Table Of ContentCATALOG
OF
FOSSIL SPORES AND POLLEN
Volume 42
Cretaceous Spores and Pollen
from the U.S.A. and the U.S.S.R.
PREPARED BY
H.T. AMES and W. SPACKMAN
Coal Research Section
THE PENNSYLVANIA STATE UNIVERSITY
Published at
University Park, Pennsylvania
1981
Copyright ©
by The Pennsylvania State University (1981)
Copying of material
in the Catalog of Fossil Spores and Pollen
for com mercial exploitation is prohibited.
CATALOG OF FOSSIL SPORES AND POLLEN
Editors
H.T. Ames W. Spackman
Editorial Committee
W.G. Chaloner, Bedford College, University of London, London, England
A.T. Cross, Michigan State University, East Lansing, Michigan
J. Jansonius, Imperial Oil Co., Calgary, Alberta
R.M. Kosanke, U.S. Geological Survey, Denver, Colorado
G.O.W. Kremp, Palynodata, Inc:, Tucson, Arizona
E.A. Stanley, Indiana University of Pennsylvania, Indiana,
Pennsylvania
Business Manager
Karen Copenhaver
TABLE OF CONTENTS
Compiler's Preface. ii
Brenner, G.J., 1963
The spores and pollen of the Potomac Group of Maryland • • •• 1
Khlonova, A.F., 1961
Spores and pollen of the upper half of the Upper
Cretaceous of the eastern part of the West-Siberian
lowland . . . . • . . . . • . 125
Appendix
Part I
Generic Emendations 209
Part II
List of New Combinations . 211
Reference 217
Index 220
List of Nomenclatural Comments
DisyUabipoUis 150
LongaevipoUis . 169
MUZtinodisporites 172
OccuZtospoPites 180
Po Zypodiisporites 195
Seductisporites 200
CicatricosispoPites araZica 215
42:i
COMPILER'S PREFACE
Volume 42 contains systematic information originally published in
Brenner, 1963, and Khlonova (Chlonova), 1961. Brenner's investigation
constitutes one of the earliest, if not the earliest, major palynological
study of the Lower Cretaceous from the Atlantic Coastal Plain of North
America. His samples included surface and subsurface sediments from
the Patuxent, Arundel, and Patapsco formations (Potomac Group) from
localities in Delaware, Maryland and Virginia. Khlonova's study was
based upon samples from 8 localities in the eastern part of the western
Siberian lowland along the Bax, Cbim, Kemb and Enuceu rivers. These
samples were dated as Late and Latest Cretaceous.
Systematics recorded in this Volume from Brenner, 1963, and
Khlonova, 1961, include 15 new genera, 112 new species, 1 emendation
and 14 new combinations. The emendation and new combinations are
presented as Part I and Part II of the Appendix. New taxa from Brenner
include one genus (Decussosporites) and 65 species, while new taxa from
Khlonova include 14 genera and 47 species. The following names were
proposed by Khlonova for her new genera. Accuratipollis, Circumflexi
pollis, Compactipollis, Deplexipollis, Disyllabipollis, Expressipollis,
Fibulapollis, MUltinodisporites, Occultosporites, Ocellipollis,
Orbiculapollis, Seductisporites, Translucentipollis and Tricerapollis.
Brenner's emendation of Exesipollenites Balme 1957 is reproduced in Part
I of the Appendix, while the 5 and 9 new combinations proposed by
Brenner and Khlonova, respectively are identified along with their
basionyms in Part II of the Appendix.
42:ii
Brenner's new species were assigned to 40 genera while Khlonova
assigned hers to 28 different genera. About one third of the genera.
to which KhlonoV'a assigned her new or previously described species.
refer to extant plant taxa.
To locate any significant specimen recorded in his 1963 publi
cation. Brenner provided the following method to convert his published
coordinates for use with other mechanical stages. As a base from
which to determine a conversion factor he inscribed an "X" with known
coordinates. on each of three blank slides. which he calls "Key slides".
(In all cases Brenner recorded the horizontal component for any of his
coordinates first.) The difference between the "known" coordinates
on any of his "key slides" and those observed for that "X" position
on the vernier readings of an alternative stage established a conversion
factor between the two stages. Proper "directional" application of the
horizontal and vertical components of this conversion factor to the
published coordinates for any specimen recorded in Brenner. 1963.
will convert his coordinates to vernier readings that will properly
locate that specimen for observation on the alternative stage.
The coordinates of each X is as follows:
Key slide A. coordinates 28.7 x 119.4
Key slide B. coordinates 28.5 x 119.3
Key slide C. coordinates 32.2 x 120.1
Key slides are retained by the author in his personal collection of
type slides.
With regard to Klonova's identification of holotypes repetition
of numerical designations purporting to identify holotypes in Khlonova.
1961. and in the CFSP references. indicate that these designations of
themselves. cannot actually identify an individual specimen. However.
42 :iii
they apparently do identify the slide which carries the type (personal
communications from E. Stanley to H. T. Ames.) In addition to the
numerical designations, Khlonova also provided illustrative
identification of her holotypes, in that figures illustrating the
holotype are so noted in Russian by the word ronoTHn in the accompany
ing captions. CFSP cites these illustrations by plate and figure
as part of its identification of the holotype.
Khlonova's treatment of her proposed new monotypic genera in
1961 would not have provided valid publication of their names when
published, per a strict interpretation of the requirements of Article
PB6 1956 ICBN. This Article reads: "The name of a monotypic genus
of fossil plants published after 1 Jan. 1953, must be accompanied by
a description of the genus indicating its differences from other
genera." Following deletion in 1959 of the Appendix entitled:
"Special provision concerning fossil plants", the essence of Article
PB6 was combined with Article 42 of subsequent Codes until publication
of the 1978 edition of the ICBN. In this edition the special and
restrictive limitation imposed for the names of monotypic genera of
fossil plants was also eliminated. As Khlonova's treatment is now
in compliance with requirements pertaining to the names of monotypic
new genera of both fossil and extant plants, her names must be recog
nized as validly published.
The editors acknowledge, with gratitude, permission from the
authors and publishers to reproduce systematic material from their
publications. We also wish to express our appreciation to G.J.
Brenner and A.F. Khlonova for their aid and helpful suggestions
which contributed to the preparation of Volume 42.
42:iv
Abietineaepollenites1minimus Brenner 1963
2a 2b 2c
FiJ:ure 2 . ..t biclil/cllrp"lIwilcs m;,,;I/III.<, n. sp.
lIolotyp<" slid .. LI·-lL\. l'CIordinates 21lA· 11.1.1. lall'ral vil·w. 2a. low foclIs. lh, mediulll
focus, sh"winJ: the ck texine absent on the dislal surface hl'lwl'l'n I he .hladders. 2e, hiJ:h
focus shows Ih .. lincl), rUJ:ulate nature of the proximal cap.
Description:
Bisaccate pollen; central body broadly oval to circular in lateral
view; bladders distally disposed with the proxirnal attachment proximal from
the equator; exine of the proximal cap is rugulate, ca. 1.5 Ji thick; end exine
smooth, exposed on the distal surface between the bladders,. ca. 0.5 p. thick;
bladders small in relation to central1Jody, attached at their maximum height.
Dimensions.' (5 specimens)
Body width. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 22(25)33 p.
Body height. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 18(21)24 p.
Bladder length ..... , ............................... 6-7 p.
Total length of grain. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34(36)37 p.
Comments: Abietineaepollenites minimus can be distinguished from other
bisaccate pollen by its small size, small sacs in relation to the body, and rugulate
proximal cap.
Occurrence.' Rare; sample frequency infrequent, has been found only in
Subzone B. 2
Holotype: Slide L1-41A, coordinates 28.4-113.1; plate 26, fig. 2.
CATALOG OF FOSSIL SPORES AND POLLEN 42:1
Author: Brenner, G.J., 1963, p. 77, pI. 26, fig. 2
Reference: The spores and pollen of the Potomac Group of Maryland: Dept.
Geol., Mines and Water Resources, Bull. 27, 215 pp., 43 pIs.,
7 text figs., 7 tabs.
Type locality: Maryland, Prince Georges Co., ca. 15 mi. E of Washington, D.C.,
Belair Estate (U.S. 411.3 N, 871.9 E), Sydnor
Army~oord.
Pump &W ell Co., Probe Hole No.1, depth 447.0-448.5 ft.
Type horizon: Cretaceous, Potomac Group, Patapsco Fm.
Preparation: All samples collected from unconsolidated sediments which were
easily disaggregated by crushing in a mortar with distilled H20.
Organics separated by flotation in ZnC12 sol. (2.0 sp. gr.).
Organic float treated HF 23 min., oxidized 20 min. in 5.25%
NaOCl sol. (C10rox), treated with 10% 3 min., stained in
~~40H
a dilute tincture of safranin Y 10-20 min. Diaphane mounts.
Repository: Not indicated
Holotype: PI. 26, fig. 2, Slide Ll-4IA, coord. 28.4xl13.l2
1 [Brenner assigned this species to '~bietineaepoZZenites Potonie
1951". The generic name was not validated, however, until treated
in Potonie, 1958, p. 61. It should be cited as AbietineaepoZZenites
Potonie 1951 ex Potonie 1958.]
[See Compiler's Preface for explanation of location coordinates
2
of holotypes.]
CATALOG OF FOSSIL SPORES AND POLLEN 42:2
Species n.me: AZsophilidites pannuaeus Brenner 1963
Sa Sb 6
Figures 5, 6. Alsopililidites pannllcells, n. sp.
Fig. S. Holotype, slide St. lOB, coordinates 21.2-125.5. Sa, high focus on the proximal
surface. Sh, low focus on the distal surface. The finely wrinkled surface is characteristic
of this species.
Fig. 6. Slidc 1.2-58/\, coordinates 38.2-108.2, mcdiulll focus, :1 slightly larger specimen
with a more strongly developed margo.
Description:
Trilete spore; laesurae lI'eaching the margin, commissure bord
ered by a dark lip 1.4-2.4 Il wide and tapering slightly toward the corners;
outline in polar view triangular with straight to slightly convex sides and acute
corners; the equatorial outline in the interradial areas arc simlOu,; or indented.
Exine smooth ca. 1.5 JL thick; surfat:e finely wrinkled, the wrinkles showing
no preferred orientation.
DiIllCltsicJl/s: l\Iaximllm diameter 11 (U)34 JL (13 specimens).
Commel/ts: Spores of similar Iliorpholot;r an: found in members of the
C)'utlzeaccae, Glciclleuiaccae, and Polypodiaceac.
Occurrel/ce: Rare to ocra:<ional; "ample frequency high. This "pecic,; is found
in Lot h Zones I :lli'd 11, but occurs more often, and in slightly greater specimen
fn:quellcies in Zone I.
lIolotypc: Slide St. 1013, coordinates 21.2-125.5t plate 12, fig. 5.
CATALOG OF FOSSIL SPORES AND POLLEN 42:3
