Table Of ContentBull.Br.arachnol.Soc.(1999)11(5),161–184 161
Bibliographic check list of the Hungarian spider Time period: The check list is based on the last
fauna published check list from Hungary by Chyzer &
Kulczyn´ski(1918a,b).Thosepapersgaveacomprehen-
Ferenc Samu sive and critical list of all previous arachnological
PlantProtectionInstitute, records, since the pioneering work by Herman (1876,
HungarianAcademyofSciences, 1878, 1879). For records after 1900 the authors have
P.O.Box102,Budapest,H-1525Hungary* extensively searched the literature between 1900 and
1998.
and
Sources: Concerning the Chyzer & Kulczyn´ski
(1918a,b) papers, it has to be mentioned that they were
Csaba Szinetár
completed in 1896 and 1900 respectively, but remained
BerzsenyiCollege,DepartmentofZoology,
unpublished until 1918. These papers synthesise all
4KárolyiGáspártér,
records from the better known three-volume Araneae
Szombathely,H-9700Hungary
Hungariae (Chyzer & Kulczyn´ski, 1891, 1894, 1897).
Thecoverageofliteratureafter1900isalmostcomplete.
Summary
Szinetár & Samu (1995) gave a comprehensive list of
The first check list of the Hungarian spider fauna is Hungarian arachnological literature. In the present list,
presentedsincethe1918checklistbyChyzerandKulczyn´- to our knowledge we have included all records from
ski.Allpre-1900recordsareincludedfromtheChyzerand Balogh, Loksa and other contemporary arachnologists.
Kulczyn´skilist.Fortheperiod1900–1996theliteraturehas
Kolosváry was an extremely prolific author, and we
been extensively searched, and records are included from
have been able to examine c. 95% of his papers, includ-
theoriginalsources.Recordsarelistedonlyfromlocations
within the present borders of Hungary. Altogether the ing all important faunistical reports. The reason why
presentlistisbasedon108literatureitems,inwhichthere many papers which might be well known in the arach-
were6010recordsfor725spiderspeciesfromthecountry. nological literature, but to which no reference can be
foundhere,issimplythatthesepaperscontainedrecords
Introduction only from outside Hungary.
The preservation of global and local biodiversity
requires knowledge about patterns of diversity. Fauna Data validation procedures
lists of spiders have traditionally served this purpose.
Species identity: The taxonomic names mainly follow
Suchlistscanbeestablishedatdifferentscales,forwhole
the nomenclature in Platnick (1997). Original names
countries (Merrett et al., 1985; Pesarini, 1994; Buchar
in the sources were often synonyms. These were
etal.,1995;Mikhailov,1996),orsmallerregions,suchas
checked and if possible cleared using the catalogues of
a national park (Coddington et al., 1996). Fauna lists
Bonnet (1945, 1955–1959, 1961), Roewer (1942, 1955),
areinventories,whichnotonlyhelptoestablishpatterns
Brignoli (1983) and Platnick (1989, 1993, 1997),
of biodiversity, but also serve as raw material for more
but other sources, such as Prószyn´ski’s computer
detailed ecological studies. Their electronic publication
catalogue (1993) and original species descriptions, were
viatheInternet(e.g.Langemark;Workinggrouparach-
also used.
nology, 1996; Vanuytven, 1998) or in other database
In cases where one or more sibling species have been
formats (Hanggi et al., 1995) makes their availability
described based on one original species, or where sub-
wider and easier.
species have been elevated to species rank, the original
Arachnological studies in Hungary have a long tra-
speciesnameislistedeithersensustrictoorsensulato.In
dition. Listing the spider fauna was started by Ottó
caseswhereweknowthatthespecieshasbeenidentified
Herman (1876–1879) and was continued by Cornél
according to the narrower definition, we list it as sensu
ChyzerandLászlóKulczyn´ski(1891,1894,1897).Based
stricto. We have indicated sensu lato in cases where
on their three-volume seminal work, the first and until
either the records originated before the sibling species
now the only fauna list of Hungarian spiders was
were described, or in cases where it was impossible to
publishedin1918byChyzer&Kulczyn´ski(1918a,b).In
establish whether the species names have been used in
the present list we set out to revise this list with
the wider or in the narrower sense. For species given as
published spider data originating after 1900, in a
sensu lato, the names of sibling species included under
format where all known references are given for each
thatnamearelisted.Insomespeciesrecordsareknown
species included in the list. We hope that an electronic
in both senses, and these are shown as separate entries,
publication of our results will follow in the near future.
but counted only as one species.
We have classified species as invalid if their identity
Material and methods could not be established, or if their validity was ques-
tionable (nomen dubium), or in one case because it was
Scope of data included in check list
stated to be a nomen nudum in the literature (Loksa,
Area: Only records from locations within the present 1972).
borders of Hungary are included. Location validity: Each record in the literature was
checked,astowhethertheoriginallocationiswithinthe
*Addressforcorrespondence. present borders of Hungary, using old locality lists and
162 ChecklistofHungarianspiders
Speciesname Reference Presentdesignation Reference
fordesignation
BalluslendliKolosváry,1934 Kolosváry,1934a Nomendubium presentpaper
CheiracanthiumcuniculumHerman,1879 Chyzer&Kulczyn´ski,1918b Nomendubium Chyzer&Kulczyn´ski,1918b
CyclosastrandiKolosváry,1934 Kolosváry,1934a Nomendubium presentpaper
DrassodesstriatusL.Koch,1866 Kolosváry,1933b Nomendubium presentpaper
EnoplognathahungaricaKolosváry,1934 Kolosváry,1934a,1935b,d Nomendubium presentpaper
LepthyphantesanguliformisLoksa Loksa,1966a Nomendubium presentpaper
LophommarufipesBösenberg,1902 Kolosváry,1928b,c,1933b Nomendubium Braun,1982
LycosaconicaKolosváry,1928 Kolosváry,1928b Nomennudum Loksa,1972
PoecilochroahungaricaKolosváry,1934 Kolosváry1932b,1934a,1935b Nomendubium presentpaper
TheridionsilvaticumKoch Loksa,1966a Nomendubium presentpaper
Table1: Listofinvalidspecies.
maps. A complete locality list in the second volume of dated, given the unknown location of type material and
Chyzer & Kulczyn´ski (1897) was especially useful. the poor quality of the drawings.
Record validity: There was no exact way to check the In the list of invalid species (Table 1) the original
validity of the records. We declare the present paper to species name is given, as it occurred in the source
be a literature survey. Accordingly no specimens from literature. It sometimes lacked the year of description.
collectionswerechecked.However,werealisethatthere
was a wide range of different quality data. In the
Probably invalid records (not in check list)
overwhelming majority of cases we had no reason to
doubt the precision and validity of the records. Special Based on the criteria described in the data validation
care was taken of species found in one or very few section,therecordspresentedinTable2areregardedas
records. Many of these were supported by our own invalid.
unpublished data. Others were less certain, but we had
no grounds to exclude them. A separate list of these
uncertain species is given below. Species that deserve Possibly uncertain records (in check list)
special attention because they were originally described
Wegivealistof‘‘somewhatuncertainspecies’’,which
from Hungary are also listed separately.
we feel might be problematic. Species were included in
A record was classified as invalid if any of the
thislistonthebasisofcommentsonpreviousversionsof
following circumstances were met: (i) Occurrence is
the list by T. Blick, K. Thaler and P. Merrett; known
considered highly unlikely on the basis of known biol-
distribution made occurrence questionable, but still
ogy, habitat preference and distribution. These species
possibletosomedegree(cf.listofinvalidrecords).These
usually were restricted in known occurrence to high
species are listed here to indicate that some caution
mountains, seashores, or were Mediterranean species.
should be paid when accepting the record as valid for
They have not been mentioned from any habitat prob-
Hungary.
able in Hungary in the check lists of neighbouring
countries (Drenski, 1936; Fuhn & Oltean, 1970; Nikolic
SpeciesName Reference
& Polenec, 1981; Weiss, 1987; Buchar, 1992; Pesarini,
1994; Buchar et al., 1995; Platen et al., 1995; Kropf & Centromerussellarius(Simon,1884) Loksa,1971
Centromerussilvicola Kolosváry,1943
Horak, 1996; Mikhailov, 1996). (ii) Records examined
(Kulczyn´ski,1887)
byLoksa(1969a,1972)anddeclaredtobenotvalid.We
Drassodesheeri(Pavesi,1873) Kolosváry,1930
are not sure, but it is likely, that this declaration was Drassodeshypocrita(Simon,1878) Kolosváry,1933b
basedontheexaminationofspecimens.(iii)Revisionof Helophorainsignis(Blackwall,1841) Kolosváry,1935a
theidentificationmadebytheauthorofthegivenrecord Lepthyphantesistrianus Kolosváry,1934c
Kulczyn´ski,1914
in a later paper, or unpublished self-revision in the case
LepthyphantesliguricusSimon,1929 Loksa,1966a
of Samu and Szinetár.
MicariasilesiacaL.Koch,1875 Kolosváry,1930
Porrhommaerrans(Blackwall,1841) Balogh&Loksa,1953
Porrhommarosenhaueri Loksa,1961b
Results of species and record validation
(L.Koch,1872)
Silometopuscurtus(Simon,1881) Loksa,1991b
Invalid species (not included in check list)
Sitticusrupicola(C.L.Koch,1837) Chyzer&Kulczyn´ski,1918a
We classified most of the species described by TrachelasmaculatusThorell,1875 Kolosváry,1932b,1935b
TrichoncushackmaniMillidge,1956 Kerekes,1988
Kolosváry from Hungary as nomina dubia (many other
Zelotesclivicola(L.Koch,1870) Kolosváry,1932b,1935b,1943
species were described by him from outside the area of
Hungary,andthusfalloutsideourscope).Thesespecies
have seldom if ever been mentioned after the original
Notes on some valid records
description, and in many cases only one sex was found
byKolosváry.Consideringtheoriginaldescriptions,itis Porrhommaspeciesareprobablythemostproblematic
hardly likely that these Kolosváry species will be vali- group in the list. Miller & Kratochvil (1940) examined
F.Samu&C.Szinetár 163
Species Reference Reason Speciesin
foruncertainty checklist
Argiopelobata(Pallas,1772) Balogh,1935 self-revision yes
ClubionareclusaO.P.-Cambridge,1863 Szinetár,1992e self-revision yes
Dasumiaamoena(Kulczyn´ski,1897) Kolosváry,1932b,1935b Loksa,1969a no
DictynamajorMenge,1869 Kolosváry,1932b Loksa,1969a no
Evarchajucunda(Lucas,1846) Kolosváry,1933b,1934b occurrenceunlikely no
Gnaphosamontana(L.Koch,1866) Kolosváry,1935c occurrenceunlikely no
LarinialineataLucas,1846 Szinetár,1993 self-revision no
Marilyniabicolor(Simon,1870) Kolosváry,1928c Loksa,1969a no
Nemesiapannonica(Herman,1879) Chyzer&Kulczyn´ski,1918a self-revision yes
Neosconasubfusca(C.L.Koch,1837) Chyzer&Kulczyn´ski,1918a Loksa,1969a no
Nigmapuella(Simon,1870) Kolosváry,1934b Loksa,1969a no
Nucteneasilvicultrix(C.L.Koch,1844) Kolosváry,1935d Loksa,1969a no
Pardosaalbatula(Roewer,1951) Kolosváry,1933b occurrenceunlikely no
Pardosablanda(C.L.Koch,1834) Kolosváry,1928b,1933b Loksa,1972 no
Pardosamonticola(Clerck,1757) Samu&Lövei,1995 self-revision yes
Pardosanigriceps(Thorell,1856) Kolosváry,1933b Loksa,1972 no
PardosapurbeckensisF.O.P.-Cambridge,1895 Loksa,1988 occurrenceunlikely no
PhilodromusglaucinusSimon,1870 Szinetár,1992d self-revision no
Philodromuspoecilus(Thorell,1872) Szinetár,1992c,d self-revision yes
TegenariapictaSimon,1870 Kolosváry,1933d,1934c Loksa,1969a no
TetragnathapineaSeeley,1928 Kolosváry,1935d occurrenceunlikely no
TheridionpetraeumL.Koch,1872 Kerekes,1988 occurrenceunlikely no
Table2: Listofprobablyinvalidrecords.Aspecieswasincludedinthechecklistiftherewereothervalidrecords,butwasexcludedifonlythe
revisedrecordwasfoundforthegivenspecies.
several Porrhomma specimens originating from Xysticus embriki Kolosváry, 1935c
Hungary. According to their results all Porrhomma Cryptodrassus hungaricus (Balogh, 1935)—this species
rosenhaueri records before that study were in fact was described as Zelotes hungaricus Balogh and trans-
Porrhommaprofundum.TherecordofPorrhommaerrans ferred to the genus Cryptodrassus by Weiss et al., 1998.
by Kolosváry (1928a) is also P. profundum (Miller &
Kratochvil, 1940). The subspecies Porrhomma rosen-
List of species
haueri hungaricum Loksa, 1970 is a junior synonym of
P. profundum (Thaler & Plachter, 1983). Thus only
one record each of P. errans and P. rosenhaueri
Family ATYPIDAE
remained which we regard as valid (but draw attention Atypus affinis Eichwald, 1830
to them, by listing them above); both records were
Balogh, 1938c; Chyzer & Kulczyn´ski, 1918a; Kisbenedek, 1991;
identified by Loksa. Porrhomma thorelli mentioned by Kolosváry,1934b;Loksa,1966a,1969a,1971,1988,1991b.
Chyzer & Kulczyn´ski (1918a)=P. convexum. P. errans Atypus piceus (Sulzer, 1776)
sensu Chyzer & Kulczyn´ski (1894)=P.proserpina Balogh,1938b;Balogh&Loksa,1953;Loksa,1966a,1969a,1981b,
(Miller & Kratochvil, 1940). Since Porrhomma proser- 1991a;Szinetár,1992b.
pina is regarded as a junior synonym of P. convexum
(Holm, 1944), all records of P. proserpina are listed
Family NEMESIIDAE
under P. convexum. However, we noted that the
Nemesia pannonica Herman, 1879
reported habitats were systematically different between
Balogh, 1938c; Kisbenedek, 1991; Loksa, 1958, 1966a,b, 1969a,
the two nominal species; P. proserpina having always 1971,1984,1988;Samuetal.,1996.
beenreportedfromcavesandP.convexumfromvarious
non-cave habitats. Only a future examination of
Family SCYTODIDAE
the Hungarian cave Porrhomma spp. can resolve this
Scytodes thoracica (Latreille, 1802)
problem.
Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928b; Loksa, 1958,
1969a;Pillich,1914;Szinetár,1992e.
NewspeciesdescribedfromHungaryafter1900(incheck
list)
Family PHOLCIDAE
The following new species, described from Hungary Hoplopholcus forskali (Thorell, 1871)
after 1900, are regarded as valid: Balogh & Loksa, 1946; Chyzer & Kulczyn´ski, 1918a; Kolosváry,
Altella hungarica Loksa, 1981b 1928b,1930;Loksa,1969a,1984;Szinetár,1992e.
Pholcus opilionoides (Schrank, 1781)
Altella orientalis Balogh, 1935
Balogh, 1933, 1935, 1938a; Balogh & Loksa, 1953; Chyzer &
Brommella falcigera (Balogh, 1935)
Kulczyn´ski, 1918a; Kisbenedek, 1991; Kolosváry, 1928c, 1930,
Metopobactrus deserticola Loksa, 1981b
1933d, 1934c; Loksa, 1962, 1966a, 1969a, 1971, 1977, 1984, 1988,
Sintula spiniger (Balogh, 1935) 1991a;Pillich,1914;Szinetár,1991,1992e.
164 ChecklistofHungarianspiders
Pholcus phalangioides (Fuesslin, 1775) Ero tuberculata (De Geer, 1778)
Balogh & Loksa, 1946; Chyzer & Kulczyn´ski, 1918a; Kolosváry, Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Loksa, 1977,
1928b,1930;Loksa,1966a,1969a,1984;Szinetár,1992e. 1984;Szinetár,1992e.
Psilochorus simoni (Berland, 1911) Mimetus laevigatus (Keyserling, 1863)
Szinetár,1992e. Balogh,1933;Chyzer&Kulczyn´ski,1918a.
Family SEGESTRIIDAE
Family ERESIDAE
Segestria bavarica C. L. Koch, 1843
Eresus cinnaberinus (Olivier, 1789)
Balogh, 1933; Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1930;
Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Kerekes, 1988;
Loksa,1966a,1969a;Szinetár,1992d,1996.
Kisbenedek, 1991; Loksa, 1958, 1961c, 1966a, 1969a, 1971, 1977,
Segestria senoculata (Linnaeus, 1758)
1988,1991a;Loksa&Loksa,1991b;Pillich,1914;Szinetár,1991.
Balogh, 1938b; Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1930;
Loksa,1966a,1969a,1971,1984;Szinetár,1992b.
Family ULOBORIDAE
Family DYSDERIDAE Hyptiotes paradoxus (C. L. Koch, 1834)
Dasumia canestrinii (L. Koch, 1876) Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;Loksa,1969a,
1984;Szinetár,1988,1992c,d,1996.
Loksa,1969a,1971.
Uloborus plumipes Lucas, 1846
Dysdera crocota C. L. Koch, 1838
Szinetár,1992e.
Kolosváry,1930;Loksa,1966a,1969a;Szinetár,1992a,b.
Uloborus walckenaerius Latreille, 1806
Dysdera erythrina (Walckenaer, 1802)
Balogh,1938d;Balogh&Loksa,1946,1948b;Chyzer&Kulczyn´ski,
Balogh, 1933; Chyzer & Kulczyn´ski, 1918a; Kisbenedek, 1991;
1918a; Kolosváry, 1935d, 1943; Loksa, 1967, 1969a, 1984, 1987;
Kolosváry, 1933b; Loksa, 1966a, 1969a, 1971, 1978, 1988, 1991b;
Pillich,1914;Szinetár,1992b,1996.
Szinetár,1991.
Dysdera hungarica Kulczyn´ski, 1897
Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918b; Loksa, 1966a,
1969a,1971,1973a,1977,1981b;Pillich,1914;Tóthetal.,1996. Family NESTICIDAE
Dysdera longirostris Doblika, 1853 Nesticus cellulanus (Clerck, 1757)
Balogh,1938d;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a; Balogh & Loksa, 1946; Chyzer & Kulczyn´ski, 1918a; Kolosváry,
Kolosváry,1928b,1933b;Loksa,1966a,1969a,1971,1973a,1981b, 1928a,b, 1933d, 1934c, 1939a; Loksa, 1958, 1962; Szinetár, 1992e;
1988,1991b;Szinetár,1992b. Szombathy,1917.
Dysdera ninnii Canestrini, 1868
Balogh,1938b;Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;
Kasper,1992;Loksa,1966a,1969a,1971,1984,1991a;Szathmáry, Family THERIDIIDAE
1995. Achaearanea lunata (Clerck, 1757)
Dysdera westringi O. P.-Cambridge, 1872
Balogh,1938b;Balogh&Loksa,1946,1953;Chyzer&Kulczyn´ski,
Balogh,1935,1938a;Chyzer&Kulczyn´ski,1918a,b;Loksa,1969a, 1918a; Kolosváry, 1933d, 1934c; Loksa, 1958, 1962, 1981b, 1984,
1977,1984,1988. 1991a;Loksa&Loksa,1991a;Szinetár,1992a,b,c,d,e.
Harpactea hombergi (Scopoli, 1763) Achaearanea riparia (Blackwall, 1834)
Balogh, 1933, 1938b; Chyzer & Kulczyn´ski, 1918a; Kolosváry, Chyzer&Kulczyn´ski,1918a,b;Szinetár,1992c.
1928a, 1930; Loksa, 1966a, 1968, 1969a, 1971, 1984, 1988, 1991b; Achaearanea simulans (Thorell, 1875)
Szinetár,1991.
Kolosváry,1928b,c,1933b.
Harpactea lepida (C. L. Koch, 1838)
Achaearanea tepidariorum (C. L. Koch, 1841)
Bologh,1938b;Kolosváry,1928c;Loksa,1969a.
Balogh, 1933, 1938b; Chyzer & Kulczyn´ski, 1918a; Kolosváry,
Harpactea rubicunda (C. L. Koch, 1838)
1928c;Loksa,1981b,1984;Pillich,1914;Szinetár,1992e.
Balogh, 1933, 1935, 1938a; Balogh & Loksa, 1946, 1948a, 1953; Anelosimus aulicus (C. L. Koch, 1838)
Chyzer & Kulczyn´ski, 1918a; Kerekes, 1988; Kisbenedek, 1991;
Kolosváry,1928b,1933b.
Loksa, 1961c, 1966a, 1969a, 1971, 1977, 1978, 1981b, 1984, 1987,
Anelosimus pulchellus (Walckenaer, 1802)
1988,1991a,b;Pillich,1914;Samu&Lövei,1995;Samuetal.,1996;
Szathmáry,1995;Szinetár,1991,1992e,1996. Kolosváry,1928c.
Harpactea saeva (Herman, 1879) Anelosimus vittatus (C. L. Koch, 1836)
Kolosváry,1928b,1933b;Loksa,1966a,1969a. Chyzer&Kulczyn´ski,1918a;Szinetár,1992a,d,1996.
Crustulina guttata (Wider, 1834)
Balogh,1938b;Balogh&Loksa,1946,1953;Chyzer&Kulczyn´ski,
Family OONOPIDAE 1918a;Loksa,1966a,1971,1973a,1981b,1984,1987,1991a;Pillich,
Oonops pulcher Templeton, 1835 1914;Szinetár,1992b,1995a.
Crustulina sticta (O. P.-Cambridge, 1861)
Balogh,1938c;Loksa,1969a.
Chyzer&Kulczyn´ski,1918a;Kolosváry,1943.
Dipoena braccata (C. L. Koch, 1841)
Family MIMETIDAE Balogh, 1935, 1938a,b; Balogh & Loksa, 1946; Chyzer &
Ero aphana (Walckenaer, 1802) Kulczyn´ski,1918a;Loksa,1984;Pillich,1914.
Dipoena coracina (C. L. Koch, 1837)
Balogh & Loksa, 1946; Chyzer & Kulczyn´ski, 1918a; Kolosváry,
1928c; Loksa, 1981b, 1984, 1988, 1991b; Loksa & Loksa, 1991a; Balogh,1935,1938a;Loksa,1977,1988;Szinetár,1992c.
Szathmáry,1995;Szinetár,1992a,c,d,e,1995a,1996. Dipoena erythropus (Simon, 1881)
Ero furcata (Villers, 1789) Balogh,1938b;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;
Balogh,1933,1935,1938a,b;Chyzer&Kulczyn´ski,1918a;Kasper, Loksa,1988.
1992; Kolosváry, 1930; Loksa, 1966a, 1977, 1978, 1981a,b, 1984, Dipoena inornata (O. P.-Cambridge, 1861)
1987,1988,1991a,b;Szathmáry,1995;Szinetár,1992a,b. Chyzer&Kulczyn´ski,1918a.
F.Samu&C.Szinetár 165
Dipoena melanogaster (C. L. Koch, 1837) Pholcomma gibbum (Westring, 1851)
Balogh, 1935, 1938a,b,d; Balogh & Loksa, 1946; Chyzer & Balogh,1938b,c;Chyzer&Kulczyn´ski,1918a;Loksa,1966a,1984,
Kulczyn´ski, 1918a; Loksa, 1961c, 1966a, 1968, 1971, 1977, 1984, 1991b;Szinetár,1992b.
1987,1988;Pillich,1914;Szinetár,1992a,b,c,d,1995a,1996. Robertus arundineti (O. P.-Cambridge, 1871)
Dipoena nigroreticulata (Simon, 1879) Balogh & Loksa, 1953; Chyzer & Kulczyn´ski, 1918a; Kolosváry,
Chyzer&Kulczyn´ski,1918a. 1930;Loksa,1961c,1966a,1968,1971,1979,1981a,b,1987,1991a;
Dipoena prona (Menge, 1868) Samuetal.,1996;Szathmáry,1995;Tóthetal.,1996.
Robertus lividus (Blackwall, 1836)
Chyzer&Kulczyn´ski,1918a;Szinetár,1995a.
Dipoena torva (Thorell, 1875) Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1943; Loksa, 1961a,c,
1981b,1984,1987,1988,1991a,b;Pillich,1914;Samu&Lövei,1995;
Balogh & Loksa, 1946; Chyzer & Kulczyn´ski, 1918a; Szinetár,
Szinetár,1991,1992a.
1992a.
Robertus neglectus (O. P.-Cambridge, 1871)
Enoplognatha ambigua Kulczyn´ski, 1894
Loksa,1966a,1971.
Chyzer&Kulczyn´ski,1918a;Loksa,1977.
Rugathodes bellicosus (Simon, 1873)
Enoplognatha mandibularis (Lucas, 1846)
Balogh,1933;Chyzer&Kulczyn´ski,1918a;Kolosváry,1928c.
Balogh,1935,1938a;Kerekes,1988.
Simitidion simile (C. L. Koch, 1836)
Enoplognatha mordax (Thorell, 1875)
Balogh & Loksa, 1946; Chyzer & Kulczyn´ski, 1918a; Kolosváry,
Chyzer&Kulczyn´ski,1918a;Kolosváry,1930;Szathmáry,1995.
1930, 1932b, 1935b,d; Loksa, 1966a, 1981b, 1984; Pillich, 1914;
Enoplognatha oelandica (Thorell, 1875)
Szinetár,1996.
Balogh & Loksa, 1946; Chyzer & Kulczyn´ski, 1918a; Kolosváry, Steatoda albomaculata (De Geer, 1778)
1932b,1935b;Loksa,1977,1981b,1984. Balogh, 1935, 1938a; Balogh & Loksa, 1946, 1948b; Chyzer &
Enoplognatha ovata (Clerck, 1757) sensu lato, probably
Kulczyn´ski, 1918a; Jenser et al., 1997; Kolosváry, 1935a; Loksa,
incl. Enoplognatha latimana Hippa & Oksala, 1982 1966a,1977,1984,1987;Tóthetal.,1996.
Balogh,1933,1935,1938a,b,d;Balogh&Loksa,1946,1953,1956; Steatoda bipunctata (Linnaeus, 1758)
Chyzer & Kulczyn´ski, 1918a; Kasper, 1992; Kolosváry, 1928b,c, Balogh,1933,1935,1938a,b;Balogh&Loksa,1946,1953;Chyzer&
1930, 1943; Loksa, 1977, 1981b, 1984, 1991a; Loksa & Loksa, Kulczyn´ski, 1918a; Kolosváry, 1928c; Loksa, 1958, 1977, 1981b,
1991b;Pillich,1914;Szathmáry,1995;Szinetár,1992a,b,c,d,e. 1984,1991a;Szinetár,1992e,1996.
Enoplognatha ovata (Clerck, 1757) sensu stricto Steatoda castanea (Clerck, 1757)
Samuetal.,1996;Szinetár,1995a,1996. Balogh, 1933, 1938b; Balogh & Loksa, 1946, 1953; Chyzer &
Enoplognatha thoracica (Hahn, 1833) Kulczyn´ski,1918a;Kolosváry,1928b,c;Loksa,1984,1991a;Pillich,
Balogh, 1935, 1938a,b; Balogh & Loksa, 1946; Chyzer & 1914.
Steatoda grossa (C. L. Koch, 1838)
Kulczyn´ski,1918a,b;Kerekes,1988;Kisbenedek,1991;Kolosváry,
1933c;Loksa,1977,1981b,1987,1988,1991a,b;Pillich,1914;Samu Chyzer&Kulczyn´ski,1918a;Kolosváry,1928b;Szinetár,1992e.
&Lövei,1995;Samuetal.,1996;Szathmáry,1995;Szinetár,1991; Steatoda meridionalis (Kulczyn´ski, 1894)
Tóthetal.,1996. Chyzer&Kulczyn´ski,1918a.
Episinus angulatus (Blackwall, 1836) Steatoda phalerata (Panzer, 1801)
Balogh, 1938b; Loksa, 1981a, 1987, 1988; Pillich, 1914; Szinetár, Balogh, 1935, 1938a,b; Balogh & Loksa, 1946; Chyzer &
1992c,1996. Kulczyn´ski,1918a;Kerekes,1988;Kolosváry,1928b,1930;Loksa,
Episinus truncatus Latreille, 1809 1966a, 1977, 1981b, 1984, 1987, 1988; Pillich, 1914; Samu et al.,
Balogh, 1933, 1935, 1938a; Balogh & Loksa, 1946; Chyzer & 1996;Szinetár,1992b,1995a.
Kulczyn´ski, 1918a; Kisbenedek, 1991; Kolosváry, 1930; Loksa, Steatoda triangulosa (Walckenaer, 1802)
1958, 1966a, 1968, 1971, 1977, 1978, 1981b, 1984, 1988, 1991a,b; Balogh,1935,1938a,b;Chyzer&Kulczyn´ski,1918a;Loksa,1958,
Loksa & Loksa, 1991b; Samu et al., 1996; Szathmáry, 1995; 1977,1984;Pillich,1914;Szinetár,1992d,e.
Szinetár,1992a,b. Theridion blackwalli O. P.-Cambridge, 1871
Euryopis flavomaculata (C. L. Koch, 1836)
Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918b.
Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;Loksa,1961c, Theridion familiare O. P.-Cambridge, 1871
1966a,1971,1973a,1981b,1984,1987,1991a;Pillich,1914;Szinetár, Kolosváry,1933b,1934b.
1991,1992a,b. Theridion impressum L. Koch, 1881
Euryopis laeta (Westring, 1861)
Balogh,1933,1935,1938a,b;Balogh&Loksa,1946,1953;Chyzer&
Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Kolosváry, Kulczyn´ski, 1918a; Jenser et al., 1997; Kolosváry, 1933c; Loksa,
1928b;Loksa,1977. 1977,1984,1987,1991a;Pillich,1914;Samuetal.,1996;Szinetár,
Euryopis quinqueguttata Thorell, 1875 1992d,1995a,1996;Tóthetal.,1996.
Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Kerekes, 1988; Theridion melanurum Hahn, 1831
Loksa,1977. Balogh, 1935, 1938a,b; Balogh & Loksa, 1946, 1953; Chyzer &
Lasaeola tristis (Hahn, 1833) Kulczyn´ski, 1918a; Loksa, 1977, 1984, 1991a; Loksa & Loksa,
Balogh,1933;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a; 1991b;Pillich,1914;Szinetár,1992e.
Szinetár,1995a. Theridion mystaceum L. Koch, 1870
Neottiura bimaculata (Linnaeus, 1767) Kolosváry,1928c,1930;Szinetár,1992a,d,1996.
Balogh, 1938b; Balogh & Loksa, 1953; Chyzer & Kulczyn´ski, Theridion nigrovariegatum Simon, 1873
1918a,b; Kisbenedek, 1991; Loksa, 1984, 1987, 1988, 1991a,b; Balogh,1935,1938a,b;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,
Pillich, 1914; Samu et al., 1996; Szathmáry, 1995; Szinetár, 1918a;Loksa,1996a,1977,1984;Pillich,1914;Szinetár,1996.
1992a,b,c,d,1995a,1996;Tóthetal.,1996. Theridion pictum (Walckenaer, 1802)
Neottiura suaveolens (Simon, 1879)
Chyzer&Kulczyn´ski,1918a;Loksa,1981b,1984;Loksa&Loksa,
Balogh, 1935, 1938a,b,c; Kisbenedek, 1991; Loksa, 1977; Samu 1991b;Szathmáry,1995;Szinetár,1992a,b.
etal.,1996. Theridion pinastri L. Koch, 1872
Paidiscura pallens (Blackwall, 1834)
Balogh, 1935, 1938a,b; Balogh & Loksa, 1946; Chyzer &
Chyzer & Kulczyn´ski, 1918a; Loksa, 1984; Szathmáry, 1995; Kulczyn´ski,1918a;Jenseretal.,1997;Kolosváry,1933b,c;Loksa,
Szinetár,1996. 1977;Szathmáry,1995;Szinetár,1992c,d,1996.
166 ChecklistofHungarianspiders
Theridion sisyphium (Clerck, 1757) Centromerus cavernarum (L. Koch, 1872)
Balogh, 1933, 1938b; Chyzer & Kulczyn´ski, 1918a; Kolosváry, Balogh&Loksa,1947b;Loksa,1971,1991b.
1928b,c, 1930, 1933b,c, 1935d; Loksa, 1984, 1989; Pillich, 1914; Centromerus incilium (L. Koch, 1881)
Szathmáry,1995;Szinetár,1996. Chyzer&Kulczyn´ski,1918a;Loksa,1961c,1988.
Theridion tinctum (Walckenaer, 1802) Centromerus sellarius (Simon, 1884)
Balogh, 1935, 1938a,b; Balogh & Loksa, 1946; Chyzer & Chyzer&Kulczyn´ski,1918a;Loksa,1971,1991b.
Kulczyn´ski, 1918a; Kolosváry, 1933c; Loksa, 1977, 1984; Pillich, Centromerus serratus (O. P.-Cambridge, 1875)
1914;Szinetár,1988,1992a,b,c,1996.
Balogh & Loksa, 1947b; Kisbenedek, 1991; Loksa, 1961c, 1966a,
Theridion varians Hahn, 1833
1968,1971,1988;Szinetár,1991.
Balogh, 1933, 1935, 1938a; Balogh & Loksa, 1946; Chyzer & Centromerus silvicola (Kulczyn´ski, 1887)
Kulczyn´ski,1918a;Jenseretal.,1997;Kolosváry,1928b,c;Loksa,
Kolosváry,1943.
1977, 1981b, 1984, 1987, 1991a; Loksa & Loksa, 1991a; Pillich,
Centromerus sylvaticus (Blackwall, 1841)
1914;Szathmáry,1995;Szinetár,1988,1992a,b,c,d,1996.
Balogh, 1935, 1938a,b; Balogh & Loksa, 1946, 1947b; Chyzer &
Kulczyn´ski,1918a;Kerekes,1988;Kolosváry,1943;Loksa,1961a,c,
Family THERIDIOSOMATIDAE 1966a,1968,1973a,1977,1978,1981a,b,1987,1988,1991a,b;Samu
&Lövei,1995;Szinetár,1992a,b.
Theridiosoma gemmosum (L. Koch, 1877)
Ceratinella brevipes (Westring, 1851)
Balogh,1933,1938c,d;Balogh&Loksa,1953;Loksa,1972,1973b,
Loksa,1981a,1987,1988,1991b.
1981a,1991a.
Ceratinella brevis (Wider, 1834)
Balogh, 1935, 1938a; Balogh & Loksa, 1947a, 1953; Chyzer &
Family ANAPIDAE Kulczyn´ski,1918a;Kerekes,1988;Kisbenedek,1991;Loksa,1961a,
Comaroma simoni Bertkau, 1889 1968,1973a,1977,1981b,1991a;Szathmáry,1995;Szinetár,1991.
Ceratinella major Kulczyn´ski, 1894
Chyzer&Kulczyn´ski,1918a.
Balogh,1938b;Kolosváry,1939b;Loksa,1966a,1979,1987.
Ceratinella scabrosa (O. P.-Cambridge, 1871)
Family MYSMENIDAE Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;Loksa,1971,
Mysmenella jobi (Kraus, 1967) 1987,1991a,b;Szinetár,1991.
Ceratinopsis romana (O. P.-Cambridge, 1872)
Balogh,1938c;Loksa,1973b,1981a,b,1987,1991b;Szinetár,1995b.
Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Kerekes, 1988;
Loksa,1977,1981b.
Family LINYPHIIDAE Ceratinopsis stativa (Simon, 1881)
Abacoproeces saltuum (L. Koch, 1872) Kolosváry,1930;Loksa,1981a,1987;Pillich,1914;Szinetár,1993.
Balogh & Loksa, 1953; Kolosváry, 1933b, 1934b; Loksa, 1971, Dicymbium nigrum (Blackwall, 1834)
1973a,1981a,b,1987,1988,1991a,b;Szinetár,1991,1992b. Balogh&Loksa,1947a;Chyzer&Kulczyn´ski,1918a;Pillich,1914;
Acartauchenius scurrilis (O. P.-Cambridge, 1872) Szinetár,1992a,1995a.
Balogh & Loksa, 1946, 1947a; Chyzer & Kulczyn´ski, 1918a; Diplocephalus alpinus (O. P.-Cambridge, 1872)
Kerekes,1988;Kisbenedek,1991;Loksa,1984. Balogh,1938b;Balogh&Loksa,1947a.
Allomengea vidua (L. Koch, 1879) Diplocephalus cristatus (Blackwall, 1833)
Loksa,1981a;Szinetár,1993. Balogh & Loksa, 1946, 1947a; Chyzer & Kulczyn´ski, 1918a;
Araeoncus anguineus (L. Koch, 1869) Kolosváry,1939a.
Balogh&Loksa,1956. Diplocephalus latifrons (O. P.-Cambridge, 1863)
Araeoncus humilis (Blackwall, 1841) Loksa,1991b.
Balogh, 1935, 1938a; Balogh & Loksa, 1947a, 1953; Chyzer & Diplocephalus picinus (Blackwall, 1841)
Kulczyn´ski, 1918a; Kerekes, 1988; Loksa, 1961c, 1966a, 1977, Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;Loksa,1968,
1981b,1984,1987,1991a;Pillich,1914;Samuetal.,1996;Szinetár, 1971,1978,1981b,1987,1991a,b;Szathmáry,1995.
1995a;Tóthetal.,1996. Diplostyla concolor (Wider, 1834)
Bathyphantes approximatus (O. P.-Cambridge, 1871)
Balogh,1935,1938a,b;Balogh&Loksa,1946,1947b,1953;Chyzer
Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;Loksa,1981b, &Kulczyn´ski,1918a;Kerekes,1988;Kisbenedek,1991;Kolosváry,
1984,1991a. 1928b;Loksa,1961a,1966a,1971,1973a,1977,1981a,b,1987,1988,
Bathyphantes gracilis (Blackwall, 1841) 1991a,b;Loksa&Loksa,1991a;Samu&Lövei,1995;Samuetal.,
Balogh,1933;Balogh&Loksa,1947b;Chyzer&Kulczyn´ski,1918a; 1996,Szathmáry,1995;Szinetár,1991,1992a,b,1995b;Tóthetal.,
Kerekes,1988;Loksa,1973a,1981b,1984,1987,1991a;Samuetal., 1996.
1996;Szinetár,1993. Dismodicus bifrons (Blackwall, 1841)
Bathyphantes nigrinus (Westring, 1851) Balogh&Loksa,1953;Loksa,1984,1991a.
Balogh & Loksa, 1947b; Chyzer & Kulczyn´ski, 1918a; Kolosváry, Dismodicus elevatus (C. L. Koch, 1838)
1933b, 1934b; Loksa, 1981a, 1984, 1991a; Szathmáry, 1995; Balogh&Loksa,1947b;Chyzer&Kulczyn´ski,1918a.
Szinetár,1992b,c,1993,1995b. Donacochara speciosa (Thorell, 1875)
Bathyphantes similis Kulczyn´ski, 1894
Balogh & Loksa, 1947b, 1953; Chyzer & Kulczyn´ski, 1918a,b;
Szinetár,1995a. Loksa,1987,1991a;Szathmáry,1995;Szinetár,1993,1995b.
Bolyphantes alticeps (Sundevall, 1833) Drapetisca socialis (Sundevall, 1833)
Chyzer&Kulczyn´ski,1918a. Balogh,1938b;Balogh&Loksa,1947b,1953;Chyzer&Kulczyn´ski,
Bolyphantes luteolus (Blackwall, 1833)
1918a;Kolosváry,1928c,1933b;Loksa1966a,1981a,b,1984,1991a.
Chyzer&Kulczyn´ski,1918a;Loksa,1961c,1984. Entelecara acuminata (Wider, 1834)
Centromerita bicolor (Blackwall, 1833) sensu lato,
Balogh & Loksa, 1946, 1947b; Chyzer & Kulczyn´ski, 1918a;
probably incl. Centromerita concinna (Thorell, 1875) Kolosváry,1928b;Loksa,1968,1981b,1984;Pillich,1914;Szinetár,
Chyzer&Kulczyn´ski,1918a;Loksa,1966a,1971. 1992a,c.
Centromerus aequalis (Westring, 1851) Entelecara congenera (O. P.-Cambridge, 1879)
Loksa,1981b,1991a. Szinetár,1992d.
F.Samu&C.Szinetár 167
Entelecara flavipes (Blackwall, 1834) Lepthyphantes alacris (Blackwall, 1853)
Loksa,1981a;Szinetár,1992d. Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928b, Loksa, 1966a,
Erigone atra Blackwall, 1833 1984.
Balogh, 1935, 1938a,b; Balogh & Loksa, 1953; Chyzer & Lepthyphantes angulatus (O. P.-Cambridge, 1881)
Kulczyn´ski, 1918a; Loksa, 1961c, 1977, 1981b, 1984, 1987, 1991a; Kolosváry,1928b.
Pillich,1914;Szathmáry,1995;Tóthetal.,1996. Lepthyphantes angulipalpis (Westring, 1851)
Erigone dentipalpis (Wider, 1834)
Balogh & Loksa, 1947b; Chyzer & Kulczyn´ski, 1918b; Kerekes,
Balogh,1933,1938b;Balogh&Loksa,1947b;Chyzer&Kulczyn´ski, 1988; Kolosváry, 1933b; Loksa, 1966a, 1971, 1978, 1988, 1991a,b;
1918a;Kerekes,1988;Kolosváry,1928b;Loksa,1958,1981b,1984, Szinetár,1991,1992a.
1987;Loksa&Loksa,1991a;Samuetal.,1996;Szathmáry,1995; Lepthyphantes cristatus (Menge, 1866)
Szinetár,1992d;Tóthetal.,1996.
Balogh&Loksa,1947b;Kolosváry,1939a;Loksa,1971,1984,1988,
Erigonella ignobilis (O. P.-Cambridge, 1871)
1991b;Szinetár,1992a.
Balogh&Loksa,1953;Loksa,1987,1991a. Lepthyphantes crucifer (Menge, 1866)
Erigonoplus globipes (L. Koch, 1872)
Chyzer&Kulczyn´ski,1918a;Kolosváry,1930;Loksa,1966a,1971,
Balogh & Loksa, 1946, 1947b; Kolosváry, 1932b, 1935b; Loksa, 1984,1988,1991b.
1966a,1984. Lepthyphantes flavipes (Blackwall, 1854)
Floronia bucculenta (Clerck, 1757)
Balogh & Loksa, 1947b; Chyzer & Kulczyn´ski, 1918a,b; Kerekes,
Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928b,c, 1933b; Loksa, 1988; Kisbenedek, 1991; Loksa, 1966a, 1971, 1973a, 1978, 1979,
1991a;Szinetár,1992c. 1981a,b,1987,1988,1991a,b;Loksa&Loksa,1991b;Samuetal.,
Frontinellina frutetorum (C. L. Koch, 1834) 1996;Szinetár,1991,1992a,b.
Balogh,1938b;Chyzer&Kulczyn´ski,1918a;Kolosváry,1930,1943; Lepthyphantes geniculatus Kulczyn´ski, 1898
Loksa,1984,1988;Szinetár,1996. Kerekes,1988.
Glyphesis servulus (Simon, 1881) Lepthyphantes istrianus Kulczyn´ski, 1914
Loksa,1981a;Szinetár,1993. Kolosváry,1934c.
Glyphesis taoplesius Wunderlich, 1969 Lepthyphantes keyserlingi (Ausserer, 1867)
Loksa, 1981a (described as Glyphesis conicus Loksa); Szinetár, Balogh, 1935, 1938a; Balogh & Loksa, 1946, 1947b; Chyzer &
1995b. Kulczyn´ski,1918a;Loksa,1996a,1977,1984,1988.
Gnathonarium dentatum (Wider, 1834) Lepthyphantes leprosus (Ohlert, 1865)
Balogh,1933;Balogh&Loksa,1947b;Chyzer&Kulczyn´ski,1918a; Balogh, 1933, 1935, 1938a; Balogh & Loksa, 1947b; Chyzer &
Loksa, 1981b, 1987, 1991a; Loksa & Loksa, 1991a; Szathmáry, Kulczyn´ski,1918a;Kolosváry,1928b,c,1933d,1934c,1935a;Loksa,
1995;Szinetár,1993,1995b. 1960a,1962,1977,1984,1988;Szinetár,1992e.
Gonatium hilare (Thorell, 1875) Lepthyphantes lepthyphantiformis (Strand, 1907)
Balogh&Loksa,1947b. Loksa,1970,1971.
Gonatium paradoxum (L. Koch, 1869) Lepthyphantes liguricus Simon, 1929
Balogh & Loksa, 1947b, 1953; Loksa, 1987, 1988, 1991a,b; Loksa,1966a.
Szathmáry,1995. Lepthyphantes mansuetus (Thorell, 1875)
Gonatium rubellum (Blackwall, 1841)
Chyzer&Kulczyn´ski,1918a,b;Loksa,1961a,1966a,1988,1991b.
Chyzer&Kulczyn´ski,1918a;Loksa,1981a,1984. Lepthyphantes mengei Kulczyn´ski, 1887
Gonatium rubens (Blackwall, 1833)
Chyzer & Kulczyn´ski, 1918a,b; Kolosváry, 1935a; Loksa, 1966a,
Chyzer&Kulczyn´ski,1918a;Loksa,1984. 1968,1971,1978,1981a,1987,1988,1991b;Szinetár,1992d.
Gongylidiellum murcidum Simon, 1884 Lepthyphantes minutus (Blackwall, 1833)
Balogh & Loksa, 1947b, 1953; Chyzer & Kulczyn´ski, 1918a; Chyzer&Kulczyn´ski,1918a;Kolosváry,1935a;Loksa,1984.
Kerekes, 1988; Loksa, 1961a, 1973a, 1978, 1981a,b, 1987, 1991a; Lepthyphantes mughi (Fickert, 1875)
Pillich,1914;Szathmáry,1995;Szinetár,1996.
Chyzer&Kulczyn´ski,1918a;Kolosváry,1928c;Loksa,1984.
Gongylidium rufipes (Linnaeus, 1758)
Lepthyphantes nanus Kulczyn´ski, 1898
Balogh,1933;Balogh&Loksa,1947b,1953;Chyzer&Kulczyn´ski,
Balogh&Loksa,1947b;Kisbenedek,1991;Loksa,1987,1988.
1918a;Kolosváry,1930;Loksa,1981a,1991a;Szinetár,1992a,b.
Lepthyphantes pallidus (O. P.-Cambridge, 1871)
Helophora insignis (Blackwall, 1841)
Balogh,1938b;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;
Kolosváry,1935a.
Loksa, 1959b, 1960a,b, 1961a,c, 1962, 1966a, 1968, 1970, 1971,
Hylyphantes graminicola (Sundevall, 1830)
1981a,b,1984,1987,1991a,b;Szinetár,1991,1992b.
Szinetár,1988,1992c. Lepthyphantes pillichi Kulczyn´ski, 1915
Hylyphantes nigritus (Simon, 1881)
Loksa,1978,1988;Samuetal.,1996;Tóthetal.,1996.
Kolosváry,1930;Szinetár,1992b,1995a. Lepthyphantes tenebricola (Wider, 1834)
Hypomma bituberculatum (Wider, 1834)
Kolosváry,1928a,b,1930,1932b,1935b;Loksa,1966a,1968,1984,
Balogh & Loksa, 1953; Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1991a,b;Szinetár,1991,1992a,b.
1928b,1933b;Loksa,1981b,1991a;Szinetár,1992b,1993. Lepthyphantes tenuis (Blackwall, 1852)
Hypomma cornutum (Blackwall, 1833)
Balogh & Loksa, 1946, 1947b; Chyzer & Kulczyn´ski, 1918a;
Balogh&Loksa,1946,1947b,1953;Chyzer&Kulczyn´ski,1918a; Kerekes,1988;Kolosváry,1928b,1930,1935a;Loksa,1981b,1984,
Loksa,1981b,1991a;Szinetár,1992a,b. 1987, 1988, 1991b; Samu et al., 1996; Szathmáry, 1995; Szinetár,
Hypomma fulvum (Bösenberg, 1902) 1992d,1996.
Loksa,1987. Lepthyphantes zimmermanni Bertkau, 1890
Kaestneria dorsalis (Wider, 1834) Loksa,1988;Szinetár,1992e.
Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;Loksa,1984, Linyphia hortensis Sundevall, 1830
1987,1991a;Szinetár,1992a. Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;Kerekes,1988;
Kaestneria pullata (O. P.-Cambridge, 1863)
Kolosváry, 1928c; Loksa, 1984, 1987, 1991a,b; Szathmáry, 1995;
Balogh&Loksa,1953;Loksa,1991a;Szathmáry,1995. Szinetár,1991,1992a,b.
168 ChecklistofHungarianspiders
Linyphia triangularis (Clerck, 1757) Minicia marginella (Wider, 1834)
Balogh,1933,1935,1938a,b;Balogh&Loksa,1946,1953;Chyzer& Balogh & Loksa, 1946; Loksa, 1961c, 1966a, 1971, 1984, 1987,
Kulczyn´ski,1918a;Kolosváry,1928b,c,1930,1933b,1935a,d,1943; 1991b.
Loksa, 1977, 1981b, 1984, 1987, 1991a; Loksa & Loksa, 1991b; Minyriolus pusillus (Wider, 1834)
Samuetal.,1996;Szathmáry,1995;Szinetár,1988,1992a,b,c,1996. Chyzer&Kulczyn´ski,1918a.
Lophomma punctatum (Blackwall, 1841) Moebelia penicillata (Westring, 1851)
Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;Loksa,1987, Balogh&Loksa,1947a;Chyzer&Kulczyn´ski,1918a;Pillich,1914.
1991a. Nematogmus sanguinolentus (Walckenaer, 1841)
Macrargus rufus (Wider, 1834)
Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928b; Loksa, 1981a,b,
Kolosváry,1943;Loksa,1968,1971,1991b;Szinetár,1991,1992b. 1987;Szathmáry,1995.
Maro minutus O. P.-Cambridge, 1906 Neriene clathrata (Sundevall, 1830)
Loksa,1973a,1981a,b. Balogh,1938b;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;
Maso sundevalli (Westring, 1851) Loksa, 1959c, 1973a, 1978, 1981a,b, 1984, 1987, 1988, 1991a,b;
Balogh&Loksa,1946,1947a,1953;Loksa,1981a,b,1987,1991a. Szathmáry,1995;Szinetár,1992a,b,c,e.
Mecopisthes silus (O. P.-Cambridge, 1872) Neriene emphana (Walckenaer, 1841)
Balogh,1935,1938a,b;Kisbenedek,1991;Kolosváry,1939b;Loksa, Chyzer&Kulczyn´ski,1918a;Kolosváry,1928c;Loksa,1984.
1977,1988. Neriene furtiva (O. P.-Cambridge, 1870)
Megalepthyphantes collinus (L. Koch, 1872) sensu lato, Balogh,1933;Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a.
probably incl. Megalepthyphantes pseudocollinus Saaristo, Neriene montana (Clerck, 1757)
1997 Balogh, 1938b; Chyzer & Kulczyn´ski, 1918a; Kasper, 1992;
Balogh,1933;Chyzer&Kulczyn´ski,1918a;Kolosváry,1928b,1930, Kolosváry, 1928b,c, 1930; Loksa, 1984, 1991a; Szinetár, 1991,
1933b,1943;Loksa,1961c,1966a,1971,1988,1991b. 1992b.
Megalepthyphantes nebulosus (Sundevall, 1830) Neriene peltata (Wider, 1834)
Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Loksa, 1977, Balogh,1938b;Chyzer&Kulczyn´ski,1918a;Loksa,1984.
Neriene radiata (Walckenaer, 1841)
1984;Szinetár,1992e.
Megalepthyphantes pseudocollinus Saaristo, 1997 Balogh, 1938d; Chyzer & Kulczyn´ski, 1918a; Loksa, 1981b, 1987;
Saaristo,1997. Szinetár,1995a.
Meioneta affinis (Kulczyn´ski, 1898) Oedothorax agrestis (Blackwall, 1853)
Samuetal.,1996. Balogh,1938b;Balogh&Loksa,1947b;Kolosváry,1935c;Loksa,
Meioneta fuscipalpus (C. L. Koch, 1836) 1981b,1984;Pillich,1914.
Oedothorax apicatus (Blackwall, 1850)
Balogh,1938b;Balogh&Loksa,1946,1948b;Chyzer&Kulczyn´ski,
1918a;Kolosváry,1928b,1939c;Loksa,1981b,1988. Balogh,1938b;Balogh&Loksa,1946,1947b;Chyzer&Kulczyn´ski,
Meioneta mollis (O. P.-Cambridge, 1871) 1918a;Kerekes,1988;Kolosváry,1928b,1930,1939c;Loksa,1981b,
1984; Pillich, 1914; Samu et al., 1996; Szathmáry, 1995; Szinetár,
Balogh & Loksa, 1947b; Chyzer & Kulczyn´ski, 1918a; Kolosváry,
1992a,e,1995a,b;Tóthetal.,1996.
1928c,1930;Loksa,1981b.
Oedothorax fuscus (Blackwall, 1834)
Meioneta rurestris (C. L. Koch, 1836)
Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;Loksa,1991a;
Balogh, 1933, 1935, 1938a; Balogh & Loksa, 1946, 1953, 1956;
Pillich,1914.
Chyzer & Kulczyn´ski, 1918a; Jenser et al., 1997; Kerekes, 1988;
Oedothorax gibbosus (Blackwall, 1841)
Kisbenedek,1991;Kolosváry,1928b,c;Loksa,1961c,1966a,1971,
1973a, 1977, 1979, 1981a,b, 1987, 1991a; Samu et al., 1996; Balogh,1938b;Szathmáry,1995.
Oedothorax retusus (Westring, 1851)
Szathmáry,1995;Szinetár,1992a,d,1996;Tóthetal.,1996.
Meioneta saxatilis (Blackwall, 1844) Balogh & Loksa, 1946, 1947b; Chyzer & Kulczyn´ski, 1918a;
Loksa,1981b. Kolosváry,1928b;Loksa,1984,1991a;Pillich,1914.
Meioneta simplicitarsis (Simon, 1884) Panamomops fagei Miller & Kratochvil, 1939
Samuetal.,1996. Loksa,1991b.
Metopobactrus ascitus (Kulczyn´ski, 1894) Panamomops mengei Simon, 1926
Loksa,1966a,1971. Balogh & Loksa, 1946, 1953; Loksa, 1971, 1973a, 1979, 1981a,b,
Metopobactrus deserticola Loksa, 1981 1987,1991a.
Panamomops sulcifrons (Wider, 1834)
Loksa,1981b.
Metopobactrus prominulus (O. P.-Cambridge, 1872) Chyzer&Kulczyn´ski,1918a;Loksa,1961c,1966a.
Pelecopsis elongata (Wider, 1834)
Loksa,1981a.
Metopobactrus rayi (Simon, 1881) Balogh,1938b;Balogh&Loksa,1946,1947a;Chyzer&Kulczyn´ski,
1918a;Loksa,1991b,Pillich,1914.
Loksa,1966a.
Pelecopsis mengei (Simon, 1884)
Micrargus herbigradus (Blackwall, 1854)
Szinetár,1995b.
Balogh, 1935, 1938a,b; Balogh & Loksa, 1953; Kisbenedek, 1991;
Pelecopsis parallela (Wider, 1834)
Loksa,1961a,1971,1977,1981a,b,1987,1991a,b;Szathmáry,1995.
Micrargus laudatus (O. P.-Cambridge, 1881) Balogh & Loksa, 1947a; Chyzer & Kulczyn´ski, 1918a; Kerekes,
1988;Kolosváry,1932b,1935b;Szinetár,1992b.
Szathmáry,1995;Tóthetal.,1996.
Pelecopsis radicicola (L. Koch, 1872)
Microlinyphia impigra (O. P.-Cambridge, 1871)
Loksa,1981a,1987,1988,1991a,b.
Chyzer&Kulczyn´ski,1918a.
Peponocranium orbiculatum (O. P.-Cambridge, 1882)
Microlinyphia pusilla (Sundevall, 1830)
Loksa,1987.
Chyzer&Kulczyn´ski,1918a;Kerekes,1988;Kolosváry,1928c;Loksa,
Pityohyphantes phrygianus (C. L. Koch, 1836)
1981b,1984;Samuetal.,1996;Szathmáry,1995;Tóthetal.,1996.
Microneta viaria (Blackwall, 1841) Balogh,1938b;Chyzer&Kulczyn´ski,1918a.
Pocadicnemis pumila (Blackwall, 1841) sensu lato,
Balogh & Loksa, 1947b; Chyzer & Kulczyn´ski, 1918a; Kerekes,
probably incl. Pocadicnemis juncea Locket & Millidge,
1988; Loksa, 1966a, 1968, 1971, 1973a, 1978, 1979, 1981b, 1987,
1991a,b;Szathmáry,1995;Szinetár,1991,1992a,b. 1953
F.Samu&C.Szinetár 169
Balogh, 1938b; Balogh & Loksa, 1947a; Loksa, 1966a, 1973a, Theonina cornix (Simon, 1881)
1981a,b,1987,1991a,b. Balogh&Loksa,1947b;Chyzer&Kulczyn´ski,1918a;Kisbenedek,
Pocadicnemis pumila (Blackwall, 1841) sensu stricto 1991.
Samuetal.,1996;Szinetár,1992b. Thyreosthenius biovatus (O. P.-Cambridge, 1875)
Poeciloneta variegata (Blackwall, 1841) Loksa,1991b.
Loksa,1966a. Thyreosthenius parasiticus (Westring, 1851)
Porrhomma convexum (Westring, 1851) Balogh,1938b;Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;
Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1933d, 1934c, 1939a; Kolosváry,1932b,1935b;Loksa,1981b,1991a.
Loksa, 1959b, 1960a, 1962, 1981b, 1988, 1991b; Szathmáry, Tiso aestivus (L. Koch, 1872)
1995. Balogh&Loksa,1953;Loksa,1991a.
Porrhomma errans (Blackwall, 1841) Tiso vagans (Blackwall, 1834)
Balogh&Loksa,1953. Chyzer&Kulczyn´ski,1918a;Szathmáry,1995.
Porrhomma microphthalmum (O. P.-Cambridge, 1871) Trematocephalus cristatus (Wider, 1834)
Chyzer & Kulczyn´ski, 1918b; Kerekes, 1988; Loksa, 1991a; Samu Balogh & Loksa, 1947b; Chyzer & Kulczyn´ski, 1918a; Kolosváry,
etal.,1996;Szinetár,1995a,1996;Tóthetal.,1996. 1928b,1933b;Szinetár,1992b.
Porrhomma montanum Jackson, 1913 Trichoncoides piscator (Simon, 1884)
Szinetár,1992a. Balogh&Loksa,1947a;Tóthetal.,1996.
Porrhomma profundum (Dahl, 1939) Trichoncus affinis Kulczyn´ski, 1894
Chyzer&Kulczyn´ski,1918a;Kolosváry,1928a,1939a;Loksa,1970. Balogh & Loksa, 1947b; Chyzer & Kulczyn´ski, 1918a; Kerekes,
Porrhomma pygmaeum (Blackwall, 1834) 1988;Kisbenedek,1991;Loksa,1966a,1981b.
Chyzer&Kulczyn´ski,1918a;Loksa,1981b,1991a;Szinetár,1991. Trichoncus auritus (L. Koch, 1869)
Porrhomma rosenhaueri (L. Koch, 1872) Balogh&Loksa,1947b;Chyzer&Kulczyn´ski,1918a;Loksa,1988.
Loksa,1961b. Trichoncus hackmani Millidge, 1956
Prinerigone vagans (Audouin, 1826) Kerekes,1988.
Balogh, 1933, 1935, 1938a; Balogh & Loksa, 1947b; Chyzer & Trichopterna cito (O. P.-Cambridge, 1872)
Kulczyn´ski,1918a;Loksa,1977,1984,1987. Balogh&Loksa,1946,1947b;Kerekes,1988;Loksa,1981b;Samu
Saloca diceros (O. P.-Cambridge, 1871) etal.,1996.
Balogh&Loksa,1947b. Trichopterna thorelli (Westring, 1861)
Saloca kulczynskii Miller & Kratochvil, 1939 Loksa,1991a.
Loksa,1971. Troxochrus scabriculus (Westring, 1851)
Silometopus curtus (Simon, 1881) Chyzer&Kulczyn´ski,1918a;Loksa,1968.
Loksa,1991b. Typhochrestus digitatus (O. P.-Cambridge, 1872)
Silometopus elegans (O. P.-Cambridge, 1872) Loksa,1987.
Loksa,1981a. Walckenaeria acuminata Blackwall, 1833
Silometopus reussi (Thorell, 1871) Balogh,1938b;Kolosváry,1933d,1934c,1939a.
Balogh & Loksa, 1947a, 1953; Chyzer & Kulczyn´ski, 1918a; Walckenaeria antica (Wider, 1834) sensu lato, probably
Jenser et al., 1997; Kerekes, 1988; Loksa, 1991a; Samu et al., incl. Walckenaeria alticeps (Denis, 1952)
1996. Balogh & Loksa, 1946, 1947b, 1953; Chyzer & Kulczyn´ski,
Sintula retroversus (O. P.-Cambridge, 1875) 1918a; Kisbenedek, 1991; Loksa, 1966a, 1971, 1973a, 1981a,b,
Chyzer&Kulczyn´ski,1918a. 1984,1987,1988,1991a,b;Pillich,1914;Szathmáry,1995;Szinetár,
Sintula spiniger (Balogh, 1935) 1991,1992a.
Balogh, 1935, 1938a; Balogh & Loksa, 1947b; Kerekes, 1988; Walckenaeria atrotibialis O. P.-Cambridge, 1878
Kisbenedek, 1991; Loksa, 1966a, 1977, 1988; Loksa & Loksa, Loksa,1968,1971,1981a,1987,1991a,b;Szinetár,1992b.
1991b;Szinetár,1996. Walckenaeria capito (Westring, 1861)
Stemonyphantes lineatus (Linnaeus, 1758) Balogh&Loksa,1947b,1953;Chyzer&Kulczyn´ski,1918a;Loksa,
Balogh, 1933, 1935, 1938a; Kerekes, 1988; Loksa, 1966a, 1977, 1966a,1981b,1984,1991a,b;Tóthetal.,1996.
1981b,1984,1987,1988,1991b;Szathmáry,1995. Walckenaeria corniculans (O. P.-Cambridge, 1875)
Syedra gracilis (Menge, 1869) Kolosváry,1932b,1935b,d.
Balogh&Loksa,1947b;Loksa,1968,1981b,1988,1991a. Walckenaeria cucullata (C. L. Koch, 1836)
Tallusia experta (O. P.-Cambridge, 1871) Balogh & Loksa, 1953; Loksa, 1966a, 1981a, 1984, 1988, 1991a;
Balogh&Loksa,1953;Kerekes,1988;Loksa,1973a,1981a,b,1987, Szinetár,1991,1992b.
1991a;Szathmáry,1995;Tóthetal.,1996. Walckenaeria cuspidata Blackwall, 1833
Tallusia vindobonensis (Kulczyn´ski, 1898) Balogh&Loksa,1947b.
Balogh&Loksa,1947b;Loksa,1981b. Walckenaeria dysderoides (Wider, 1834)
Tapinocyba affinis (Lessert, 1907) Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;Loksa,1968,
Loksa,1991b. 1971,1981a,1987,1988;Szinetár,1992a.
Tapinocyba insecta (L. Koch, 1869) Walckenaeria furcillata (Menge, 1869)
Balogh & Loksa, 1947a; Chyzer & Kulczyn´ski, 1918a; Kerekes, Balogh&Loksa,1947b;Kerekes,1988;Kisbenedek,1991;Loksa,
1988;Loksa,1961a,1968,1973a,1978,1979,1981b,1987,1991a,b; 1958,1961c,1966a,1971,1978,1981b,1984,1988,1991b;Szinetár,
Pillich,1914;Szathmáry,1995. 1991,1992b.
Tapinocyboides pygmaeus (Menge, 1869) Walckenaeria incisa (O. P.-Cambridge, 1871)
Balogh&Loksa,1947b;Loksa,1966a,1973a,1981b. Loksa,1978.
Tapinopa longidens (Wider, 1834) Walckenaeria kochi (O. P.-Cambridge, 1872)
Balogh&Loksa,1947b;Chyzer&Kulczyn´ski,1918a;Kisbenedek, Szathmáry,1995.
1991;Loksa,1966a,1984,1991b. Walckenaeria mitrata (Menge, 1868)
Taranucnus setosus (O. P.-Cambridge, 1863) Balogh, 1938b; Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928a;
Balogh&Loksa,1953;Loksa,1981a,1987,1991a. Loksa,1966a,1984,1991a,b;Szinetár,1991,1992a.
170 ChecklistofHungarianspiders
Walckenaeria nodosa O. P.-Cambridge, 1873 Tetragnatha reimoseri (Rosca, 1939)
Loksa,1981a,1991b. Balogh, 1938c; Kolosváry, 1933b; Loksa, 1967; Szinetár, 1993,
Walckenaeria nudipalpis (Westring, 1851) 1995a.
Balogh & Loksa, 1947b; Chyzer & Kulczyn´ski, 1918a; Kolosváry, Tetragnatha shoshone Levi, 1981
1933d,1934c;Loksa,1981a,b,1987,1991a;Szinetár,1992b. Szinetár,1995b;Uhletal.,1992.
Walckenaeria obtusa Blackwall, 1836 Tetragnatha striata L. Koch, 1862
Chyzer&Kulczyn´ski,1918a;Loksa,1981a,1991a,b. Chyzer&Kulczyn´ski,1918a,b;Lendl,1886b;Szinetár,1993,1995b.
Walckenaeria simplex Chyzer, 1894 Zygiella atrica (C. L. Koch, 1845)
Balogh&Loksa,1947b;Loksa,1971,1988,1991b. Szathmáry,1995.
Walckenaeria unicornis O. P.-Cambridge, 1861 Zygiella montana (C. L. Koch, 1834)
Balogh&Loksa,1953;Szathmáry,1995. Balogh,1938b;Chyzer&Kulczyn´ski,1918a;Loksa,1984.
Walckenaeria vigilax (Blackwall, 1853) Zygiella stroemi (Thorell, 1870)
Balogh & Loksa, 1947b, 1953; Loksa, 1981b, 1991a; Szathmáry, Balogh,1933,1938b;Kolosváry,1930;Loksa,1972.
1995;Tóthetal.,1996. Zygiella thorelli (Ausserer, 1871)
Balogh, 1933; Balogh & Loksa, 1946; Hörömpöly, 1967; Loksa,
1972;Szinetár,1992e.
Family TETRAGNATHIDAE
Zygiella x-notata (Clerck, 1757)
Meta menardi (Latreille, 1804)
Chyzer&Kulczyn´ski,1918a;Loksa,1972,1984.
Balogh, 1933; Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928a,
1933d, 1934c, 1935c, 1939a; Loksa, 1959b, 1960a, 1962, 1972,
1984.
Family ARANEIDAE
Metellina mengei (Blackwall, 1869)
Aculepeira armida (Audouin, 1826)
Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928b,c, 1930, 1939a;
Loksa,1972;Szathmáry,1995;Szinetár,1995a. Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Loksa, 1972,
Metellina merianae (Scopoli, 1763) 1977.
Aculepeira ceropegia (Walckenaer, 1802)
Balogh,1938b;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;
Kolosváry,1928a,1933b,d,1934c,1939a,1943;Loksa,1972,1984. Balogh, 1938b; Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1935a;
Metellina segmentata (Clerck, 1757) Loksa,1972,1984;Szinetár,1995a.
Balogh,1933,1935,1938a,b;Balogh&Loksa,1946,1953;Chyzer& Agalenatea redii (Scopoli, 1763)
Kulczyn´ski, 1918a; Kolosváry, 1928a, 1935a, 1939a, 1943; Loksa, Balogh, 1935, 1938a,b,d; Balogh & Loksa, 1946; Chyzer &
1972, 1977, 1981b, 1984, 1987, 1991a; Szathmáry, 1995; Szinetár, Kulczyn´ski, 1918a; Kolosváry, 1930, 1933c; Loksa, 1972, 1977,
1988,1992a,b,c,d,e. 1984,1987;Szathmáry,1995;Szinetár,1992a,b,1995a,1996.
Pachygnatha clercki Sundevall, 1823 Araneus alsine (Walckenaer, 1802)
Balogh,1933,1938b;Balogh&Loksa,1953;Chyzer&Kulczyn´ski, Balogh,1938b;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;
1918a; Kerekes, 1988; Kolosváry, 1928b,c; Lendl, 1886b; Loksa, Kolosváry, 1928c, 1933b; Loksa, 1972, 1981b; Szinetár, 1992c,
1981b,1984,1987,1991a;Szathmáry,1995;Szinetár,1992b,e. 1995a.
Pachygnatha degeeri Sundevall, 1830 Araneus angulatus Clerck, 1757
Balogh, 1933, 1935, 1938a,b; Balogh & Loksa, 1946; Chyzer & Balogh, 1933, 1935, 1938a,b,d; Balogh & Loksa, 1946; Chyzer &
Kulczyn´ski, 1918a; Kerekes, 1988; Kolosváry, 1928b,c; Lendl, Kulczyn´ski, 1918a; Kasper, 1992; Kolosváry, 1928b; Loksa, 1972,
1886b; Loksa, 1973a, 1977, 1981b, 1984, 1987, 1991a,b; Samu & 1977,1981b,1984,1989;Szinetár,1992a,b,c.
Lövei, 1995; Samu et al., 1996; Szathmáry, 1995; Szinetár, Araneus circe (Audouin, 1826)
1992b,c,d,1995a;Tóthetal.,1996.
Balogh, 1938d; Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928b,
Pachygnatha listeri Sundevall, 1830
1935d;Loksa,1958,1972,1984,1987.
Balogh, 1938b; Chyzer & Kulczyn´ski, 1918a; Kerekes, 1988; Araneus diadematus Clerck, 1757
Kolosváry,1932b,1935b;Lendl,1886b;Loksa,1981a,b,1984,1987;
Balogh, 1933, 1935, 1938a,b; Balogh & Loksa, 1946, 1953, 1956;
Szathmáry,1995;Szinetár,1991,1992a,b.
Chyzer & Kulczyn´ski, 1918a; Kasper, 1992; Kolosváry, 1928b,c,
Tetragnatha dearmata Thorell, 1873
1930, 1943; Loksa, 1972, 1977, 1981b, 1984, 1987, 1989, 1991a;
Kolosváry,1928c,1932b,1935b,d;Lendl,1886b;Szathmáry,1995. Szathmáry,1995;Szinetár,1988,1992a,b,c,d,e,1995a,1996.
Tetragnatha extensa (Linnaeus, 1758) Araneus grossus (C. L. Koch, 1844)
Balogh, 1933, 1938b,d; Balogh & Loksa, 1946, 1953; Chyzer & Balogh, 1933, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Loksa,
Kulczyn´ski, 1918a; Kasper, 1992; Kolosváry, 1928b,c, 1930, 1943; 1958,1972,1977,1984.
Lendl,1886a,b;Loksa,1981b,1984,1987,1991a;Loksa&Loksa, Araneus marmoreus Clerck, 1757
1991a; Samu et al., 1996; Szathmáry, 1995; Szinetár, 1988,
Balogh,1938b;Balogh&Loksa,1946,1953;Chyzer&Kulczyn´ski,
1992a,b,c,d,1993.
1918a; Kolosváry, 1928b,c; Loksa, 1972, 1981b, 1984, 1987, 1989,
Tetragnatha montana Simon, 1874
1991a;Szathmáry,1995;Szinetár,1992a,b,c,1995a.
Balogh, 1933, 1938b; Balogh & Loksa, 1946, 1953; Chyzer &
Araneus quadratus Clerck, 1757
Kulczyn´ski, 1918a; Kolosváry, 1928b,c, 1930, 1935d, 1943; Lendl,
1886b;Loksa,1987,1991a;Szinetár,1992a,b,1993,1995a. Balogh,1938b;Balogh&Loksa,1953;Chyzer&Kulczyn´ski,1918a;
Tetragnatha nigrita Lendl, 1886 Loksa,1972,1984,1987,1991a;Szathmáry,1995;Szinetár,1992b,
1995a.
Balogh, 1938d; Chyzer & Kulczyn´ski, 1918a; Kasper, 1992;
Araneus sturmi (Hahn, 1831)
Kolosváry, 1928c, 1930; Lendl, 1886b; Loksa, 1984; Szathmáry,
1995;Szinetár,1993. Balogh, 1935, 1938a; Chyzer & Kulczyn´ski, 1918a; Kasper, 1992;
Tetragnatha obtusa C. L. Koch, 1837 Kolosváry,1928b,1935d;Loksa,1972,1977,1984;Loksa&Loksa,
1991b;Szinetár,1988,1992a,c,d,1996.
Balogh,1933;Balogh&Loksa,1946;Chyzer&Kulczyn´ski,1918a;
Araneus triguttatus (Fabricius, 1775)
Kolosváry, 1928b,c, 1930; Lendl, 1886b; Loksa, 1984; Szinetár,
1988,1992c,1996. Balogh, 1933, 1938b; Balogh & Loksa, 1946, 1953; Chyzer &
Tetragnatha pinicola L. Koch, 1870 Kulczyn´ski,1918a;Loksa,1972,1984,1991a;Szinetár,1992a,b,c.
Araniella alpica (L. Koch, 1869)
Chyzer & Kulczyn´ski, 1918a; Kolosváry, 1928b,c, 1943; Loksa,
1984;Szathmáry,1995;Szinetár,1995a. Balogh,1938b;Chyzer&Kulczyn´ski,1918a;Loksa,1972.
