Attached algal vegetation in running waters of J amtland, Sweden

92 Pages·2012·5.67 MB·English

ACTA PHYTOGEOGRAPHICSAU ECICA7 1
EDITD I
SVENSKVAA XTGEOGRFASIKA ASLLSAKPET
CatarJionhaa nsson
Attacheadl gavle getation
in runniwnagt ers
ofJ a mtlanSdw,e den
UPPASLA1 892

ACTA PHYTOGEOGRAPHICSAU ECICA7 1
EDIDIT
SVENSKVAA X TGEOGRFASIKA ASLLSKAPET
CatarJionhaa nsson
Attachalegdal v egetation
in
runninwga ters
ofJ fun tlnad,S weden
Almqv&i Wsitk sIenltle rnaSttioocnkahlo,l m
UPPSAL1A89 2
SuggecsitteadtJ ioohna:n sCs.1o 98n2,A. t tachaeldg vaeltg aetiinor nu nnwiantgeo fr s
JiimtlaSnwde,d Aecnt.Pa h ytogSeuoeg7cr1..U. p psala.
IBSN9 1-702-7101-(0p aperback)
IBSN9 1-702-7141 -(6c loth)
ISS0N08 4-5914
Doctotrhaelas tiU sp psaUlnai ve1r9s8i2t.y
CatarJionhaa ns1s9o8n2
©
SvensVkaax tgeogSraalfilsskkaa pet
Box5 59S,- 75212 U ppsala
EditEorriS:kj ogren
TechneidciatGlou rn:nS ejlo rs
Phototypbeys etting
TextgriuU ppppesnaA lBa
PrinitnSe wde d1e9n8 b2y
BorgstTrroymcskA eBr,Mi o tala
Contents
Introduction
1.
1.1 Aims
1.E2a rlSiweerd irsehs eaornac tht acahlegidan re u nnwiantge rs
1.T3h ien vestairgeaat ion
Bedrogceko l1o,gSy o dielp o5s,iW tast erco5u,rC sleism8 a,t e
Ve getatrieogni1ao0ln s
1.O4n t ercmosn cerantitnagca hlegcdao lm munities 10
2.M ethods 12
2.S1 amplsitnagt ions 12
2.2A nalyosfai tst acahlegda e 12
2.3E nvironamneanltyasli s 12
3.T heo ccurorfea ntctea calhgeiadnJe ii mtlsatnrde ams 14
3.1G enesruarlv ey 14
3.S2p ecdiiesst ribution 16
Districbuurtviaeocsnc ortdoit nhgKe o lmogoarpopvr o1a6ch,
Thee colocghiacraalc toefrt ihsietn idtcioscar al g1a,e9 (Oedo­
gonisupmp 1.,9 Clapdhoorgal omer2a0t,aZ y gnemba 2 1,
Mougeoet2 i2a,S pirogay2 r2a, S pirogcy2 r3a, S piryorga
d 23V,au chesrpip2a.4 ,C yclotceolmlt2aa4 ,C yclotkeililta­
zingiakniiat zin2g5i,aC nycal otkeililtaz ivn.pg liaanneat o-
v.
pho2r5a ,A mphpielurpaellu2c6i,Ad naom oeonberiasc hysira 26,
Anomoeobnreaicsh ysliarnac e2ol7a,At noam oeobnreaiycs-h
v.
sirva.t herlmi2as7, Ceratoanrecivus.sl ine2a8r,iCs o cco-
neipsl acenvt.eu ulygpalt a2 8,C ymbeldleala itacu 2l9,C ym-
bellluan a2t9a,C ymbelmliac roce3p0,hD ailaat ohmiae male
v.
mesod3o0n, Diatohmiae mavl.g ee nuifn.ut mu rgi3d1ul,a
Epitheamriga3u s1 ,E unotilau rnia3s2 ,E unoitap ectliivns.a
pectlii3ns,2aE unotpieac tliinsma in3o3r, Fr agicloanrsiat ruens
v.
constr3u3e,n Fsr ulsitrau homboidrehso mibdoe3s4 ,
v. v.
Frustruhloimaib doevs.c paita3t4a, F rustruhloimaibd oevs.
saxon3i5c,aG opmhoneimnat rivc.ait nutmr i3ca5t,Gu omm ­
phonema invpt.ur iimcl3aa6 Mt,eu rmi ndc iiarcu3l6a,Nr aev icula
crpytopchealva.c rpytocheapl3a7 ,N aviccurylpat pohcaelva.
vene37t ,(l Pinnulianrtieaar r7u3,p Stt auropnheoicesen nite-
ro3n 8,S ynedurlan3 a8 T,a blelafrenieas tr3a8t,Ta ab ellaria
jolccul3o9s,aB atracpheorusmmm oinloifrm3e,9 Stigonema
mamill4o0s,uR miv ulabriiaas ole40t,t Riiavnlaaur ihaa ema-
tit4e)1,s D iscriamnianlafynost5re 3 as l gal4 1t axa
3.3M ultivaaprpiraotaec hes 45
Speciaesss ociation 46
4.
4.1C luster oafsn paelcyiseiss 46
4.F2a ctor aonnsa pleycsiiess 4 7
4.F3a ctaonra loyfss ipse cainedes n vironfmaecnttoarls 50
Actpah tyogesouger7c1..
5.S tatrieosne mblance 52
5.F1a ctaonra laypspilsti ote hd5e 0 1 stations 52
5.O2r dinaantcdil oans siofifcs autbisoeofttn hs we h odlaest eat 54
TheTA BORDm eth5o4d, O vercahlalr acteorfti hzweah toiloen
TA BORDr es5u4l,tC haracteroiftz haietn idoinvT iA dBuORaDl
clus5t4e,rC so mparibseotnwste ehne sfiuvbep aorftt hswe h ole
dataasnedtt h see lepcatre5td,5 Uniqauleg caolm muni(t=i es
stat5io6n s)
5.A3 descriopftt hiweoh no ldea tawsietrthe spteoac ltg iane-
clud-eadv erhye terogseenotefd o autsa 56
5.O4r dinaotfai s ounb soefst t atwiiotnahs P rincCiopmaplo nents
Analysis 57
55. Ordionfat thwieho ondl aet awsietrthe ciparvoecraal(g Ri)An 6g4
6.C alssificaantdoi rodni noafta ligocanoe m miutnies 66
6.T1 owaradc sl assifiocfsa ttrieoanm s 66
6.T2 hec ommunciotnyc ept 67
6.U3 seo fn umermiectahlo ds 67
6.T4h em aianl gcaolm mun-itcioemsp arbiestowndesie ffne rent
multivaanrailaytsee s 67
65. Indicsapteocri es 73
Conclurdeimnagr ks 73
Lisofts pecies 74
7.
Rfeerences 77
Abstract
JohansCs.1o 98n2,,A ttachaeldgv aelga ettiionrn u nnwiantgeo frJ sii mntdlS,aw eden.
Actpah ytogseuoeg7cr1,..U ppsa8l4p ap..
Thes tudy wdiettahhlq esu antiatnaatlioyvfsae it sta cahlegvdae lg etiantt ihroeun n ning
wateorfJs ii mtlAalngdwa.ee r e coalnld1e4 ec ntveidr onpmaernatmaemlte earssuo rreesd­
timaitn1e 9d7 51-976T. hteo tdaalt aosf5e0 1ts treaanm3ds14 algsaple cwiaetssr eated
witnhu mermiectahlo ds inclcuodmipnogn aencnddlti u ssctreiram,ni anlaaynnstwdi i st h
statitsetsiitcnsac ll utdhieKn ogl mogosrtoavt iTshtiisc s.r eisntu hleetc eodl ogical
characteorfei azcaahlt gisaoplne cfiouens(d s feuer tJhoehra ns19s8o2in,np re)ap n.tdh e
establiosf1h 6mm aeintnty poefas l gcaolm muniitnicelsut,dh ieenicgro locgoincdailt ions
antdh dei stincitnidoincs aptoeofcfr iro ea slt istiuozdfee ,s ttreemapme,r paHtc,uo rne­,
ducticvailtcyci,ou nmt ceonlto,iu nrs,o lsautriroonu,vn edgientgaa ntcdiu ornr veenlto city.
Catarina JohanofsE scoonl,o BgoiItcnaasUlntpp yis,ta uUltnaei veBrosxi5 t5y9,,
S-72521U pspalSaw,e edn.
Actpah ytogseuoe7gc1r ..
1.
Introduction
1.1 Aims northpearrontfS wedweansc arroiuebtdy S ku(j91a6 4).
Them ajorri voefrn so rthSewrend heanv bee ebnr iefly
descrbiybQ eude nner(s16t95ewd)it t rhe spteobc ott h
Them aiani omf t hsitsu wdayts o fi nd etxhtete ont
macropahnyadtl igcva elg etaEtxipoenr.i msetnutdoainle s
whilcohc aatlt acahlegvdae lg etoarts iionangl lgea l
migrapteiroino odifc uintiyc ellular algae were carried out
taxcaa nb eu seads i ndicafort ocresr teani­nb yM iiller-(H91a7e6Sc)hk.see t lu ddireidft ial.sgeea.se ­,
vironmceonntdailti inro unnsn wiantge brost,ah s sialleg daeet acfohrmet dh esiurb stirnaas t terse,a m flow­
regards abioitnti hcse t croenaadnmibdst i iootniisnci g n ttoh e RiveLru laSelt tvoh,Kre aa l jtoiksuks,i enxg­
condiatliootnnhgse biorr deArlgsva.el g etwaitliploo ns meidc rosscloibpdoeet insh a quawriitflaho w-through
anidn t hset ream.T hietassleeg ldafreid fotwt nh set ream
bet reahteerbdeo tihnt eromfsa lgcaolm munities
aftetrh ecierfil sls iaonnfd o ramn ewc oloonny another
ando fi ndiviindduiaclsa pteocri es. substBraactkel.&u Rnedn be(r17g93 w)o rkpeadr tilny
Thper ovionfJac met l(aFnid1g )i. nt hmei ddolfe t hsea mperj oecBto.tK ha ltiasnjVdoe krkk etahlnea, t ter
locaitneS dc ansioau,t hSewrend ewne,rs et udied by
Swedewna sc hosaesnr eseaarrcebhae cauiste
Klas(v91i7k4 w)h,of lolowtehddee veloipnVm eernkt ean
proviald aersvg aer iaitnbi eodnr aoncvdke getation
durianw gh olyee aLra.k Hej almairnce enn tSrwaeld en
typeHse.r wei,t hai rna thleirm iatreedaa g,r eat
and its wterriiebn uvteasrtwiiiegtsrah et sepdte oc tat th­e
varioeftr yu nnwiantge (rFsi g2s-.5w) i tcho m­tachaeldg vaelg etabtyiK orno nbo(r17g95) .S ome
munitoifae tst acahlegcdaa ebn e s tudTiheaedr .e as treiancm esn tSrwaeldh eanvb ee ecnl assbiySfi jteedr na­
Poot(h17 98)a sr egards cboemnmtuhniiBcte irezsi.n s
hamso untaanildno sw lavnadrsi,bo eudsr toycpke s,
(1799)st udbioetdph e riphayntdpo lna nkitnoY nb ­
incluldiimnegs atnoddni eff,e tryepnoetfss u rround­
barbssaonu,t hSewrend en.
inwgo odland. A survoefy litoenra atttuarcaehl egvdae lg etiant ion
runnwiantge rsf raoplmas root fts h weo roludt sSicdaen ­
dinawvaisca a rroiuebtdy J ohansetsa ol(n.91 77).
12. EarliSewre dsihr esrecaho na ttached
algaienr unning waters
1.3T hei nvesattiigoanr ea
Thefi rst rsetluedvaiaretnesht o osfKe y l(i1n 9)o1 nt2 he
reda lgBaaet roascrphemuma ndS irodfortoimmaa ny1 .3.B1e drogcekol ogy
parotfsS wedeFno.n t(e1l79l)i 1 nvestfiergsahtweadt er
diatfoormmsn orthpearrontfJ s a mtlEaxntde.n wsoirvkesT hefol lowgienogl oignoifrcmaalt iisso unm marized
formt hneo rthpearrnot fsS wedceonn taibnoitanhtg ­ formH ogbo(m1 92a0n)dL undq(v1i9s6t9 ).
tachaendpd l ankatloganar ete h ofsoremt he Sraergeiko n
Then orthaenrdwn e stpearrnot fts h per ovince,
byS tmm( 9123a)n d Hu(s9t142e)dBt.o obe(r91g03 )
apafrortm a s maplalri tnb etweceonn,so ifas ltp in�
studtiheed ailng aGteih ses emilraJesa , m tlaMnadi.n ly
centpraarlot fsS wedewenr ien vestaisgr aetgeadr sdcsh i(sFti6sg) T..h ee astpearrcnto nsmiasitnsol fy
fershwaPtheare ophFylcoeraiead,ne dZa yeg nembayl egsr anaintdge nse isse7s)a n(tdFh isego .u thernmost
Israe(l91s3o81n,49 21,49 9)H.ec lasstihfiSeew de dipsah rmta inolfqy u artaznidst peasr ag(mFiit8ge).s.
runnwiantge rsa noi lnitgoo ttryopcpeah liZlcyeg dn ema
Thea reianb etweceonn simsatisno lfyC ambro­
streaamnsda, e utroopnheiV,ac u crhiesat reaTmhse.
Silurroica(knFs i 9g)e.,s pecairaolulLnyad kS et or­
Abisakroei ant hneo rtherpnamrootfSs wte dweans tax­
onomicaanldle yc ologiicnavlelsyt ibgyHa utsetde sdjto n.
(9142H)ea. n alyszaemdpb loetfshr olma kaensd streTamhsei nflueonfbc eed ruopcoktn h fere tiolfti htey
andr egisctheermeidac nadpl h ysipcaarla meTtheer sso.ii sil n dicwaittEehkd l un(d15'92ss) c a(lFei1 g0.) .
mosuts uelwf orfko cro mpariitssh ovene rtyh oroiung­h
Graniatnegdsn eifsorsmev se rrye sisbteadnrto cks,
vestigoaftt hdieio ant iontm hsle a koefs Ltfhmeg rainv er
whipcrho dmuocrea i(nteioslf llsfo)rew t iTlhiietn y­.
systienmJ amtlbaynQ du enner(s91t55e.)dA t c om­
prehesntsuiodvfbye o tphl atniakcn adt tacahlegidant e h ef lueonftc hseec hiosnvt esg etiassto imoenw mhoarte
Actpa hygteoosgure.7c 1.
2 CatarJiohnaan sson
Fig1.M. a po ft h5e0 s3t atsiaomnpsil ne1 d795 -1796 i nJii mtlaNnadm.ea sn dde scriopftt hiseot nasta irogeni svi enn
Johan(s1s 89o2,ni np re).p .
Actaph ytogseuoe7gc1r. .
Attachageladvl e egtatiinro unnn ngwi ate3r s
a
b
Fig2..M ontasnter eaa)Tm hseK rpujte-stnreeaarm A Mnta.rb i)Ts h.eL eipikoanal lvp,i nei nts hlNeaW t ceo,r noefr
Jamtland.
Actpa hytogseuoe7gc1r. .
Fig3.T. h Bea kvattas back,
streasmu rrounbdye d
denssper ufcroee st.
Fig4..T heB jorkbaa ck,
stresaumr roubnybd oetdh
conifearnoddu esc iduous
trees.
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