Table Of ContentBEYOND CLADISTICS
SPECIES AND SYSTEMATICS
www.ucpress.edu/go/spsy
The Species and Systematics series will investigate fundamental and
practical aspects of systematics and taxonomy in a series of comprehensive
volumes aimed at students and researchers in systematic biology and in the
history and philosophy of biology. The book series will examine the role of
descriptive taxonomy, its fusion with cyber-infrastructure, its future within
biodiversity studies, and its importance as an empirical science. The
philosophical consequences of classification, as well as its history, will be
among the themes explored by this series, including systematic methods,
empirical studies of taxonomic groups, the history of homology, and its
significance in molecular systematics.
Editor in Chief: Malte C. Ebach (University of New South Wales, Australia)
Editorial Board
Marcelo R. de Carvalho (Universidade de São Paulo, Brazil)
Anthony C. Gill (Arizona State University, USA)
Andrew L. Hamilton (Arizona State University, USA)
Brent D. Mishler (University of California, Berkeley, USA)
Juan J. Morrone (Universidad Nacional Autónoma de México, Mexico)
Lynne R. Parenti (Smithsonian Institution, USA)
Quentin D. Wheeler (Arizona State University, USA)
John S. Wilkins (University of Sydney, Australia)
Kipling Will (University of California, Berkeley, USA)
David M. Williams (Natural History Museum, London, UK)
University of California Press Editor: Charles R. Crumly
BEYOND CLADISTICS
THE BRANCHING OF A PARADIGM
Edited by
David M. Williams
and
Sandra Knapp
UNIVERSITY OF CALIFORNIA PRESS
BERKELEY LOS ANGELES LONDON
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distinguished university presses in the United States,
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information, visit www.ucpress.edu.
Species and Systematics, Vol. 3
For online version, see www.ucpress.edu
University of California Press
Berkeley and Los Angeles, California
University of California Press, Ltd.
London, England
© 2010 by The Regents of the University of California
Library of Congress Cataloging-in-Publication Data
Beyond cladistics: the branching of a paradigm/edited
by David M. Williams and Sandra Knapp.
p. cm.—(Species and systematics; v. 3)
Includes bibliographical references and index.
ISBN 978-0-520-26772-5 (cloth : alk. paper)
1. Cladistic analysis. 2. Biogeography.
I. Williams, D. M. II. Knapp, Sandra.
QH83.B49 2010
578.01'2—dc22 2010010575
Manufactured in the United States of America
19 18 17 16 15 14 13 12 11 10
10 9 8 7 6 5 4 3 2 1
The paper used in this publication meets the minimum
requirements of ANSI/NISO Z39.48-1992 (R 1997)
(Permanence of Paper).
Cover illustration: Sunflowers (Helianthus annuus) in
Enfield, North London, UK. Photo by Lorraine Williams.
Contents
Contributors vii
Preface xi
PART ONE
ON CHRIS
Bibliography of Works by Chris Humphries 3
1 Chris Humphries, Cladistics, and Connections 19
David M. Williams, Kåre Bremer, and Sandra Knapp
2 Ontogeny and Systematics Revisited: Developmental
Models and Model Organisms 35
Stephen Blackmore and Alexandra H. Wortley
3 Rooted in Cladistics: Chris Humphries,
Conservation—and Beyond? 47
Richard I. Vane-Wright
4 Do We Need to Describe, Name, and Classify All Species? 67
Quentin D. Wheeler
5 Floras to Phylogenies: Why Descriptive Taxonomy Matters 77
Sandra Knapp and J. Robert Press
vi / CONTENTS
PART TWO
BOTANY
6 Island Hot Spots: The Challenge of Climate Change 91
David Bramwell
7 Endemism and Evolution of the Macaronesian Flora 101
Mark A. Carine, Arnoldo Santos-Guerra, I. Rosana Guma,
and J. Alfredo Reyes-Betancort
8 Early British Collectors and Observers of the
Macaronesian Flora: From Sloane to Darwin 125
Javier Francisco-Ortega, Arnoldo Santos-Guerra, Charlie E. Jarvis,
Mark A. Carine, Miguel Menezes de Sequeira, and Mike Maunder
PART THREE
CLADISTICS
9 Monophyly and the Two Hierarchies 147
Olivier Rieppel
10 Beyond Belief: The Steady Resurrection of Phenetics 169
David M. Williams, Malte C. Ebach, and Quentin D. Wheeler
11 Monographic Effects on the Stratigraphic Distribution
of Brachiopods 197
Gordon B. Curry
12 The Eukaryote Tree of Life 219
Diana Lipscomb
PART FOUR
BIOGEOGRAPHY
13 Tethys and Teleosts 243
Peter L. Forey
14 East–West Continental Vicariance in Eucalyptus
SubgenusEucalyptus 267
Pauline Y. Ladiges, Michael J. Bayly, and Gareth J. Nelson
15 Wallacea Deconstructed 303
Lynne R. Parenti and Malte C. Ebach
Index 319
About the Editors 331
Contributors
Michael J. Bayly
The University of Melbourne, Australia
[email protected]
Stephen Blackmore
Royal Botanic Garden Edinburgh, United Kingdom
[email protected]
David Bramwell
Jardín Botánico Viera y Clavijo, Spain
[email protected]
Kåre Bremer
Stockholm University, Sweden
[email protected]
Mark A. Carine
The Natural History Museum, United Kingdom
[email protected]
Gordon B. Curry
University of Glasgow, United Kingdom
[email protected]
vii
viii / CONTRIBUTORS
Malte C. Ebach
University of New South Wales, Australia
[email protected]
Javier Francisco-Ortega
Florida International University;
Fairchild Tropical Botanic Garden
[email protected]
Peter L. Forey
The Natural History Museum, United Kingdom
[email protected]
I. Rosana Guma
Unidad de Botánica Aplicada, Instituto Canario de Investigaciones
Agrarias, Jardín de Aclimatación de La Orotava, Spain
[email protected]
Charlie E. Jarvis
The Natural History Museum, United Kingdom
[email protected]
Sandra Knapp
The Natural History Museum, United Kingdom
[email protected]
Pauline Y. Ladiges
The University of Melbourne, Australia
[email protected]
Diana Lipscomb
George Washington University
[email protected]
Mike Maunder
Al Ain Wildlife Park and Resort, Abu Dhabi;
Florida International University;
Fairchild Tropical Botanic Garden
[email protected]
Miguel Menezes de Sequeira
Universidade da Madeira Campus da Penteada, Portugal
[email protected]
CONTRIBUTORS / ix
Gareth J. Nelson
The University of Melbourne, Australia
[email protected]
Lynne R. Parenti
National Museum of Natural History, Smithsonian Institution
[email protected]
J. Robert Press
The Natural History Museum, United Kingdom
[email protected]
J. Alfredo Reyes-Betancort
Unidad de Botánica Aplicada, Instituto Canario de Investigaciones
Agrarias, Spain
[email protected]
Olivier Rieppel
The Field Museum
[email protected]
Arnoldo Santos-Guerra
Jardín de Aclimatación de La Orotava, Spain
[email protected]
Richard I. Vane-Wright
National Endowment for Science, Technology and the Arts,
United Kingdom;
Durrell Institute of Conservation and Ecology (DICE), University
of Kent, Canterbury;
The Natural History Museum, United Kingdom
[email protected]
Quentin D. Wheeler
Arizona State University
[email protected]
David M. Williams
The Natural History Museum, United Kingdom
[email protected]
Alexandra H. Wortley
Royal Botanic Garden Edinburgh, United Kingdom
[email protected]