific activi
that he succeeded in reaching an administrative decision ties, his efforts for the joumal zoologischer Anzeiger
from the state of Nordrhein-Westfalia and the German played an outstanding part. He was editor-in-chief since
Federal Govemment for a new building. He would defi 1995. Without his energy and optimism, this joumal
nitely not have succeeded without the recruitment to ac would certainly have ended in 1994, when Dr. Johanna
cept a position of füll professor at the München Univer Schlüter from Gustav Fischer Verlag, Jena (the publish
sity and at the same time director of the Zoologische ers at that time) asked Professor Naumann to accept
Staatssammlung München. When negotiating with both the task of saving the oldest German zoological jour-
Clas M. NAUMANN: in memorian 3
nal. The present profile of the joumal and the boards of
editors and advisors still bear Professor Naumann's
mark.
Also, Clas Naumann was very much interested in im
proving the museum's own publication series. lt was his
explicit desire that Bonner zoologische Beiträge should
grow into an intemationally renowned, fully peer re
viewed joumal of high standards in appearance and
content. We, the team of editors and advisors since vol
ume 51, are proud to say that Clas Naumann was highly
satisfied when he saw the newly designed issues of
vol. 51.
Clas Naumann worked tremendously hard for the good of
Museum Koenig, for all the persons working here, for a
better standing of the science region of Bonn in a wider
context, for taxonomy and systematics, and for science
in general. He did so finally at the cost ofhis private life
and even his health. Who ever knew him weil enough to
be able to judge, and close enough to care, felt relieved Fig. 2. Clas Naumann on the Isle of Skye (Scotland), during
an International Zygaenidae Symposium, September 11, 1996.
when he informed us about his decision to retire ahead
Left: Gerhard Tarmann (Innsbruck), right: Jean-Marie Desse
of schedule in April 2004, and he and we hoped he had
(Angers) (Photo C. Esch, Münster).
some time to devote to his family and his hobbies. But
regrettably, he passed away some two months before
Scientists, technicians, other staff members and students
that date. Thus, he could neither live to see the new
of the Zoologisches Forschungsmuseum Alexander
building being completed nor could he introduce his
Koenig are deeply indebted to Clas Naumann for his
successor smoothly into his duties, as he wished.
tireless efforts. The board of directors and scientific
staff unanimously decided to dedicate the new building
Two fields of interest absorbed Clas Naumann espe
to the memory of their former director and to name it
cially besides his scientific and administrative work:
"Clas M. Naumann-Bau".
Bumet Moths and Afghanistan (and adjacent countries).
In his spare time and on several expeditions, he col Clas Michael Naumann zu Königsbrück is survived by
lected and investigated Zygaenidae. lt was one of his bis wife Storni Naumann-Nawabi and his children Rox
highest satisfactions to discover a new species or to re ana and Alexander, to whom we express our deep sym
discover a lost one. He published numerous taxonomic, pathy.
morphological and zoogeographical studies on that
group of moths, and he enjoyed meeting other Zygaeni
Acknowledgements. Storni Naumann-Nawabi (Wacht
dae enthusiasts (Fig. 2).
berg-Pech) read the manuscript of this obituary and relin
quished a !ist of her husband's publications to me, Renate
Until the last weeks of his life, he was very much con
Feist (Bielefeld) compiled a !ist of Clas Naumann's di
cemed about the political, social and scientific devel
ploma and doctoral students in Bielefeld, Christoph Häuser
opments in Afghanistan, especially at the University of
(Stuttgart) provided me with a draft of an extensive necrol
Kabul. While many of his public lectures were either ogy meant for a lepidopterists' joumal, Christoph Esch
characterised by a certain scientific distance or by tacti (Münster) gave permission to use the photograph ofFigure
cal considerations (e.g., in order to convince his audi 2, Axel Hofmann (Breisach) identified the persons showed
ence of a certain goal), he gave an entirely different im on this photo, and Brad Sinclair (Bonn) improved the Eng
pression when giving a talk on countryside, people, lish of my manuscript.
culture and history of Afghanistan. His voice, his smile,
bis gesture changed. No doubt was left that this was
his true passion. He was significantly involved in
the revitalisation of the partnership between the Rhei
nische Friedrich-Wilhelms-Universität Bonn and the
Kabul University, and he invested not only just some
thoughts in writing several proposals, but he travelled to
Kabul and after returning organised campaigns for as
sistance.
4 Bonner zoologische Beiträge 53 (2004)
PUBLICATIONS OF CLAS M. NAUMANN 18 NAUMANN, C. M. (1977): Biologie, Verbreitung und
(AS OF 01.02.2005, COMPILED BY MICHAEL Morphologie von Praezygaena (Epizygaenella)
caschmirensis (Kollar, 1848) (Lepidoptera, Zygaeni
SCHMITT)
dae). Spixiana 1: 45-84.
1 NAUMANN, C. (1958): Schmetterlinge aus der Umge 19 NAUMANN, C. M. (1977): Stammesgeschichte und
bung von Maria-Taferl (Niederösterreich). Zeitschrift tiergeographische Beziehungen der Zygaenini (Insecta,
der Wiener entomologischen Gesellschaft 43: 33-36. Lepidoptera, Zygaenidae). Mitteilungen der Münchner
2 NAUMANN, C. (1966): Zygaena (Subg. Zygaena F.) entomologischen Gesellschaft 67: 1-25.
problematica nov. sp. Zeitschrift der Wiener ento 20 NAUMANN, C. M. (1977): Rasterelektronenoptische
mologischen Gesellschaft 51: 10-19. Untersuchungen zur Feinstruktur von Lepidopteren
3 NAUMANN, C. (1969): Zur Kenntnis der Mesem Gespinsten. Mitteilungen der Münchner entomolo
brynus-Arten Afghanistans (Lep., Zygaenidae). Bon gischen Gesellschaft 67: 27-37.
ner zoologische Beiträge 20: 266-278. 21 NAUMANN, C. M. (1977): Studies on the Systematics
4 NAUMANN, C. (1970): Bemerkungen zur Biologie und and Phylogeny of Holarctic Sesiidae (Insecta, Lepi
zur systematischen Stellung von Zygaena graslini doptera) [Untersuchungen zur Systematik und Phy
Lederer, 1855 (Lep., Zygaenidae). Bonner zoologische logenese der holarktischen Sesiiden (Insecta, Lepidop
Beiträge 21: 133-136. tera)]. 139 pp., Amerind Publishing Co. Pvt. Ltd., New
5 NAUMANN, C. (1971): 6. Schmetterlinge. Pp. 547-549 Delhi, Washington.
in: NIETHAMMER, G.: Die Fauna der Sahara und ihrer 22 NAUMANN, C. M., JUNGE, G. & ROSE, K. (1977): Ein
Randgebiete, Darstellung eines Naturgroßraumes, Bd. 1: weiterer Beitrag zur Zygaenen-Fauna Kappadokiens
Physiogeographie. München (Weltforum-Verlag). (Lep. Zygaenidae). Atalanta 8: 282-289.
6 NAUMANN, C. (1971): Untersuchungen zur Systematik 23 NAUMANN, C. M., & NAUMANN, S. (1978): Eine neue
und Phylogenese der holarktischen Sesiiden (Insecta, ostanatolische Unterart der Zygaena (Zygaena) loti
Lepidoptera). Bonner zoologische Monographien 1: 1- ([Denis und Schiffermüller], 1775) (Lep., Zygaeni
133. dae). Entomologische Zeitschrift Frankfurt am Main
7 NAUMANN, C. M. & NOGGE, G. (1973): Die Zoolo 88: 37-40.
gischen Projekte Afghanistans - erfolgreiches Neben 24 NAUMANN, c. M., Huss, H. & NAUROZ, M. K. (1978):
produkt einer Universitätspartnerschaft in Afghanistan. Zur Tierwelt des "Großen Pamir". Pp. 202-213 in: DE
GA WI-Rundbrief 4/73: 25-28. GRANCY, R. S. & KOSTKA, R. (eds): Grosser Pamir.
8 NAUMANN, C. & NOGGE, G. (1973): Die Großsäuger Verlagsanstalt, Graz.
Afghanistans. Zeitschrift des Kölner Zoo 16: 79-93. 25 NAUMANN, C. M. & RACHELI, T. (1978): Taxonomie
9 NAUMANN, C. & NIETHAMMER, J. (1973): Zur and ecological observations on Zygaena F. from the
Säugetierfauna des afghanischen Pamir und des Wak Province of Fars, Southem Iran (Lepidoptera, Zygae
han. Bonner zoologische Beiträge 24: 237-248. nidae ). Atalanta 9: 208-218.
10 NAUMANN, C. M. (1973): Ein ehemaliges Wildyak 26 NAUMANN, C. M. (1978): Zur Systematik, Verbreitung
Vorkommen im afghanischen Pamir. Bonner zoolo und Biologie von Zygaena pamira Sheljuzhko, 1919
gische Beiträge 24: 249-253. (Lepidoptera, Zygaenidae). Atalanta 9: 355-372.
11 NAUMANN, C. M. (1974): Beobachtungen über den 27 NAUMANN, C. M. (1979. Hans Burgeffs Beitrag zur
Verlauf einer Lepidopteren-Immigration in Afghani Zygaenenkunde. Atalanta 10: 85-88.
stan 1972. Atalanta 5: 82-88. 28 RACHELI, T. & NAUMANN, C. M. (1979): On Zygaena
12 NAUMANN, C. M. (1974): Pamir und Wakhan - Kurz Fabricius (Zygaenidae) from Fars, south Iran. Nota
bericht zweier Expeditionen ( 1971 und 1972) nebst lepidopterologica 2: 53-55.
einigen allgemeinen Bermerkungen. Afghanistan 29 NAUMANN, C. M. & BENDER, R. (1980): Eine neue
Journal 1: 91-104. Arctia intercalaris-Subspezies aus dem Hindukusch
13 NAUMANN, C. M. (1974): Neue Zygaena-Unterarten (Lepidoptera, Arctiidae) . Atalanta 11: 21-28.
aus Afghanistan. Entomologische Zeitschrift Frankfurt 30 NAUMANN, C. M. & EISNER, C. (1980): Pamassiana
am Main 84: 29-36. Nova LVII. Beitrag zur Ökologie und Taxonomie der
c.
14 NAUMANN, & NIETHAMMER, J. (1974): Neu afghanischen Pamassiidae (Lepidoptera). Zoologische
nachweise von Säugetieren aus dem nördlichen Af Verhandelingen Leiden 178: 35 pp., 9 pls.
ghanistan. Säugetierkundliche Mitteilungen 22: 295- 31 NAUMANN, C. M. & TREMEWAN, W. G. (1980): On
298. the biology of Zygaena (Mesembrynus) tamara Chris
15 NAUMANN, C. M. & NAUROZ, M.K. (1975): Be toph, 1889 (Lepidoptera: Zygaenidae) . Entomologist's
merkungen zur Verbreitung des Markhors, Capra fal Gazette 31: 113-121.
c.
coneri (Wagner, 1839) in Afghanistan. Säugetierkund 32 NAUMANN, M. & NAUMANN, S. (1980): Neue Zy
liche Mitteilungen 23: 81-85. gaenen aus den Aufsamrnlungen W. Eckweiler's aus
16 NAUMANN, C. M. & SCHULTE, A. (1977): Eine weitere dem Iran (Lepidoptera Zygaenidae). Mitteilungen der
neue Subspecies von Zygaena (Agrumenia) shivacola Basler entomologischen Gesellschaft N. F. 30: 47-54.
Reiss & Schulte, 1962 (Lepidoptera, Zygaenidae). En 33 NAUMANN, C. M. & NAUMANN, S. (1980): Ein Beitrag
tomologische Zeitschrift Frankfurt am Main 87: 89- zur Kenntnis der Zygaenen-Fauna Nord- und Ost
93. Anatoliens (Lep„ Zygaenidae). Entomofauna 1: 302-
17 NAUMANN, C. M. (1977): Zygaena (Mesembrynus) 353.
halima n.sp. und einige Bemerkungen zur stammes 34 NAUMANN, C. M. & SCHROEDER, D. (1980): Ein
geschichtlichen Gliederung der Gattung Zygaena F. weiteres Zwillingsarten-Paar mitteleuropäischer Se
(Lepidoptera: Zygaenidae). Zeitschrift der Arbeitsge siiden: Chamaesphecia tenthrediniformis ([Denis &
meinschaft österreichischer Entomologen. 29: 35-40. Schiffetmüller], 1775) und Chamaesphecia empi-
Clas M. NAUMANN: in memorian 5
formis (Esper, 1783) (Lepidoptera, Sesiidae). Fabricius, 1775 (Lepidoptera, Zygaenidae). Theses
Zeitschrift der Arbeitsgemeinschaft Österreichischer Zoologicae 5: 1-97 (Verlag J. Cramer, Braunschweig).
Entomologen 32: 29-46. 51 FRANZL, S. & NAUMANN, c. M. (1984): Morphologie
35 NAUMANN, C. M. & ROSE, K. (1981): Eine bisher un und Histologie der Wehrsekretbehälter erwachsener
bekannte peucedanoide Population der Zygaena Raupen von Zygaena trifolii (Lepidoptera, Zygaeni
ephialtes (Linnaeus, 1763) aus Griechenland (Lepi dae ). Entomologische Abhandlungen und Berichte aus
doptera, Zygaenidae). Entomofauna 2: 113-124. dem Staatlichen Museum für Tierkunde Dresden 18:
36 NAUMANN, C. M. (1981): Die zoologischen Forschun 1-12.
gen der letzten zwei Jahrzehnte. Pp. 105-114 in: 52 WITTHOHN, K. & NAUMANN, C. M. (1984): Qualita
RATHJENS, C. (ed.) Neue Forschungen in Afghanistan. tive and quantitative studies on the compounds of the
Leske-Verlag + Budrich GmbH, Opladen. larval defensive secretion of Zygaena trifolii (Esper,
37 NAUMANN, C. M. (1982): Zur Kenntnis der von Man 1783) (Insecta, Lepidoptera, Zygaenidae). Compara
fred Koch beschriebenen Taxa des Zygaena tive Biochemistry and Physiology 79C: 103-106.
c.
purpuralis-Komplexes (Lepidoptera, Zygaenidae). En 53 WITTHOHN, K. & NAUMANN, M. (1984): The dis
tomofauna 3: 225-242. tribution of ß-cyano-L-alanine in cyanogenic Lepidop
38 NAUMANN, C. M. (1982): Zur Kenntnis der von Otto tera. Zeitschrift für Naturforschung 39C: 837-840.
Holik beschriebenen Taxa des Zygaena-purpuralis 54 NAUMANN, C. M. (1984): Zur Kenntnis der von Fran
Komplexes. Entomofauna 3: 371-405. cis Dujardin beschriebenen Taxa des Zygaena
39 NAUMANN, C. M. (1982): Zur Kenntnis der von Hans purpuralis-Komplexes (Lepidoptera, Zygaenidae). En
Burgeff beschriebenen Taxa des Zygaena-purpuralis tomofauna 5: 343-354.
Komplexes (Lepidoptera, Zygaenidae). Entomofauna 55 PRIESNER, E., NAUMANN, C. M. & STERTENBRINK, J.
3: 411-437. (1984): Specificity of synthetic sex-attractants in Zy
40 NAUMANN, c. M. & TARMANN, G. (1983): Ergebnisse gaena moths. Zeitschrift für Naturforschung 39C:
der tschechoslowakisch-iranischen entomologischen 841-844.
Expeditionen nach dem Iran 1973 und 1977 (Lepidop 56 HÄUSER, C. L., NAUMANN, C. M. & TREMEWAN, W.
tera, Zygaenidae). Acta entomologica Musei Nationa G. (1985): On the biology of Parnassius charltonius
lis Pragae 41: 273-276. Gray, 1852 (Lepidoptera: Papilionidae) . Entomolo
41 NAUMANN, C. M. (1982): Zur Kenntnis der von Hugo gist's Gazette 36: 5-13.
Reiss beschriebenen Taxa des Zygaena-purpuralis 57 NAUMANN, C. M. & EDELMANN, A. (1984): Insects of
Komplexes (Lepidoptera, Zygaenidae). Entomofauna Southem Arabia -The life history, ecology and distri
3: 459-498. bution of Reissita simonyi (Rebe!, 1899) (Zygaenidae,
42 NAUMANN, C. M. (1983): Zur Kenntnis der von A. Lepidoptera). Fauna ofSaudi Arabia 6: 473-509.
c.
Costantini und Ubaldo Rocci beschriebenen Taxa des 58 FRANZL, S. & NAUMANN, M. (1985): Cuticular
Zygaena purpuralis- Komplexes (Lepidoptera, Zygae cavities: Storage chambers for cyanoglucoside
nidae ). Entomofauna4: 97-107. containing defensive secretions in larvae of a zygaenid
43 NAUMANN, C. M. (1983): Zur Kenntnis der von Rug moth. Tissue & Cell 17: 267-278.
gero Verity beschriebenen Taxa des Zygaena 59 NAUMANN, c. M. (1985): Zur Ökologie und Biologie
purpuralis-Komplexes (Lepidoptera, Zygaenidae). En von Zygaena (Zygaena) christa Reiss & Schulte, 1967.
tomofauna 4: 141- 155. Nota lepidopterologica 8: 42-50.
c.
44 NAUMANN, C. M. (1983): Zur Kenntnis der von A. 60 NAUMANN, M. & NAUMANN, S. (1985): Zur mor
Przegendza beschriebenen Taxa des Zygaena phologischen Differenzierung asiatischer Populationen
purpuralis-Komplexes (Lepidoptera, Zygaenidae). En des Zygaena-purpuralis-Komplexes (Lepidoptera, Zy
tomofauna 4: 173-180. gaenidae). Entomofauna 6: 265-358.
45 NAUMANN, C. M., RICHTER, G. & WEBER, U. (1983): 61 NAUMANN, C. M. (1985): Phylogenetische Systematik
Spezifität und Variabilität 1m Zygaena-purpuralis und klassisch-typologische Systematik - mit einigen
Komplex (Lepidoptera, Zygaenidae). Theses Zoologi Arrrnerkungen zu stammesgeschichtlichen Fragen bei
cae 2: 1-263 (Verlag J. Cramer, Braunschweig). den Zygaenidae (Lepidoptera). Mitteilungen der
46 NAUMANN, C. M. (1983): Zur Kenntnis der von 0. Münchner entomologischen Gesellschaft 74: 1-35.
Staudinger und H. Rebe! beschriebenen Taxa des Zy 62 FRANZL, S., NAHRSTEDT, A. & NAUMANN, C. M.
gaena-purpuralis-Komplexes (Lepidoptera, Zygaeni (1986): Evidence for site of biosynthesis and transport
dae). Entomofauna 4: 201-208. of the cyanoglucosides linamarin and lotaustralin in
47 NAUMANN, C. M. (1984): Zur Kenntnis der von Leo larvae of Zygaena trifolii Insecta, Lepidoptera). Jour
Sheljuzhko und Otto Holik & Leo Sheljuzhko nal oflnsect Physiology 32: 705-709.
beschriebenen Taxa des Zygaena-purpuralis 63 NAUMANN, c. M. & WITTHOHN, K. (1986): Cyano
Komplexes (Lepidoptera, Zygaenidae). Entomofauna genese bei Zygaeniden (Insecta, Lepidoptera) und
5: 1-19. ihren larvalen Nahrungspflanzen: Co-Evolution oder
48 NAUMANN, C. M. & TREMEWAN, W. G. (1984): Zy einseitige Strategie-Optimierung? Verhandlungen der
gaena anthyllidis Boisduval, [ 1828] (Insecta, Lepidop deutschen zoologischen Gesellschaft 79: 181-182.
c.
tera): Proposed Conservation, Z. N. (S.) 2442. Bulletin 64 BODE, W. & NAUMANN, M. (1987): Structure of a
ofzoological Nomenclature 41: 73-76. newly discovered glandular organ in Neurosymploca
49 NAUMANN, c. M. & TREMEWAN, w. G. (1984): Das larvae (Lepidoptera, Zygaenidae). Zoologische Jahr
Biospecies-Konzept in seiner Anwendung auf die Gat bücher Anatomie 115: 319-329.
tung Zygaena Fabricius, 1775. Spixiana 7: 161-193. 65 NAUMANN, C. M. (1987): Distribution pattems of Zy
50 NAUMANN, C. M., FEIST, R„ RICHTER, G. & WEBER, gaena moths in the Near and Middle East (Insecta,
U. (1984): Verbreitungsatlas der Gattung Zygaena Lepidoptera, Zygaenidae). Beihefte zum Tübinger At-
6 Bonner zoologische Beiträge 53 (2004)
las des Vorderen Orients, Reihe A (Naturwissen (Esper, 1783) (Insecta, Lepidoptera). Verhandlungen
w.
schaften) 28 (KRUPP, F„ SCHNEIDER, & KlNZEL der deutschen zoologischen Gesellschaft 81: 258.
BACH, R., eds., Proceedings of the Symposium on the 80 FRANZL, S., NAUMANN, C. M. & NAHRSTEDT, A.
Fauna and Zoogeography of the Middle East): 200- (1988): Cyanoglucoside storing cuticle of Zygaena
212. larvae (Insecta, Lepidoptera). Morphological and
66 NAUMANN, C. M. & FEIST, R. (1987): The structure cyanoglucoside changes during the moult. Zoomor
and distribution of cyanoglucoside-storing cuticular phology 108: 183-190.
cavities in Pryeria sinica Moore (Lepidoptera, Zygae 81 TREMEWAN, W. G. & NAUMANN, C. M. (1989): The
nidae ). Zoologica Scripta 16: 89-93. nominal taxa described by G. Grum-Grshimailo in the
c.
67 WITTHOHN, K. & NAUMANN, M. (1987): Cyano genus Zygaena Fabricius, 1775 (Lepidoptera: Zygae
genesis - a general phenomenon in the Lepidoptera? nidae). Entomologist's Gazette 40: 109-113, pl. 9, 10.
Journal of chemical Ecology 13: 1789-1809. 82 TREMEWAN, W. G. & NAUMANN, C. M. (1989): Are
68 NAUMANN, C. M. & POVOLNY, D. (1987): Zur Lage vision of Zygaena (Agrumenia) magiana Staudinger,
beziehung und Funktion von W ehrsekretbehältem und 1889 (Lepidoptera: Zygaenidae). Entomologist's Ga
Sinneshaaren im larvalen Integument von Zygaena zette 40: 114-122,pls 11, 12.
(Insecta, Lepidoptera, Zygaenidae). Entomologische 83 NAUMANN, C. M. (1990): Stammesgeschichtliche Re
Abhandlungen und Berichte aus dem Staatlichen Mu konstruktion und ökologische Beziehungen der Organ
seum für Tierkunde Dresden 50: 193-198. ismen - Ansätze für interdisziplinäre Kooperation.
69 BODE, W. & NAUMANN, C. M. (1987. On a pair oflit Verhandlungen der deutschen zoologischen Gesell
tle-known accessory glands in female Zygaena moths schaft 83: 291-301.
(Lepidoptera, Zygaenidae). Zoological Journal of the 84 0CKENFELS, P., NAUMANN, C. M., SCHMITZ, J.,
Linnean Society 92: 27-42. SCHMIDT, W. & FRANCKE, W. (1990): Antennale
70 NAUMANN, C. M. (1987): On the phylogenetic signi Reaktionen von Zygaena trifolii (Zygaenidae, Lepi
ficance of two Miocene zygaenid moths Insecta, doptera) auf Blüteninhaltsstoffe von Knautia arvensis.
Lepidoptera). Palaeontologische Zeitschrift 61: 299- Verhandlungen der deutschen zoologischen Gesell
308. schaft 83: 424.
c.
71 WITTHOHN, K. & NAUMANN, M. (1987): Active 85 ESCH, C. B. & NAUMANN, C. M. (1990): Morpholo
cyanogenesis - a new phenomenon in arthropods. gische und ethologische Differenzierung bei Zygaena
Zeitschrif für Naturforschung 42C: 1319-1322. filipendulae (Linnaeus, 1758) und Z. lonicerae
72 NAUMANN, C. M. (1988): The intemal female genita (Scheven, 1777) (Lepidoptera, Zygaenidae). Verhand
lia of some Zygaenidae (Insecta, Lepidoptera) their lungen der deutschen zoologischen Gesellschaft 83:
morphology and remarks on their phylogenetic signifi 505.
c.
cance. Systematic Entomology 18: 85-99. 86 SCHADE, R. & NAUMANN, M. (1990): Morphologie
73 NAUMANN, C. M. (1988): Functional morphology of und phylogenetische Bedeutung der Sieboldschen
the extemal male and female genitalia in Zygaena Fab Drüse einiger Nymphalidae (Lepidoptera). Verhand
ricius, 1775 (Lepidoptera: Zygaenidae). Entomologica lungen der deutschen zoologischen Gesellschaft 83:
Scandinavica 18: 213-219. 513.
74 NAUMANN, C. M. (1987): Epizygaenella e1ythrosoma 87 HILLE, A., MÜLLER-TAPPE, S. & NAUMANN, C. M.
(Hampson, [1893]), with notes on the taxonomic (1990): Parameter genetischer Variabilität bei
treatment of the genus Epizygaenella Tremewan & isolierten Zygaena-Populationen am nord-westlichen
Povolny, 1968 (Lepidoptera, Zygaenidae). Mitteilun Arealrand (Insecta, Lepidoptera). Verhandlungen der
gen der Münchner entomologischen Gesellschaft 77: deutschen zoologischen Gesellschaft 83: 609-610.
139-147. 88 NAUMANN, C. M., 0CKENFELS, P., FRANCKE, W.,
75 FRANZL, S., NAHRSTEDT, A. & NAUMANN, C. M. SCHMIDT, F. & SCHMITZ, J. (1991): Reactions of Zy
(1987): Chemische Abwehr bei Zygaena-Larven gaena moths (Lepidoptera: Zygaenidae) to volatile
(Lepidoptera, Zygaenidae). Einfluß der Häutungsperi compounds of Knautia arvensis. Entomologia gener
odik auf Morphologie und Inhaltsstoffe kutikularer alis 15: 255-264.
Wehrsekretbehälter.Verhandlungen der deutschen zo 89 NAUMANN, C. M. & TREMEWAN, W. G. (1991): A
ologischen Gesellschaft 80: 208-209. new subspecies of Zygaena (Zygaena) rhadamanthus
76 NAUMANN, C. M. (1988): Revision der Esper'schen (Esper, [1789]) from southern Portugal (Lepidoptera:
Zygaena-Typen (Insecta, Lepidoptera, Zygaenidae). Zygaenidae). Entomologist's Gazette 42: 85-88.
Mitteilungen des internationalen entomologischen 90 NAUMANN, C. M. & TREMEWAN, W. G. (1991): The
Vereins 13: 1-22. type-material and type-locality of Zygaena (Agrume
77 NAUMANN, C. M. (1988): Zur Evolution und adap nia) fausta faustina Ochsenheimer, 1808 (Lepidoptera:
tiven Bedeutung zweier unterschiedlicher Partner Zygaenidae). Entomologist's Gazette 42: 157-160.
c.
findungsstrategien bei Zygaena trifolii (Esper, 1783) 91 RILLE, A. & NAUMANN, M. (1992): Allozyme dif
(Insecta, Lepidoptera). Verhandlungen der deutschen ferentiation in Zygaena transalpina Esper, 1780)
zoologischen Gesellschaft 81: 257-258. (Lepidoptera: Zygaenidae). Pp. 59-87 in: DUTREIX, C.
c.
78 FÄNGER, H. & NAUMANN, M. (1988): Aufbau des et al. (eds) Recent advances in bumet moth research
männlichen Genitaltraktes und Spermatophoren (Lepidoptera, Zygaenidae). Proceedings of the 4'h
bildung bei Zygaena trifolii (Esper, 1783) (Insecta, Symposium on Zygaenidae, Nantes 11-13 September
Lepidoptera). Verhandlungen der deutschen zoolo 1987. Költz, Königstein.
c.
gischen Gesellschaft 81: 249-250. 92 WIPKING, W. & NAUMANN, M. (1992): Diapause
c.
79 PRrNZ, J. & NAUMANN, M. (1988): Optische Pa and related phenomena in zygaenid moths. Pp. 107-
rameter bei der Partnerfindung von Zygaena trifolii 128 in: DUTREIX, C. et al. (eds) Recent advances in
Clas M. NAUMANN: in memorian 7
burnet moth research (Lepidoptera, Zygaenidae). Pro taktzone zwischen Zygaena angelicae und Z. hip
ceedings of the 4'h Symposium on Zygaenidae, Nantes pocrepidis (Insecta, Lepidopt„ Zygaenidae) in Süd
11-13 September 1987. Költz, Königstein. deutschland. Verhandlungen der deutschen zoologi
93 LAMBERT, B. & NAUMANN. C. M. (1992): Zygaena schen Gesellschaft 87.1: 225.
c.
(Zygaena) nevadensis Rambur, 1844 -new for Turkey 106 SPEIDEL, W. & NAUMANN, M. (1995): Phylogenetic
(Zygaenidae, Lepidoptera). Pp. 184-186 in: DUTREIX, aspects in the higher classification of the subfamily
C. et al. (eds) Recent advances in burnet moth research Catocalinae (Lepidoptera, Noctuidae). Beiträge zur
(Lepidoptera, Zygaenidae). Proceedings of the 4'h Entomologie 45: 109- 118.
Symposium on Zygaenidae, Nantes 11-13 September 107 SPEIDEL, W. & NAUMANN, C. M. (1995): Further
1987. Költz, Königstein. morphological characters for a phylogenetic classifica
94 HrLLE, A„ ÜCKENFELS, P„ BöHMER, T. & NAUMANN, tion of the Noctuidae (Lepidoptera). Beiträge zur En
C. M. (1993): Untersuchungen zur Pheromonspezifität tomologie 45: 119-135.
zweier parapatrischer Zygaena Species. Verhandlun 108 WEYER, M. & NAUMANN, C. M. (1996): Möglich
gen der deutschen zoologischen Gesellschaft 86: 244. keiten zur Integration ökologischer Ursache-Wir
95 ÜCKENFELS, P„ NAUMANN, C. M. & FRANCKE, W. kungsbeziehungen in ein naturwissenschaftliches Mu
(1993): Die Rolle von Blüteninhaltsstoffen und ihren seum zur Verbesserung von Umweltaufklärung und
Derivaten im Paarungsverhalten von Zygaena trifolii -erziehung. Umwelt-Bundesamt (Berlin): Texte 4/96:
(Lepidoptera: Zygaenidae). Verhandlungen der deut 423 pp.
schen zoologischen Gesellschaft 86: 266. 109 SPEIDEL, W„ FÄNGER, H. & NAUMANN, C. M. (1996):
96 SOBOTTA, J„ ÜCKENFELS, P. & NAUMANN, C. M. The ultrastructure of the noctuid proboscis (Lepidop
(1993): Die Nutzung von Pyrolizidin-Alkaloiden durch tera: Noctuidae). Zoologischer Anzeiger 234: 307-
Oreina elongata (Coleoptera: Chrysomelidae). Ver 315.
handlungen der deutschen zoologischen Gesellschaft 110 SPEIDEL, W„ FÄNGER, H. & NAUMANN, C. M. (1996):
86: 273. The phylogeny of the Noctuidae (Lepidoptera). Sys
97 NAUMANN, C. M. (1993): Die Russische Entomolo tematic Entomology 21: 219-251.
gische Gesellschaft. Mitteilungen der Arbeitsgemein 111 NAUMANN, C.M. & TREMEWAN, W. G. T. (1996):
schaft Rheinischer Koleopterologen (Bonn) 3: 63-65. Geographical variation in the last instar larvae of Zy
98 FÄNGER, H. & NAUMANN, C. M. (1993): Correlation gaena (Mesemblynus) corsica Boisduval, [1828]
between the mesodermal male genital system and the (Lepidoptera, Zygaenidae). Linnaea belgica 15: 286-
structure of the spermatophore of Zygaena trifolii (Es 288.
per, 1783) (Insecta, Lepidoptera, Zygaenidae ). Acta 112 SPEIDEL, W„ FÄNGER, H. & NAUMANN, C. M. (1997):
Zoologica 74: 239-246. On the. systematic position of Cocytia (Lepidoptera,
99 ÜCKENFELS, P„ PRINZ, J. & NAUMANN, C. M. (1993): Noctuidae). Deutsche entomologische Zeitschrift N. F.
Reactions of Aleioides cf. assimilis (Hymenoptera: 44: 27-31.
Braconidae) to the larval foodplant of its host Zygaena 113 FÄNGER, H. & NAUMANN, C. M. (1998): Genital mor
trifolii (Esper, 1783) (Lepidoptera: Zygaenidae). En phology and copulatory mechanism in Zygaena trifolii
tomologist's Gazette 44: 125- 133. (Esper, 1783) (Insecta, Lepidoptera, Zygaenidae). Acta
100 NAUMANN, C. M., SCHURIAN, K. & WEISS, J.-C. zoologica 79: 9-24.
(1993): Notes on the ecology and early stages of Zy 114 NAUMANN, C. M. (1998): Biodiversity - Is there a sec
gaena (Mesembrynus) lydia Staudinger, 1887 (Lepi ond chance? Pp. 3-11 in: BARTHLOTT, W. & WINIGER,
doptera: Zygaenidae). Entomologist's Gazette 44: M. (eds): Biodiversity. A Challenge for Developmen
225-229, 1 pl. tal Research and Policy. Springer, Berlin, Heidelberg
101 HÄUSER, C. L„ NAUMANN, C. M. & KREUZBERG, A. etc.
c.
V.-A. (1993): Zur taxonomischen und phyloge 115 TREMEWAN, W. G. & NAUMANN, M. (1998): Notes
netischen Bedeutung der Feinstruktur der Eischale der on the biology and ecology of Zygaena (Mesem
Parnassiinae (Lepidoptera, Papilionidae). Zoologische brynus) alpherakyi Sheljuzhko 1936 (Lepidoptera:
Mededeelingen Leiden 67: 239-264. Zygaenidae). Stapfia 55: 107- 112.
102 HEINE, S. & NAUMANN, C. M. (1994): Sexuelle Selek 116 NAUMANN, C. M. (1998): Biogeographie und
tion und die Evolution tageszeitlich variierender Part Stammesgeschichte. Prioritäten für die Erhaltung von
nerfindungsmechanismen bei Zygaena trifolii (Esper, Biodiversität. Pp. 46-49 in: BARTHLOTT, W. & GUT
1783) (Insecta, Lepidoptera). Mitteilungen der MANN, M. (eds) Biodiversitätsforschung in Deutsch
deutschen Gesellschaft für allgemeine und angewandte land. Potentiale und Perspektiven. Europäische
Entomologie 8: 649- 658. Akademie zur Erforschung von Folgen wissen
103 NAUMANN, C. M. & TREMEWAN, W. G. (1994): The schaftlich-technischer Entwicklungen, Graue Reihe no
holotype of Zygaena glasunovi Grum-Grshimailo, 11.
c.
1893 (Lepidoptera: Zygaenidae). Entomologist's Ga 117 RILLE, A. & NAUMANN, M. (1998): Allozyme dif
zette 45: 123- 126. ferentiation in the Zygaena trasnalpina (Esper, 1780)
104 GASSMANN, D„ RILLE, A. & NAUMANN, C. M. (1994): superspecies complex (Lepidoptera, Zygaenidae). II.
Morphometrische Variation in der Kontaktzone zweier Genetic diversity and population taxonomy of the two
Semispezies des Zygaena-transalpina-Komplexes semispecies Zygaena angelicae (Ochsenheimer, 1808)
(Lepidoptera. Zygaenidae). Verhandlungen der deut and Z. hippocrepidis (Hübner, [1799]). Theses zo
schen zoologischen Gesellschaft 87.1: 223. ologicae 30: 45-88.
105 RILLE, A. & NAUMANN, C. M. (1994): Allel 118 GASSMANN, D„ HILLE, A. & NAUMANN, C. M. (1998):
Stufenkline und Multilocus-Kopplungsgleichgewichte Morphological and morphometrical investigations
von Allzymen kennzeichnen eine parapatrische Kon- within a geographical contact zone between Zygaena
8 Bonner zoologische Beiträge 53 (2004)
angelicae Ochsenheimer, 1808 and Zygaena transal Art im Iran (Lepidoptera: Zygaenidae ). Entomolo
pina hippocrepidis (Hübner, [1799]) (Lepidoptera: gische Zeitschrift Frankfurt am Main 112: 233-236.
Zygaenidae) in southern Ge1many. Theses zoologicae 133 FÄNGER, H., ÜWADA, M. & NAUMANN, C. M. (2002):
30: 89-104. The pupa of Eterusia aedea (Clerck, 1759) (Lepidop
119 BöHMER, T., HILLE, A., NAUMANN, C. M. & ÜCK tera, Zygaenidae, Chalcosiinae): morphology and phy
ENFELS, P. ( 1998): On the specificity of female sex logenetic implications. Deutsche entomologische
pheromone compounds in the Zygaena-transalpina Zeitschrift N. F. 49: 261-271.
comp\ex (Zygaenidae, Lepidoptera, Insecta). Theses 134 KLüTSCH, C. F. C., MISOF, B. & NAUMANN, C. M ..
zoologicae 30: 105-124. (2003): Characterization of microsatellite loci for Re
120 MÜLLER, T., ÜCKENFELS, P. & NAUMANN, C. M. issita simonyi (Rebe!, 1899), (Lepidoptera, Zygaeni
(1998): Olfactory discrimination between host and dae) Molecular Ecology Notes 3: 528-531.
non-host-plants in Zygaena trifolii (Esper, 1783) and 135 NAUMANN, C. M .. (2003): Die höchste Zygaene der
Z. transalpina (Esper, 1782) (Lepidoptera, Zygaeni Welt. Entomologische Zeitschrift Frankfurt am Main
dae). Theses zoologicae 30: 125-138. 113: 362-371.
121 ESCH, C. B. & NAUMANN, C. M. (1998): Host 136 ZAGROBELNY, M., BAK, S., RASMUSSEN, A. V., J0R
searching and host-discrimination of a pupal parasitoid GENSEN, B., NAUMANN, C. M., M0LLER, B.L. (2004):
of Zygaena trifolii (Esper, 1783) (Lepidoptera, Zygae Cyanogenic glucosides and plant-insect interactions.
nidae). Theses zoologicae 30: 139-158. Phytochemistry 65 (3): 293-306.
122 FÄNGER, H. & NAUMANN, C. M. (1998): Delayed in 137 KLüTSCH, C. F. C„ GROSSE, W.-R., NASHER, A. K.,
semination in a ditrysian moth, Zygaena trifolii (Esper, MISOF, B. & NAUMANN, C. M. (accepted): The distri
1783) (Insecta, Lepidoptera, Zygaenidae). Theses zo bution of the yellow-lemon tree frog Hyla savignyi
ologicae 30: 159- 164. (Audouin, 1827): updates and extensions of previous
123 KARAMI, M., NAUMANN, C. M. & TREMEWAN, W. G. records (Zoology in the Middle East).
T. (1998): The systematics and distribution of Zygaena 138 PATT,A., MISOF, B., WAGNER, T. & NAUMANN, C. M.
Fabricius, 1775 (Lepidoptera: Zygaenidae) in Iran. (2004): Characterization of microsatellite loci in Am
Iran Journal of Entomology 15(1995): 1-22, 1 [ab phitmetus transversus (Kolbe, 1897) (Coleoptera, Cur
stract in Farsi]. culionidae). Molecular EcologyNotes 4, 2: 188-190.
124 KARAM1, M., NAUMANN, C. M. & TREMEWAN, W. G. 139 SPEIDEL, W. & NAUMANN, C. M. (2005): A survey of
T. (1998): The biology and ecology of Zygaena Fabri family-group names in noctuoid moths (Insecta: Lepi
cius, 1775 (Lepidoptera: Zygaenidae) in the Islamic doptera). Systematics and Biodiversity 2 (2): 191-221.
c.
Republic of Iran. Iran Journal of Entomology 140 SPEIDEL, W. & NAUMANN, M. (2005): The system
15(1995): 23-46, 3 [abstract in Farsi]. atic position of the genus Cetola Walker, 1855 (Lepi
125 EPSTEIN, M., GEERTSEMA, M., NAUMANN, C. M. & doptera: Noctuidae) Entomologische Zeitschrift 115
TARMANN, G. M. (1999): The Zygaenoidea. Pp. (1): 13- 14.
159- 180 in: KRISTENSEN, N. P. (ed.): Handbook of 141 KUZNETZOV, V. I., NAUMANN, C. M., SPEIDEL, W. &
Zoology 4: Arthropoda: Insecta. De Gruyter, Berlin STEKOL'NIKOV, A. A. (2005): The skeleton and muscu
etc. lature of male and female terminalia in Oenosandra
126 NAUMANN, C. M., TREMEWAN, W. G. & TARMANN, boisduvalii Newman, and the phylogenetic position of
G. (1999): The western palaearctic Zygaenidae. 304 the family Oenosandridae (Lepidoptera). SHILAP, Re
pp., Apollo Books, Stenstrup. vista de Lepidopterologia 32: 297-313.
127 FÄNGER, H., YEN, S.-H., NAUMANN, C. M. (1999): PATT, A., MISOF, B., WAGNER, T. & NAUMANN, C.
Externat morphology of the last instar larva of Phauda M.: Isolation and characterization of microsatellite loci
mimica Strand, 1915 (Lepidoptera: Zygaenoidea, Zy in Monolepta vincta Gerstaecker 1871 (Coleoptera,
gaenidae, Phaudinae). Entomologica Scandinavica Chrysomelidae, Galerucinae). Molecular Ecology
30(1998): 429-450. Notes (submitted).
128 NAUMANN, C. M. (2000): Zygaena carniolica (Sco KLüTSCH, C. F. C., MISOF, B. & NAUMANN, C. M.:
poli, 1763) forrnaflava erstmals im Iran nachgewiesen Distribution patterns of the Arabian bumet moth Reis
(Lepidoptera: Zygaenidae ). Entomologische Zeitschrift sita simonyi (Rebe!, 1899) and the wall butterfly
Frankfurt am Main 110: 57. Lasiommata felix (Warnecke, 1929) in Yemen (in
c.
129 FÄNGER, H. & NAUMANN, M. (2001): The mor prep.: Fauna of Saudi Arabia).
phology of the last instar larva of Aglaope infausta NAUMANN, C. M.: New Zygaenidae from South Af
(Lepidoptera: Zygaenidae: Chalcosiinae ). European rica (Lepidoptera, Zygaenidae) (in prep.)
Journal ofEntomology 98: 201-218. NAUMANN, C. M.: Notes on the biology and ecology
130 NAUMANN, C. M. (2001): Biodiversität - gibt es eine of Zygaena huguenini Staudinger, 1888, an endemic
zweite Chance? Pp. 191-195 in WINNACKER, E.-L., species of the Tian Shan mountain ranges (Lepidop
DICHGANS, J., ERKER, G., FRITZSCH, H„ HöLLDOBLER, tera, Zygaenidae) (in prep.).
B., WINTERFELDT, E. & DONNER, W. (eds), Unter NAUMANN, C. M. & TARMANN, G. M.: New light on
jedem Stein liegt ein Diamant. Struktur -Dynamik - the phylogeny of the Zygaenidae: proposing a new
Evolution. S. Hirzel Verlag, Stuttgart-Leipzig. cladogramm for the zygaenid subfamilies.
131 NAUMANN, C. M. (2001): Regenwald und Savanne.
Pp. 31-41 in: TüRKAY, M. (ed.) Leben ist Vielfalt.
POPULAR ARTICLES (EXTRACT)
Kleine Senckenberg-Reihe no. 41.
132 KLiR, J. & NAUMANN, C. M. (2002): Zygaena (Me DOR, R. & NAUMANN, C. M. (1978): Die Kirghisen
semblynus) purpuralis pseudorubincundus subsp. des afghanischen Pamir. Akademische Druck- und
nov.: eine weitere eurosibirisch verbreitete Zygaena- Verlagsanstalt, Graz.
Clas M. NAUMANN: in memorian 9
NAUMANN, C. M. (1986): Zur Tierwelt Afghanistans. NAUMANN, C. M. (2002): Zum Gedenken an Max
Pp. 89-191 in: BUCHERER-DIETSCHI, P. & JENTSCH, Korb (1851-1933) aus Anlass seines 150. Geburt
C. (eds): Afghanistan. Ländermonographie, bearbei stages. Entomologische Zeitschrift Frankfurt am Main
tet von der Arbeitsgemeinschaft Afghanistan. Schrif 112: 12- 19.
tenreihe der Stiftung Bibliotheca Afghanica 4 (Lies NAUMANN, C. M. (2002): Biodiversität - gibt es eine
tal). zweite Chance? Mitteilungen der deutschen Gesell
NAUMANN, C. M. (1991): Apollo und Osiris. Oder: schaft für Chirurgie 1/02: 49-53.
Sind wir noch zu retten? Pp. 42-45 in: Baden NAUMANN, C. M. (2002): Wie lernen Fische das
Württemberg 5/1991. Schwimmen ohne Wasser? Frankfurter Allgemeine
NAUMANN, C. M. (1991): Gift und Farbsignale Zeitung, 21. Juni 2002.
im Überleben der Insekten. Tier und Museum 2: 113- NAUMANN, C. M. (2002): Ohne die Erhaltung der
119. Biodiversität keine erneuerbaren Ressourcen. Pp. 119-
NAUMANN, C. M. (1992): Echolot und Signalfäl 123 in: Bayerische Akademie für Naturschutz und
schung - Krieg der Sterne in der Sommernacht. Pp. 9- Landschaftspflege (ed.) Das Ende der Biodiversität?
13 in: GA!DA, K. G. & PROKOT, S. (eds) Microchirop Grundlagen zum Verständnis der Artenvielfalt und
tera. Falter Verlag, Wien. seiner Bedeutung und der Maßnahmen, dem Arten
NAUMA.NN, C. M. (1995): Terra, quo vadis ? - oder: sterben entgegen zu wirken (5. Franz-Ruttner-Sym
Ziele und Aufgaben des Naturkunde-Museums in un posium). Laufener Seminarbeiträge 2/02.
serer Zeit. Tier und Museum 4: 55-61. NAUMANN, C. M. (2002): Im Spannungsfeld - Um
NAUMANN, C. M. (1996): Biologische Vielfalt aus zo weltforschung. Das Museum Koenig (Schriftenreihe
ologischer Sicht. Veröffentlichungen der Deutsch der Alexander-Koenig-Gesellschaft) No. 4: 4--9.
Chinesischen Gesellschaft e.V., Oktober 1996: 13- 19. NAUMANN, C. M. (2002): BMZ? AA? GTZ? DAAD?
NAUMANN, C. M. (1996): Wo findet die Zukunft der Gender-Seminare statt Damen-Toiletten: Die absurden
Menschheit statt? TenDenZen 5: 27-36. Folgen der deutschen Entwicklungshilfe in Afghani
NAUMANN, C. M. (1998): Unser blauer Planet - Leben stan. Frankfurter Allgemeine Zeitung, 21. November
im Netzwerk. Das Museum Koenig: Blick in die Zu 2002, Nr. 271, S. 40.
c.
kunft. Das Museum Koenig (Schriftenreihe der Alex NAUMANN, M. (2003): Albertus Seba (1665-1736)
ander-Koenig-Gesellschaft) No. 1: 32-35. Rerum Naturalium Thesaurus. Entomologische
NAUMANN, C. M. (1999): Der internationale Code für Zeitschrift Frankfurt am Main 113: 355-359.
zoologische Nomenklatur - 4. Ausgabe. Entomolo
gische Zeitschrift Frankfurt am Main 109: 353-358.
NAUMANN, C. M. (1999): Ras Fartak - 100 Jahre und LIST OF DIPLOMA-AND DOCTORAL THESES
ein bisschen später. Entomologische Zeitschrift Frank SUPERVISED BY C. M. NAUMANN
furt am Main 109: 393-408.
Diploma Theses
NAUMANN, C. M. (1999): In Bonn blüht die Biodiver
sitätsforschung. Das Museum Koenig (Schriftenreihe ALTHOFF, Arnhild (2001) „Fragmentierung von Kalk
der Alexander-Koenig-Gesellschaft) No. 2: 4--9. magerrasen und die Auswirkungen auf die Bestäuber
NAUMANN, C. M. (1999): Das Museum Koenig auf Pflanzen-Interaktionen am Beispiel von Campanula
dem Weg zu neuen Ufern. Kurzfassung einer zehn glomerata" (University of Bonn, jointly with Dr. M.
jährigen Odyssee. Das Museum Koenig (Schriften KRAEMER).
reihe der Alexander-Koenig-Gesellschaft) No. 2: 44- BERGFELD, Thomas ( 1996) „Dynamik von Zooplankton
57. und Jungfischen in Altarmen der Sieg" (University of
NAUMANN, C. M. (2000): Zu Besuch beim Großfür Bonn, jointly with J. FREYHOF and Dr. K. BUSSE).
sten Nikolai Mikhailovich Romanoff. Entomologische B!SCHOFF, Antje (1995) „Verteilungsmuster und Nah
Zeitschrift Frankfurt am Main 110: 12-17. rungsökologie von Jungfischen in Rauschen der Sieg"
NAUMANN, C. M. (2000): Schakuh - ein Traum wird (University ofBonn).
wahr. Entomologische Zeitschrift Frankfurt am Main BISSCHOPINCK, Thorsten (1995) „Die phytophagen Insek
110: 203-211, 12 Abb. ten auf Birke (Betula pendula Roth) unter besonderer
NAUMANN, C. M. 2000. Jede Art führt ein „Nischen Berücksichtigung der Blattminierer im Raum Bonn.
Dasein". Leibniz 3/2000: 18. Ein Standortvergleich" (University ofBonn).
NAUMANN, C. M. (2001): Schattenfreies Fotografieren BöHMER, Thomas (1994) „Untersuchungen zur Abundanz,
von präparierten Insekten. Entomologische Zeitschrift Dispersion und genetischen Nachbarschaftsgröße aus
Frankfurt am Main 111: 187, 188. gewählter Populationen des Zygaena transalpina
NAUMANN, C. M. (2001): Johann Moritz David Superspezies-Komplexes (Insecta, Lep., Zygaenidae"
Herold ( 1815): „Entwickelungsgeschichte der Schmet (University ofBonn, jointly with Dr. A. BILLE).
terlinge". Entomologische Zeitschrift Frankfurt am COUSIN, Markus (1998) „Diversität baumkronenbe
Main 111: 268-276. wohnender Arthropoden eines Tieflandregenwaldes im
NAUMANN, C. M: (2001): Laudatio für Herrn Profes westlichen Uganda (Semliki Forest)" (University of
sor Dr. sc. Bernhard Klausnitzer anlässlich der Verlei Bonn, jointly with Dr. T. WAGNER).
hung der Fabricius-Medaille 2001 der DgaaE. Mit DALBECK, Lutz (1998) „Aspekte der Lebensstrategie von
teilungen der deutschen Gesellschaft für allgemeine Hasel Leuciscus leuciscus (Linnaeus 17 58) und Döbel
und angewandte Entomologie 13: 7-9. Leuciscus cephalus (Linnaeus 1758) in der Sieg"
NAUMANN, C. M. (2001): Ein Mullah mit Gazelle zagt (University ofBonn, jointly with Dr. J. FREYHOF).
in Kabul nie. Frankfurter Allgemeine Zeitung, 21. ESCH, Christoph (1989) „Untersuchungen zur etholo
Dezember 2001. gischen, genetischen und morphologischen Differen-
10 Bonner zoologische Beiträge 53 (2004)
zierung von Zygaena filipendulae (Linnaeus, 1758) tera)" (University of Bonn), jointly with Dr. T. Wag
und Zygaena lonicerae (Scheven, 17 77) (Insecta, ner.
Lepidoptera) sowie zu deren Verbreitung in SW LEVERMANN, Eva-Maria (1998) „Vergleichend öko
Europa" (University ofBielefeld). logische Studien an Panurgus calcaratus (Scopoli
FÄNGER, Harald (1986) „Untersuchungen zum Bau des 1763) und Dasypoda hirtipes (Fabricius 1793) (Hy
weiblichen Genitalsystems und der Spermatophore menoptera, Apidae) in der Wahner Heide" (University
von Zygaena trifolii (Esper, 1783)" (University of of Bonn, jointly with I. B!SCHOFF).
Bielefeld). LINDNER, Michael (1991) „Morphologische und taxono
FELTGEN, Kerstin (1998) „Bionomie der Seidenbiene Col mische Untersuchungen an den Zikaden-Ektoparasiten
letes cunicularius (Linnaeus 1761) (Hymenoptera, Heteropsyche schawerdae (Zerny, 1929)" (University
Aculeata, Apidae )" (University of Bonn, jointly with I. of Bielefeld).
BISCHOFF). MAURER, Golo (2001) "Nestling begging behaviour and
FIELENBACH, Jörg (1996) „Zeitliche und räumliche the impact of parental alarm-calls in white-browed
Verteilungsmuster der Nasen Chondrostoma nasus scrubwrens, Sericornis frontalis (Vigors & Horsfield)"
(L.) in der Sieg" (University of Bonn, jointly with J. (University ofBonn).
FREYHOF and Dr. K. BUSSE). MEUSEMANN, Karin (2001) „Karyologische Unter-Su
FREUND, Wolfram (2000) "Revision of Bonesioides chungen zum Agrodiaetus-Komplex (Lycaenidae)"
Laboissiere, 1925 (Coleoptera: Chrysomelidae, (University ofBonn).
Galerucinae)" (University ofBonn, jointly with Dr. T. MIDDELHAUVE, Jens (2000) "Revision of Afrocrania
WAGNER). Hincks, 1949 (Coleoptera: Chrysomelidae, Galeru
FREY, Susanne (1989) „Altersbedingte Veränderungen in cinae)" (University of Bonn, jointly with Dr. T. WAG
der Feinstruktur des weiblichen Pheromonsystems von NER).
Zygaena trifolii (Esper, 1783)" (University of Biele NEEBE, Bettina (1992) „Der Einfluß von Störreizen auf die
feld). Herzschlagrate brütender Küstenseeschwalben (Sterna
FREYHOF, Jörg (1992) „Zeitliche und räumliche Verteilung paradisea)" (University ofBonn).
von Jungfischen in der Fließstrecke der Sieg" (Univer NEMITZ, Armin (1994) „Untersuchung zum Jungfisch
sity ofBonn,jointly with Dr. K. BUSSE). aufkommen in einigen Laichschongebieten der Sieg"
GRUNDMANN, Bernd (1984) „Vergleichende, ökologisch (University of Bonn, jointly with Dr. K. BUSSE).
faunistische Untersuchungen zur terricolen Käferfauna OCKENFELS, Peter (1989) „Untersuchungen zur Attraktion
der Pflanzengesellschaften eines regenerierenden Ho der Imagines von Zygaena trifoli an Dipsacaceen
chmoores" (University of Bielefeld). Blüten" (University ofBielefeld).
HERDER, Fabian (2000) „Zoogeographie und Ökologie OPPERMANN, Markus (1994) „Vergleichende Untersu
zentralvietnamesischer Süßwasserfische" (University chungen zum abdominalen Tracheen-System der Ark
of Bonn, jointly with Dr. K. BUSSE and Dr. J. FREY tiidae (Lepidoptera)" (University ofBielefeld).
HOF). PFEIFER, Carmen (1999) „Blumenvögel und Vogelblu
HEUSER, 0. ( 1994) „Zur chemosensorischen Funktion der men - wechselseitige Anpassungen und Abhängig
Pedipalpen bei der juvenilen Jagdspinne Cuppiennius keiten" (University of Bonn, jointly with Dr. M.
salei (Aranei, Ctenidae)" (Universities of Wien and KRAEMER).
Bonn). PIEPER, Benno (1991) „Untersuchungen zum Reproduk
RILLE, Axel (1986) „Morphologische und enzymelek tionserfolg der Männchen bei W eibchenpaarungen
trophoretische Merkmalsdifferenzierung im Zygaena zweiten Ranges im Paarungssystem von Zygaena trifo
transalpina-Komplex (Lepidoptera, Insecta)" (Univer lii (Esper, 1783) (Lepidoptera, Zygaenidae)" (Univer
sity ofBielefeld). sity ofBielefeld).
HÖLLING, Doris (1992) „Vergleichende Untersuchungen PRINZ, Jochen (1988) „Untersuchungen zum Partner
von Carabiden- und Curculioniden-Zönosen in gegat findungsverhalten von Zygaena trifolii (Esper, 1783) -
terten und ungegatterten Rotbuchenwäldern im Ams ein Beitrag zur Reproduktionsbiologie der Zygaeniden
berger Wald (Sauerland)" (University of Bonn, jointly (Insecta, Lepidoptera)" (University ofBielefeld).
with Prof.Dr. G. KNEnz).. PöRSCHMANN, Melanie (1986) „Morphologische und habi
HÖLLING, Michael (1990) „Uber die Eiablage, den Repro tuelle Parameter im larvalen Entwicklungszyklus von
duktionserfolg und die Steuerung des des Lockverhal Zygaena trifolii (Esper, 1783)" (University of Biele
tens von Zygaena trifolii (Esper, 1783)" (University of feld).
Bielefeld). RAMMERT, Uwe (1985) „Untersuchungen zur Wirksamkeit
KLUG, Andrea (1993) „Bestäubungsökologie neotropischer des Wehrsekretes und des aposematischen Zeich
Ericaceen" (University of Bonn, jointly with PD Dr. nungsmusters von Zygaena trifolii (Esper, 1783) auf
K.-L. SCHUCHMANN). unerfahrene Stare (Sturnus vulgaris, L.)" (University
KNEIP, Carsten (1999) „Vergleich der baumkronenbewoh ofBielefeld).
nenden Arthropodenfauna in Montanwäldern Ostafri SCHADE, Renate (1991) „Morphologie und phylogene
kas" (University of Bonn, jointly with Dr. T. W AG tische Bedeutung der Siebold'schen Drüse der Nym
NER). phalidae (Lepidoptera)" (University ofBielefeld).
KOHL, Michael (2001) „Die Arthropoden- und speziell die SCHIDELKO, Kathrin (2002) „Neozoen im Großraum Bonn
Käferfauna von Teclea (Rutaceae) in verschiedenen unter besonderer Berücksichtigung der Entenvögel"
Wäldern Ostafrikas" (University of Bonn, jointly with (University of Bonn, jointly with Dr. R. VAN DEN EL
Dr. T. WAGNER). ZEN).
KURTSCHEID, Agnes (2000) „Revision von Candeza SCHMIDT-LOSKE, Katharina (1992) „Raumeinbindung und
Chapuis, 1879 (Galerucinae, Chrysomelidae, Coleop- Biotopnutzung tagfliegender Großschmetterlinge am
