Table Of ContentANNOTATED THE VASCULAR FLORA OF THE
CHECKLIST OF
BEECH CREEK UNIT OF THE THICKET NATIONAL PRESERVE,
BIG
TYLER COUNTY, TEXAS
Brown MacRoberts and MacRoberts
Michael
Barbara
Larry H.
R.
E.
Vm, Bog 740 Columbia
Spring Branch Science Center Research,
8856 Westview Drive Shreveport, Louisiana 71 104, U.S.A.
Suzanne Birmingham
Walker
Department
c
Highway
87 South
14671 State
Texas 75973.
Texas 77005, Sheibyviile, U.S.A.
iston, U.S.A.
ABSTRACT
5S£
i^^
RESUMEN
INTRODUCTION
This manuscript the 5th part of an intended inventory, based on voucher specimens, of the vascular flora
is
we
of the Big Thicket National Preserve, southeastern Texas. In 4 previous papers, provided an annotated
Sandy and Lance (MacRoberts
checklist of the Hickory Creek, Turkey Creek, Big Creek, Rosier units et
we
2002; Brown 2005, 2006a, 2006b). In paper, provide an annotated checklist of the Beech
this
et
al. al.
Creek
Unit.
AND METHODS
SITE
West
The Gulf
Big Thicket located mostly within the longleaf pine region of the Coastal Plain in south-
is
& &
Harcombe Marks Watson Marks Harcombe Harcombe
eastern Texas 1979; 1979; 1979; 1981;
(Ajilvsgi
1993; Diggs 2006 and references therein). The Big Thicket National Preserve consists of 12
et et
al. al.
units scattered over 7 Texas counties. The units range from 222 to 10,100 hectares and total about 34,000
The and
hectares (Peacock 1994). Beech Creek Unit consists of 2,059 hectares located in western Tyler
is
County (Fig. relatively with poor drainage over most of the area. Elevation ranges from about
1). It is flat
43 meters 62 meters. Beech Creek and Beech Creek run through the unit. The Beech Creek Unit
to Little
was chosen inclusion in the Big Thicket National Preserve because of extensive stands of beech forest
for its
& &
Harcombe Marks Watson Marks Harcombe dominated by lower
(Ajilvsgi 1979; 1979; 1979; 1981). is
It
slope hardwood-pine forest (mesic lower slopes and terraces), with lesser amounts of baygall, mid-slope
hardwood
and
oak-pine floodplain hardwood-pine forest (mesic stream bottoms), floodplain forest
forest,
& &
Harcombe Van
(Harcombe Marks Marks Kley
(seasonally flooded river floodplain) 1979; 1981; 2006).
A
map
of community types can be found in Harcombe and Marks (1979). Fire suppression over the past
30 years has resulted in extensive midstory development and thus reduction of herbaceous vegetation that
t
Journal of the Botanical Research Institute of Texas 2(1)
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Location of the Beech Creek Unit, Big Thicket Nation,
1.
on September
adapted to frequent Hurricane Rita, a category 3 hurricane, passed over the unit 24
is fires.
decompose.
2005 damaging and thousands probably take decades
toppling of will for this litter to
trees.
It
Watson
Details about topography, and climate can be found in Deshotels (1978); (1979, 1982, 2006);
soils,
Harcombe and Marks Marks and Harcombe and Diggs
(2006).
(1979); (1981); et al.
been
Like most units of the Big Thicket National Preserve, the Beech Creek Unit has not the subject of
a detailed inventory.
floristic
Larry Brown 290 specimens during 9 days beginning 9 November 2006 and ending 17
collected field
October 2007. The MacRobertses collected 154 specimens during 4 field days in 2004 and 1 day in 2007.
2004 and day Sandi
Suzanne Walker collected 101 specimens during 2 days in additional in 2007. Elsik
1
and Warren Pruess together gathered 82 specimens during 2005. Geraldine Watson collected 25 specimens
3 2 3
1
Brown et a Vascular flora of the Beech Creek Unit, Big Thicket 653
I.,
over several years in the 1970s and early 1980s. Warren Pruess collected another 11 specimens over a
late
A&M
We
2 year period. Other collectors totaled 12 specimens. conducted a search of the Texas University
(TAMU)
herbarium consortium base (http://www.csdl.tamu.edu/FLORA/tracy/main/html/) ad-
data for
ditional specimens but found no additional taxa not reported here.
known
Because our purpose was to produce a of taxa to occur on the Beech Creek Unit, a vouchered
list
specimen was considered be the only evidence acceptable inclusion in the In 675 herbarium
to for
list. all,
specimens form specimens have been annotated by Larry Brown.
the data for this report. All
In most cases, nomenclature follows Jones et (1997), Kartesz (1999), Turner et al. (2003), and Diggs
al.
(2006). In cases of multiple collections, no more than 4 are listed for each taxon.
et
al.
RESULTS
The following an annotated of the Beech Creek Unit taxa. Abbreviations used are given below.
is list
SW
thaceae Chapm.,
coriacea (Pursh) 01
Ilex
Chapm. PH 002
ovata (Walter) Lindau lanceolata (Chapm.) R.W. Ilexlongipes exTrel.,
la var.
MM
SW
46 7056
ung, llexopaca Aiton,
67,
MM
SEWPLB LBSH 7058
r Gmel.) Steud., 488; llexvomitoria Sol. in Aiton,
(J.F.
2151
Araceae
MM
50
a humilis Nutt., 71
SW
Ahsaema
triphyllum Schott,
(L)
0,
1
MM
ubrum
7078
L, u
5EWp
245
spjnQsa
ArajJQ
GW
SW
2261
Aristolochiaserpentaria
L, 05;
scopallinumL,SBNP2'
Asclepiadaceae
SEWP
230
Asdepios perennis Walter,
LBRH
32061
variegata
Asclepias L,
SW Aspleniaceae
bra (Michx.) Dunal, 03; LB 3 321 11
SW
Ambrosia
artemisii
03
Centella erecta Fernald,
(Lf.)
SEWP
WP Ambrosia 265
L,
Chaerophyllum Hook. trifida
tainturieri var. tainturieri,
1 1
SW
LBRH 05
SEWP Baccharis halimifolia 32071;
maculata 414 L,
Cicuta 1
L,
33034
LB
& Bidens aristosa (Michx.) Briton,
lum leptophyllum Sprague ex
Britton
(Pers.)
P.
SEWP
33030
SW LB
Bidens bipinnata L, 252;
58
Wilson,
SW Bidens frondosa LB 33003
DC, L,
Eryngium prostratum ex 68
Nutt.
MM
99
SEWP 32783 Boltonia diffusa 71
Hydrocotyle Thunb., 262; LB Ell.,
verticillata
MK
SEWPLB
+Bradburia Semple,
pilosa 491, 501;
LBSH 32122 (Nutt.)
x.)Raf.,LB 3 1994;
SW 97
71
Sanicula canadensis
L, 71
^
WP
Sanicula
smallii E.R Bicknell,
1 1
^^
Apocynaceae
W^mMLKronquist,
J^ LB 32466 ^
Trachelospermum difforme (Walter) Gray, LB 3 998 ^J.^
A.
1 5EWp
LBJL 6a
335;
(L)
}
SW
Aquifoliaceae 063
Coreopsis lanceolata
L,
SEWP
llexambigua (Michx.) Torn, PH 001 Coreopsis tinctoria Nutt., 253
6 5 3 97
Journal of the Botanical Research Institute of Texas 2(1)
SW
32959
Elephantopus carolinianus Raeusch., 74; LBJL
Elephantopus nudatus LB 33002
Gray,
A.
Elephantopus tomentosus LB 28598a, 28353; 327;
L,
LBJL 3261
SEWP
DC, 342
ex
Erechtites hieraciifolio Raf.
(L.)
943
*Cardamine LB
hirsuta L, 3
1
MM
LBRH 32060
Muhl. ex 7139;
Erigeron strigosus Willd.,
Cardamine parviflora arenicola Schultz, LB
var. (Britt.)
L.
32457
I 31942
& LBSEWP
Eupatorium lancifolium (Torn A. Gray) Small, 3 899;
MM Bromeliaceae
7187
SEWP
395
usneoides
SEWPLB Tillandsia L,
496 (L.)
Eupatorium mohrii Greene,
MM
SW
47
Eupatorium perfoliatum
L, 83; 71 Buddlejaceae
33004
Eupatorium xpinnatifidum LB PM
Elliott, Polypremum procumbens LBJL 32601
L, 04;
SW
33039
Eupatorium LB
rotundifolium L, 39a;
Burmanniaceae
MM
SW
33009
LB
Apteria aphylla Barnhart, 7224; 88;
PM
SW43; 33016
Eupatorium serotinum LB
Michx., 02;
SEWP
422
BurmanniabifloraL,
& Greene
Euthamia ex
ieptocephala Gray) Porter
(Torn A.
LBSEWP
31894 Callitrichaceae
Britton,
MM
LBRH 32048; 7683 LB 3201
retusa (Lam.) Sch. Bip, Callitrichepeploides Nutt.,
'Facelis
SEWP
255
puichella Foug.. var.pulchella,
Gaiilardia
Campanulaceae
MM
Gamochaetacoarctata Kerguelen, 7106
MM
(Willd.)
SW
7209
Lobelia cardinalis L, 84;
SW061
Gamochaeta
purpurea Cabrera, MM
(L)
DC, LBSH 49
appendiculata 7673; 321
Lobelia A.
MM
amarum
amarum,
Heienium Rock 35
71 GW
(Raf.) H. var. SW
934
puberula Michx, 2249, 2336; LB 3
Lobelia 34;
SEWP
33023
Helianthus angustifolium 403b; LB
L,
& LBRH
bra Pavon) Greene, LB 32009; 32059;
(Ruiz
32494
Hieracium gronovii L, LB MM
7674
Hymenopappus
DC.
artemisiifolius
LBRH 32059
Nieuwl.
Triodanisperfoliata
(L.)
7148
WP
DC,
marginata LB 32487;
(Thunb.) A. 21
lia
1
;
Cab
*Hypochaeris microcephala (Sch. albiflora
Bip.) re ar.
SW110
MM
LBRH
(Kuntze) Cabrera, 32075; 7661
Caprifoliaceae
LBRH
Chambers, LB 32001; 32051
Krigiacespitosa
(Raf.)
MM
MM
Thunb, 7162
*Lonicerajaponica
7678
Willd.,
.)
SW
Sambucus
SW nigra subsp. canadensis 60,
R. Bolli,
(L.)
Mikania scandens 86 L.
(L) Willd., 33,
BT050
66
Lactuca canadensis LB 321
L,
SW
MM
Viburnum 57
acerifolium L,
Pluchea camphorata DC, 721 721 LB 32804
(L)
2;
1 ,
SW
MM
SW Viburnum dentatum 48
L,
DC, 92
Pluchea 7236;
foetida
(L.)
nudum
Viburnum nudum, 2260b; 321
Wooten &
var.
Pyrrhopappus LB L.
carolinianus (Nutt.) Standi., (
Celastraceae
WP SW
Pyrrhopappus Don) DC, 25
pauciflorus SEWP 4'
(D.
9;
1 1 1
L,
MM
LBRH 32074
Rudbeckia
hirta L, 71
2;
1
SW
Solidago LB 3301
altissima L„ 00;
1
SEWPLB
502
Solidago ludoviciana Gray) Small,
(A.
LBSEWP
Solidago rugosa aspera (Aiton) Fernald,
Mill. var.
SEWP
33024
31887; 331; LB
MM
SW SW
SEWP
*Sonchus asper (L) LB 32006; 544; Clethra alnifolia L, 20; 71
Hill,
1 1 1
MM
Nesom, 7206
(L) G.L.
i
Commelinaceae
MM
WP
&
Symphyotrichum
lateriflorum A. D. Love, 120;
(L.)
Commelina LBSH 26
321
erecta L,
GW
SEWPLB
7668; 2248; 481
MM
Commelina LBSH 20
321
virginica 71 55;
L,
LBSEWP
Symphyotrichum racemosum G.L Nesom,
(Elliott)
Convolvulaceae
31886; LB 33001
MM
Michx,
Dichondra carolinensis 71 61
Berberidaceae
Ipomoea Dennst cordatotriloba
cordatotriloba
var.
LB 33031
350;
Ipomoea pandurata Mey, LB 32493
G.
(L.)
L,SRNP3t
Betula nigra
CarpinuscarolinianaW<
Ostrya
virginiana
{M\\\)
SEWP
NyssaaquaticaL, 362
SEWP
397
Nyssa
Walter,
biflora
MM
SEWP
Marsh, 7081
Nyssa 275;
sylvatica
6 8 9
5
1 1
Brown
et al., Vascular flora of the Beech Creek Unit Big Thicket
Crassulaceae
Penthorum 32805
sedoides LB
Cupressaceae
MM MM
SW
Juniperus virginiana L var. virginiana, 7076; LB 33032 lilinum Kuhn, 96; 71 33
(L.)
;r/c/7um(L.)L.CRich,SW42
Cucurbitaceae
Matsum &
lonatus (Thunb.) Nakai LB 3301
*Citrullus
h,SW27;MM7654
Cuscutaceae
MM
Cuscuta compacta Juss. ex Choisy, 7231; LB 33007
Cyperaceae
Carex
albicans Willd. ex Spreng. var. australis (LH. Bailey)
LB31936
Rettig,
SW
Carex (Dewey) Mack, ex 27
brevior Lunell,
1
32004
Carex cherokeensis Schwein., LB
MM
complanata &
Carex LB 7677
Jorr. Hook., 31 953, 31 989;
Carex corrugate LB 3201
Fernald,
SEWP
wa(L)A.Gra
Leucothoerac 359
:emt
^Schkuhr
ex Willd. var. fioridana (L.H. Bailey) Naczi y,
MM
MM
DC,
23
& LBSH L) 71
Bryson, LB 31947, 32007; 32133; 7672
MM16
Don,
Carex fiaccosperma Dewey, LB 3201 3201 la (L) D. SE^m
4,
raL,SW89;f
33035
LB
71
32175 17;
Carex/ran/c//Kunth, LB
MM
Rhododendro hx.)Sweet,WP112
Carex giaucescens 721 LBJL 32600
Elliott, 0;
MM
SW
MM
SW Rhododendrc LBSH
7122;
escens Rudge, LBRH 32046; 7663; rorr., 18;
29;
32129
PH005
10
ZumSw
mi 31950
130,45; LB
19,
Euphorbias
GW
Acalyphagrac 2255
99;
tUSmc SEWP
Chamaesycer 247
ulata
ill,
Cyperuse
Sme SEWP
ns 251
(Lag.)
ill,
SEWPLB MM
*Cyperm 503
>eck.,
7200
Croton
capita.
rfichx.,
Cyperus 32794
erythrcrhizos Muhl., LB
SEWP
Croton glandt
us var. sep:tentrionalis Muell.-Arg.,
L.
SEWP
Cyperus haspan LB 32481
L, 356;
270
SEWP
400
Cyperus odoratus
L, SEWP
Croton monanthogynus 259
Michx.,
Cyperus pseudovegetus Steud, LB 32165
32490
Croton willdenowii Webster, LB
G.L.
MM
Cyperus 7666
virens Michx.,
32807
Euphorbia LB
coroliata
L,
LBSH 32134a
Eleocharis microcarpaTorr.,
SEWP
32460
Euphorbia dentata LB
Michx., 254;
Roem&Schult.,
berculosa (Michx.) 895;
LB:
32475
Phyllanthus caroliniensis Walter, LB
MM MM
LBRH 7662 MM
7233; 32047;
*Phyllanthus urinaria 71
L,
1
32484
KyllingaodorataVahl, LB SW
jticosa (W. Bartram) Fernald, 14
SEWP
Kyllinga pumila Michx., 399 SW
*Triadica sebifera Small, 01
(L.)
MM
Rhynchosporacaduca LBSH 32150
7127;
Elliott,
SEWP
Rhynchospora
corniculata (Lam.) A. Gray, 231
>5/nDurazz.,MM7107
+Rhynchosporadebilis LBSH LBRH 32087
32140;
Gale,
lum
Centrosema LBJH 32604
Benth,
virginiar
(L.)
Rhynchospora LBSH 32214
A.
elliottii Dietr.,
w. SEWP
L*
canadensis canadensis, 271
Cercis
Rhynchospora 32479
LB
fascicuiaris (Michx.) Vahl,
SEWP
Chamaecrista fascic ulata (Michx.) Greene, 276; LBJL
Rhynchospora (Chapm.) LBSH
globularis Small globuiaris
var.
32605
32132,32220
m
Moench,
Chamaecrista LB 32461, 321 32486,
SW nictita (L) 79,
Rhynchospora
glomerata LB 32472, 3301
Vahl, 98;
(L.)
32806
MM
Rhynchospora
inexpansa
(Michx.) Vahl, 71
1
32488
MM ..,SW 65; LB
Rhynchospora 33038
mixta 7074; LB
Britton,
Desmodium Auhl ex DC, SEWPLB 497
ciliare Willd.)
{I
Rhynchospora plumosa LB 32022; LBRH 32078;
Elliott,
m GW
Desmodium DC, 2337
glabeliu (Michx.)
SW
7676; 109
7ftimDC,MM7189
MA
Rhynchospora Chapm. LBSH 3214
ex
pusiila Curtis,
SEWP
41 21
.,
32489
LB
MM
Britton,
Scleriaoligantha Michx., LB 31 995; 7671
SEWP
43
MM T(Jacq.) Harper,
merata Michx., 7192; LB 32019; LBJL 32593;
*Kummerowia (Thumb.) LBSH 32 28
Schlindler,
striate
SW120
i 1
8
2 3
Journal of the Botanical Research Institute of Texas 2(1)
SEWPLB 495
Carya aquotica (Michx.
Nutt.,
f.)
SEWP
Carya texana Buckley, 41
31941
SW
28
& LBSH Carya tomentosa (Lam. ex Nutt.,
ex Rose) B.L Turner, Poir.)
hystricina (Small Britton
so
32119
LBRH 32050
Rhynchosialatifoiia Nutt,
LBRH LBSH
Juncus acuminatus Michx., LB 32005; 32069;
SEWP
Strophostyles umbellata (Muhl. ex Willd.) Britton, 347;
32142a
LBSH LB 32809
321
18;
31997
(Wiegand) LB
Juncus antheiatus R.L Brooks,
SEWP
& 235
Poggenb.,
Styiosanthes blflora (L) Britton, Sterns, SW
40
us Mack.,
MM
LBSH 321 52
campestre
*Trifolium Schreb., 71 51
LBRH 32070, 32077; LBSH 321
; sissimus Buckley, 23;
L.SWU4
latum
32473
LB
lappaceum 72
LB 321
*Trifolium L,
31938
Juncus effususL, LB
rem LB 999 MM
31
(L)
Poir., LBRH 32049
Juncus marginatus Rostk, 7130;
s)Swe
SEWP
355
Michx.,
MM
7665
LB 32002;
subsp. nigra
(L
rt, 32799
Lam., LB
aides
32020
Juncus LB
tenuis Willd.,
BRH: 2088
SEWP
33021
LB
408;
Wood) 935
Smyth, LB
bulbosa 31
Luzula (A.W.
Lamiaceae
SEWP
404
Shinners,
Hyptis alata
(Raf.)
SW
:a Bartr. ex Willd., 54, 55; L SEWP
Moench, 398
Lycopus rubeilus
MM
SW
32797
LB
7227;
licus L, 80;
MM
Monarda ex 7203
citriodora Cerv. Lag.,
& SEWPLB 499
Pycnanthemum
albescens Torn A. Gray,
Pycnanthemumte LB 321 76
MM
7096
L,
SEWP
LB 3201
SEWP 33010 Muhl. ex Spreng., 239;
340; LB Scutellaria elliptica
rell,
um(Sw.)Druce,SW108
69
WP
Geranium caroiinianum
L,
1 1
Grossulariaceae
SEWP
403a; LB 32024, 32784;
ilea virginica L,
IV
Haloragaceae
SEWPLB
489
_.,
Han 32594
7054
Sara, michauxii LBSH 321 36
to t Lilium
Poir.,
/I
MM
2289 33022
Nothoscordum 7216; LB
bivalve (L) Britton,
Hypericaceae texanum
(Planch.) Fernald,
Britton
var.
i.)
& &
Hook.)
Torr. A.
v.
.MM
2259;LBJL32617 LBSH 46
321
71 74;
GW
2257
Hypericum 7207;
galioide
VI
MM
SW
Aiton 7067; LB
07;
?ns (L)
f.,
SW
SEWP
31896; 95
MM
SW
Hypericum hypericoides Crantz, 73;
(L.)
& iray,SW75;LB32802
Gmel.)
Mitreola petiolata Torr. A.
(IF.
32599
32063; LBJL
SW
Lygodiaceae
Hypericum mutilum 25
L, 66,
MM
SW
SW
Sw., 87;
ray)
Triadenum Gmel.) Gleason, 85
waiter! 35,
(J.G.
SEWP
543
campestre
Sisyrinchium Bicknell,
E.P.
SW WRC
*Cuphea carthagenens /lacbr, 30;
LB
31 9i
ell,
3201
Bicknell B
E.P. i;
tj
0;SW
1
7 3
Brown etal., Vascular flora of the Beech Creek Unit
Magnoliaceae
MM
SEWP
437
Magnolia grondi 70* linata
flora L, Mill.,
Magnolia virginianaL,S\N 15
MM
LBRH 32079
P/antago 7094;
w/g/n/cfl L,
SEWP
occidental 363
Piatanus
L,
Poaceae
&
5EWPLB 504 Agrostis hyemalis (Walter) Britton, Sterns Poggenb.,
edarac/7 L,
MM
7675
32085;
LBRH 32054
*Aira caryophyllea var. capillars Mutel,
L.
MM
7'
_.,SW101
iropogon
Virginia
MM
SW
LBSH
Morella caroiiniensis Small, 21; 7090;
32121
32790
Morella Small, LB
cerifera
(L.)
Oleaceae
SEWP
409
Chionanthus
virginicus L,
GW
MM
SW
Fraxlnus americana L, 721 7063; 2252;
3,
1 1
MM
LBRH 7658
*Bromuscatharticus 32072;
Vahl,
Fraxinus pennsylvanica Marsh., LB 32795
MM
MM Chasmanthium laxum 04
(L) H.O. Yates, 71
7146
*Ligustrum sinense
Lour.,
SEWP
68
Ludwigia altemifolia L, 261 236; LB 321
,
GW
Ludwigia decurrens Walter, 2333; LB 32801
33000
LB 32459,
:u/are,
SEWP
Ludwigia glandulosa 401
Walter,
&
Gould
(Sw.) C.A. Clark var.
)
SEWP
358
Ludwigia leptocarpa Hara,
(Nutt.)
LBRH LBSH 32142
!2089;
n,
SEWP &CA
MM
Ludwigia pilosa Walter, 406; LB 32796, 3301
Gould
7165; LB
Clark,
)
Ludwigia LB 31 993
palustris (L) Elliott,
31990,32180
SEWP
Oenothera 392
biennis L,
SW
& LBRH 06
Oenothera 32052;
laciniata Torr. A. Gray,
LBSEWP
1 31949
tatum 31891; LB
LBRH 32056
Nutt,
dichotomum Gould dichotomun
hantheiium subsp.
(L)
MM
SEWP LBSH 34
Ophioglossaceae 332; 71 37, 7071; 321
SW
Underwood,
78
Botrychium biternatum
(Sav.)
Freckman & LBRH
(Muhl. ex Lelong, 32083;
L
Elliott)
Orchidaceae
SW
LBRH
praecox Watson., 32066,
Spiranthes (Walter) 41;
S.
G
v
n
dichot
(L.)
&
Freckman LBSH 32131
(Ashe) Lelong,
Gould
Dichanthelium oligosanthes oligosanthes,
(Schult.) var.
LBRH 32064
LB 32018, 32471;
MM
7136
Dichanthelium polyanthes Mohlenbr.,
(Schult.)
&
Gould
turn C.A. Clark,
(Elliott)
LBRH 32086
MM
33005
Dichanthelium scoparium (Lam.) Gould, 721 LB
8;
&
Freckmann LBRH
Lelong, 32044;
)
LBSH 32138,32123a, 32121a
MM
vr/^M
GW
Gray) Turner, 7095;
(A.
32458
KoeL, 2330; LB 32476,
Digitariaciliaris (Retz.)
SEWP
248
*Eleusineindica Gaertn.,
(L.)
32177
Elymus LB
virginicus L,
33040
Nees, LB
(Michx.)
MM
7230
Eragrostisrefracta (Muhl.) Scribn.,
33025
LB
Eragrostisspectabilis (Pursh) Steud.,
5
Journal of the Botanical Research Institute of Texas 2(1)
32010
mNutt., LB
V
LBSEWP
Mild, 31892;
LBRH 32073
L,
MM
5EWP
237
7168 Diodia virginiana L,
alter,
m
SEWPLB
492
Alton,
»//us(L)P.Beauv.subsps<
PLB 483 Houstonia pusilla Schoepf, LB 31 933
32485
LB
Michx.,
GW
SEWP
2256; LB
364;
i.,
SEWP
396
es,
MM
WRC1
33006
Panicum verrucosum MuhL, LB
402;
71
93;
1
32463
*Paspaium diiatatum LB
Poir.,
SEWPLB 494
Paspaium
Nash,
langei Fourn.)
(E.
MM
SEWP
233
*Paspalum nototum 7158 Lam.)
Fluegge, Terrell,
MM
SW
Paspaium plicatulum 90
Michx., 71
64;
32478
Paspaium setaceum LB
Michx.,
7132
MM GW
SW
*Paspalum
7202;
Steud., 70;
urvillei
Scrophulariaceae
%
LBSH
3:
MM
SW
SEWP
Bacopa monnieri Pennell, 243; 7681 112
avena 992 (L.)
Piptochaetium ;
GratiolapilosaMchx., LB 32021
L,WP117
annua
*Poa
31938
SW LB
Gratiola virginiana L„
LBSEWP
*Sacciolepis indica (L) Chase, 31897; 0'
33014
Crustacea MuelL, LB
*Lindernia (L)
F.
Dumort, 320
*Schedonorusarundinaceus LB
(Schreb.)
SW
Mecardonia acuminata (Walter) Small, 76; 7223;
IV
32803
Schizachyrium scoparium (Michx.) Nash, LB
MM
Sphenopholis intermedia (Rydb.) Rydb., LB 31 985;
SEWP
umbrosum
Micra'nthemum Gmel.) 352;
Blake,
(J.G.
LBSH 32145
Sphenopholis obtusata
(Michx.) Scribn.,
MM
GW 7667
SW
2329
Sporobolus
indicus
02;
(L.) R. Br.,
1
Mimulus LB 32787
SW alatus Aiton,
03
Tridens flavus Hitchc. var. flavus,
(L.)
1
LBRH
LB 31944;
Nuttallanthus texana (Scheele) Sutton,
LBRH 32057
Vulpia octofiora (Walter) Rydb.,
32062
Polygalaceae MM GW
SW
2254
3360a,
Scoparia dulcis L, 7221; 79;
Polygala mariana LBJL 3261
Mill.,
LB 940
"Veronica arvensis L, 31
polygama LBRH 32045
Polygala
L,
Polygonaceae
SEWP
268
Smilaxbona-noxl.,
SEWP
402a
Brunnichiaovata (Walter) Shinners, MM
7145
Smilaxglauca
Walter,
SEWPLB
Polygonum punctatum 485
Elliott, SW
SEWP
24
a 41
L,
7;
Rumexhastatulus LBRH 32055 MM
Baldw.,
7070
Smilaxpumila
Walter, 71
5,
1
SEWP
274
Polypodiaceae Smilaxrotundifolia
L,
//Morong,MM7086
pod
poly bides
Pleopeltis
f
vs&Windham,GW53;MM7105;
ara(Weatherby)Ar
SW
SEWP
402b;
81
Primulaceae
*AnagallisarvensisL,M MM
Symplocos
71
tinctoria Her.,
(L.) L.
1 <
SEWPLB
505
Halesiadiptera
Ellis,
SW
04
Styraxamericanus Lam.,
Morton,
Thelypteris kunthii (Desv.) C.V. L
7156
Crataegus
marshallii E
GW
2335
laevigata
Willd.,
Celtis
LBSH
Crataegus
ok., 3:
GW
SW
MM Ulmus 2334
atata Michx., 56;
7064
Crataegus spathulata Michx., 7166,
SEWPLB
493
Ulmus americana
L,
7Gf/ra(Andr.)Focke,MM7215
MM
7068
liana Aiton,
(Mill.)
SEWP
426
taEhrh.,
SEWPLB
482
Link,
>
3
:
Brown et Vascular flora of the Beech Creek Unit, Big Thicket
al.,
primulifoliaL,SWU
Ware,
rodiata Dufr. parviflora (Dyal) Egg. Viola
(L.)
f.
GW
2292
LBRH 32053 Viola sagittata Alton,
31986
ild.,GW 2294; LB
Verbenaceae
MM
americana 7084
Callicorpo
L,
SW
SEWP
256 Ampelopsisarborea Koehne, 29
(Sweet) Troncoso,
ilchella (L.)
SW SEWP
250
Greene, 94 Parthenocissus quinquefolia (L) Planch.,
..)
MM GW
SEWP 2428
258 7079;
HiensisVeW., chx.,
MM
MM
7055
7143
Verbena Small rotundifolia Michx.,
halei
Vitis
SEWP
257
*Verbena
Spreng.,
rigida
Xyridaceae
LBSEWP
Chapm.
Violaceae 31893,
Xyris difformis var. difformis,
MM
MM
7237
LB 3301 7204,
lanceoiata 7092;
Viola L,
MM GW
PM SEWP 32462
LB
Viotapatmata 7159; 2295; 05 +Xyris]upicai L.C. Rich., 41 354, 328;
L, 9,
Addendum
(whorled pogonia) and sugar maple (probably either Acer barbatum or Acer leucoderme)
Isotria ve ai
rticillata
on Beechwoods Guide, mimeographed guide available the entrance, b\
listed for the unit the Trail a at trail
we them during our two- year survey, and could not find herbarium material of sami
failed to collect field
making
Geraldine Watson (2006) also indicated that sugar maple (Acer barbatum) a remarkable recover
is
with removal
in this unit the of cattle grazing.
Carex The specimen was immature The
section ovales. in this collection too for specific identification.
was Perhaps was an immature
only other Carex in section ovales C. brevior. this C. brevior.
We
Bartonia texana. Currently Beech Creek the only Big Thicket unit to have this rare Texas endemic.
is
made two sphagnum mat Beechwoods and one from any near
collections, one in a along the Trail, far trail,
Beech Creek with Beech Creek.
the intersection of Little
We An
name
Bradburia pilosa. are following Semple (2006). equally correct is Chrysopsis pilosa Nutt.
Brown
Rhynchospora This Texas and the the Pineywoods. See
debilis. the third collection for for
first
is
(2007) additional information.
et for
al.
Xyris jupicai. There no consensus the native or introduced status of this yellow-eyed-grass.
as to
is
DISCUSSION
There are 103 families and 229 genera represented for the 470 taxa (418 native) on the Beech Creek Unit
list.
Lamiaceae
Cyperaceae Fabaceae
Eight families: Poaceae (59 taxa), Asteraceae (53 taxa), (40 taxa), (26 taxa),
48.9%
and account
Juncaceae
(15 taxa), Euphorbiaceae (13 taxa), Scrophulariaceae (11 taxa), (11 taxa) for
The Rhynchospora
of the taxa collected. largest genera are: Carex (15 taxa), Dichanthelium (13 taxa), (11 taxa),
and
Juncus (10 taxa), Quercus (9 taxa), Cyperus (7 taxa).
may
While is the case that our list is incomplete, as are all floras, and that a few taxa reported here no
it
longer exist on the unit, specimens of most taxa that have grown in the Beech Creek Unit during the past
We
95+ Beech Creek Unit has about
quarter century have been collected (probably %). estimate that the
450
425
native species/taxa.
to
ACKNOWLEDGMENTS
CA
Agreement 14001004
No.
This study was supported in part by a National Park Service Cooperative to
from
Harcombe and by an Taxa Inventory (ATBI) grant the author the
Paul in part All Biological to first
The
The were from author
Big Thicket Association of Saratoga, Texas. page charges paid for this grant. first
Deanna Boensch and Zhou, Thicket his passage across
appreciates Lily Big ecologists, for facilitating
fire
him
private land the northeast corner of the unit and then walking with along Little Beech Creek south
to
Journal of the Botanical Research Institute of Texas 2(1)
new
with Beech Eleven
near the confluence Creek. additic
to
Thanks Ray and
Keith
all-day hike. also to C. Telfair Eric for
II
careful reading of the paper.
A&M
and western Texas College
1979. Wild flowers of the Big Thicket, east Texas, Louisiana. Press,
G.
Ajilvsgi,
Station.
and Annotated
W.W. 2005.
Brown, MacRoberts, M.H. MacRoberts, RA. Harcombe, Pruess, D. Johnson.
L.E, B.R. I.S. Elsik,
and
Hardin
of the vascular of the Turkey Creek Unit of the Big Thicket National Preserve, Tyler
checklist flora
807-1
counties, Texas. Sida 21:1 827.
W.W. and 2006a. Annotated
Brown, MacRoberts, M.H. MacRoberts, RA. Harcombe, Pruess, S.D. Jones.
Elsik,
L.E., B.R. I.S.
Sandy County,
checklist of the vascular flora of the Big Unit of the Big Thicket National Preserve, Polk Texas.
22:705-723.
Sida
PA
W.W. and 2006b. Annotated
Brown, MacRoberts, M.H. MacRoberts, Harcombe, Pruess, S.D. Jones.
Elsik,
L.E., B.R. I.S.
checklist of the vascular flora of the Lance Rosier Unit of the Big Thicket National Preserve, Hardin County,
175-1
Texas. Sida 22:1 189.
and
and 2007. Notes on the of Texas with additions other significant
Brown, E.L DJ. Rosen, flora
Keith, Liggio.
L.E., J.
Texas 1:1255-1264.
records. Bot. Res. Inst
J.
USDA
Conservation
Deshotels, J.D. 1978. Soil survey for the Big Thicket National Preserve, Texas. Soil Service,
College
Station, Texas.
Diggs, G.M., B.L Lipscomb, M.D. Reed, and RJ. O'Kennon. 2006. Illustrated flora of east Texas. Sida, Bot. Misc.
26:1-1594.
PA
Unpublished
Harcombe, and PL Marks. 1979. Forest vegetation of the Big Thicket National Preserve. report:
New
Park Service, Santa Mexico.
U.S. Fe,
Harcombe, PA, Glitzenstein, R.G. Knox, Si. Orzell, and E.L. Bridges. 1993. Vegetation of the longleaf pine region
J.S.
of the West Gulf Coastal Plain. Proc. Ann. Tall Timbers Fire Ecol.Conf. 18:83-103.
and Montgomery. 1997. Vascular plants of Texas. Univ. Texas Press, Austin.
Jones, Wipff, P.M.
S.D., J.K.
Asynonymized and with the vascular of the United
KarteszJ.T 1999. checklist biological attributes for flora
atlas
CA
Meacham. American
Canada, and Greenland. ed. Kartesz and Synthesis of North flora.
States, st In J J.
1
North Garden. Chapel
Version Carolina Botanical
Hill.
.0.
1
and Annotated of the vascular of the Hickory
MacRoberts, M.H. MacRoberts, LE. Brown. 2002. checklist flora
B.R.,
20:781-795.
Creek Unit of the Big Thicket National Preserve, Tyler County, Texas. Sida
PL Monogr.
Marks, and PA. Harcombe. 1981. Forest vegetation of the Big Thicket, southeast Texas. Ecol.
51:287-305.
A&M
Peacock, H.H. 1994. Nature lover's guide to the Big Thicket. Texas Press, College Station.
New
Semple, J.C 2006. Bradburia. ln:The flora of North America. Vol. 20. Oxford University Press. York.
Turner, B.L, H. Nichols, G. Denny, and O. Doron. 2003. Atlas of the vascular plants ofTexas, Vol. Dicots, Vol. 2: Ferns,
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:
gymnosperms, monocots.
Sida, Bot. Misc. 24.
and O'Kennon,
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76-106.
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Watson, G.E. 1979. Big Thicket plant ecology: an introduction. Big cket Mus. Publ. Ser., No. 5, Saratoga, Texas.
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G.E. Big
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e.
