A Revision of the Subgenus Simandrena of the Genus Andrena of Eastern Asia with a Key to Palaearctic Species (Hymenoptera, Andrenidae)

23 Pages·1995·0.7 MB·English

by  Tadauchi, Osamu

九州大学学術情報リポジトリ
Kyushu University Institutional Repository
A Revision of the Subgenus Simandrena of the
Genus Andrena of Eastern Asia with a Key to
Palaearctic Species (Hymenoptera, Andrenidae)
Tadauchi, Osamu
Xu, Huan-li
徐, 環李
http://hdl.handle.net/2324/2591
出版情報：ESAKIA. 35, pp.201-222, 1995-01-31. 九州大学農学部昆虫学教室
バージョン：
権利関係：
,AIKASE :)53( .222-102 yraunaJ ,135991 201
A Revision of the Subgenus Simandrena of the Genus
Andrena of Eastern Asia with a Key to Palaearctic
Species (Hymenoptera, 2)
umasO T ADAUCHI
lacigolomot n,EyrotarobaL ytlucaF fo,erutlucirgA
,uyh ts,iuasykrKoeuvkiunFU 8 21napaJ
X U Huan-li
aimedacA ,ygolooZ fo etutitsnI ,aciniS
gnijieB ,080001anihC
Abstract. tsaE naisA seiceps fo eht sunegbus Simandrena fo eht suneg
edreasi vdenra 0s1 ediec ze eipmr nsoa,grao fncnieahrpCa Jd n.aa eereorKhTwen
,seiceps dna are described
Andrena (Simandrena) tianshana, wuae platydepressa
morf .anihC enO wen ,mynonys 1 wen sutats dna 2 wen sdrocer era .dedulcniA
yek ot seiceps fo eht sunegbus ni eht citcraealaP noiger gnidulcni9
Simandrena
.det nse essi eicnreappc sih nrtdafrneAoapNoruE
Introduction
After Smith (1869) described from Japan and Morawitz (1880)
Andrena halictoides
debircsed 4 wen seiceps morf htroN anihC dna ,ailog nloaMreve ssrohtu adetaertsiht
Andrena
usta m. u),,s69n a32e;, Y99k5d s11f1n, nnl9a.ar o A1rgieoidst.sttteuSeAsi sn(adnaeEeobgpxe
)1491( edam eht tsrif kcehc tsil fo ni nretsaE aisA gnidulcni ,anihC ,aeroK,napaJ
Andrena
fo seiceps esenapaJ deiduts )6691-2691( amihsariH .aissuR fo stcirtsiD emitiraM eht dna nawiaT
siht suneg dna ihcuadaT dna amihsariH )8891-3891( desiver lareves arenegbus gnirrucconi
.napaJ ihcuadaT )9891( detsil 38 seiceps fo
seiceps evah neeb dedrocer yb ustamusaY (1941) uW (1941), Wu (1965,
Andrena, 83
)1 stlus emRo refv int aaeyprhdaetuJpt-oSa-noi Chn CoseidutS “no ,scitametsyS no itdunlaovE
yhpar gfooe gnoaiiBsA ,. m .y,)Hae(eaddiionpe Ar.donNA .1
Andrena
akoukuF .reS( ,4 .oN.)16
Andrena
202 0. TADAUCHI & XU HUAN-LI
1982a, 1982b, 1992), miK (1985), dna ,srehto dna 44 seiceps yb miK & miK (1983), miKdna
miK )9891( dna ihcuadaT & ee L (1992), .ylevitcepser morF ,ailogn o,Maktahcm,anKeilahgaS
,rum AairebiS dna asomro Ftuoba 02 seiceps ere wdedrocer ni eht seiduts yb ztiwaroM (1880),
llerekcoC ,)1191( arumustaM (1911), Friese (1914, 1922), nekflA )9291( dna .srehtoyltneceR
eno dfeot a ep, e viim,cnic hasnitincrueataugirrpao cea adreSppnJaPhoi-Ttoa Ce1r9o9K1dna
2991 dna osla sah detrats a napaJ-anihC evitarepooC ydutS ecnis .3991 eW evah dahna
.a insrAet sdnaeiEtcel lyolce dsinwemic eepnsim ayoxtteinutroppo
eh Ttnes erreppap dna eht eru tsuefussi fo eht ema sseires fo ruo krow era ylni admesabno
eht lairetam devreserp ni eht lacigolooZ etutitsnI fo aimedacA ,aciniS ,gnijieB dna eht
.a k, oyeut Wkious sFru, lehe eavsry siuut,unytlyUKl urrufcoocoat iFalrragocAbiagLolomotnE
lliw esu eht epyt dna lam rsonnem idceewposrrob morf eht gniwo lslmoufesum dn:asetutitsni
eht larutaN yrotsiH ,muesuM ;nodnoL lanoitaN muesuM fo larutaN ,yrotsiHnainoshtimS
,noitutitsnI notgnihsaW ;.C.D naciremA muesuM fo larutaN ,yrotsiH weN ;kroYlacigolooZ
,muesuM ;nilreB lacigolooZ ,etutitsnI ymedacA fo secneicS fo ,aissuR .tS ;grubsreteP
.zniL ,smuesumsednaL nehcsihcierretsjirebO
n It ynedehus tt e estrswapu enerehtgtbus Simandrena sun ee ghfto Andrena. ehTsunegbus
si dnuof htob ni eht citcraealaP dna citcraeN .snoiger ekcnraW )8691( dedulcni 8naeporuE
seic edpnsa 2 h ntsarecoiiNcref pAnsi , seuhnt edgnbaus egreBaL )9891( desiver eht seicepsfo
eht citcraeN ,noiger gnitaert 8 .seiceps nekflA )5391( dna ekcnraW ,6691( )3791 debircsed3
de d)ks r7u es o7y mie7c9n. ocie1hn ar ecr(cariffpeytosospssnAsOi inoMathgfA Simandrena
morf naeporu E.aissuR htiW tcepser ot tsaE naisA seiceps fo Simandrena, amihsariH )2591(
dedr oecneo rhca emo rnfapa Jd in.ha acae u mr)addio3denhK8asas9Ti as1veru(ehinrtHegbus
ro fn adpea zJdinna g .o4scee irm co,erapFesr omKiK dna m di)eK9sd8er 9ioe1ccn(eeoprsbus
.seiceps 2 dedrocer )5891( uW ,anihC morF .seiceps 4 detsil )3991( eeL dna ihcuadaT dna
,epo ron usisElrau chcc o ief,hnoawo insrAet sma oEsrefic ee0pz1singo cyeedw rust isnhIt
3 esr e awidecnneap s2 e s .r edmaawzr on eioerime nncnf hyWogeowCndsoreonlcnnaaeyrs
eno wen .sutats eW tneserp a yek ot seiceps fo eht sunegbus Simandrena ni ehtcitcraealaP
,noiger gnidulcni 9 naeporuE dna htroN nacirfA seiceps elbaliava rof eht tneserp .ydutsehT
-oloh d nsdaeeptyatnagrias pendi eht tnese rrpepap e rdaevreser pni el haetctiugtoiltosonZIfo
aimedacA ,aciniS gnijieB dna emos sepytarap era ni eht lacigolomo t,nyErotarobaLuhsuyK
.akouk u,FytisrevinU
Terminology and Abbreviations
lacigolohproM smret ylniam wollof renehciM )4491( dna amihsariH .)2691(snoitaiverbbA
desu ni eht seires fo siht yduts era sa :swollof PP = ;serutcnup SI = secapsretnineewteb
punctures; ,nT nS = lamosatem mugret dna munrets n; nLF = rallegalf tnemges n; ,L W=
; )Le Wt l a=e;a1 ls hiao LpahptdLntd B btidi n nfy gtig=emaodmnwy tuooepLnmbrlaifx(am
fo gniwerof gnidulcni;aluget LH = daeh ,L morf pot fo xetrev ot rewol nigram fosuepylc
g nsisdeuclocrx pefo ;murbal l ah ithfdWcdto nH ai g dam f w=n,auW;eLemVWl VhiF=Fxam
;aevof ,lLF 2LF = htgnel fo ralleg asltfnemges 1 dna 2 fo eht elam gnola( rewol secafrusfo
;,) LdWErE a =wdr e oLhdfcnt a eW e lr;a fLaten;oPes y nLC pWe =enyGttelnn=hceawmges
W fo laneg aera nees ;yllaretal DOO = ralucolleco ;D DOP = rallecotsop ;ecnatsid DCO=
latipiccolleco ;D WsM = lamososem W neewteb( retuo smir fo ;)ealuget WtM = lamosatem.W
NOISIV E FRSOUNEGBUS F ONRETSAAIESA 203
SZMANDRENA
Subgenus Simandrena P&ez
Simandrena ,zereP 18901, setcA .toS , x u.anendirLoB 44: 174; ,ekcideH ,3391 .ttiM .looz.suM
,nilreB :91 ;812 ,llerekcoC ,6391 cificaP-naP ,.tnE :21 ;541 ,mahnaL ,9491 .vinU .filaC
.lbuP ,.tnE :8 ;591 ,egreBaL ,4691 .vinU aksarbeN .tS .suM ,.lluB :4 ;403,amihsariH
1965, J. .taF ,.rgA uhsuyK ,.vinU :31 ;794 ,ekcnraW ,8691 .meM .tsE .suM .looz .vinU
,arbmioC :)703( ;78 ,kuynhcysO ,7791 anuaF ,iniarkU 21 :)5( ;332 ihcuadaT &
,amihsariH ,3891 ,aikasE :)02( ;18 ,egreBaL ,9891 .snarT .remA .tne ,.toS 115: 1. -epyT
:seiceps Andrena propinqua ,kcnehcS 3581 = Andrena dorsata ,ybriK( )2081 detangised
y,bekc.i3d3e9H1
Platandrena ,kcereiV 1924, .danaC ,.tnE :65 ;12 ,ekcideH ,3391 .ttiM .looz .suM ,nilreB:91
;712 ,mahnaL ,9491 .vinU .filaC .lbuP ,.tnE :8 .212 epyT :seiceps Andrena nasonii
,nostreboR .5981
Stenandrena ,ekalrebmiT ,9491 ni ,mahnaL .vinU .filaC .lbuP ,.tnE :8 ;312 ,egreBaL,4691
.vinU aksarbeN .tS .suM ,.lluB :4 .403 epyT :seiceps Andrena pallidifovea (Viereck,
1904).
Diagnosis: llamS ot yletare deogmra l;sei cmeuptson otrupoht ilwarem u;hsellganeadoporp
alucibroc ni elamef ,etelpmoc tuohtiw roiretni ;sriah lasrod aera fo muedoporp deniltuoyb
etunim tub tcnitsid desiar segdir gnola eht ;seru tluasretnahco rstuccolf ;etelpmoc 1aibitapocs
yletaredom gnol ot netfo ylemertxe trohs dna ton ;degnarra dnih aibit yldaorbdednapxe
.yllacipabus
Key to the Palaearctic females of the subgenus Simandrena*
1. eni t,s ye,e lek;acrsPe uue,Tore v Gk;lsisurluf i- oxaennahloi r agloacuahgsrmtrroeestfateM
ot naeporuE aissuR lanoitidda[ lufesu :sretcarahc 1aibit apocs ,gnol llew;degnarra
suepylc ;dellud lasrod ecaf fo muedoporp ,etallesset ton denehguor htiwwollahs
;serutcnup roiretsop snoisserped fo lamosatem agret ton llew ,detacidni,etallesset
.]etatcnupmi . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . transitoria .F ztiwaroM
- lamosateM agret ;kcalb sriah no xaroht .lamron . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
2. ralle gtanleFmges 1 regnol naht stnemges 2 dna 3 ;reht e,gy oeatkirvuaTlsoguY otecnarF
[propodeal enclosure with only a few rugulae basally; supraclypeal area with
.]serutc nwuopll ahh ts,iewtalles smeutedop oef rcolpaafs r;oedalu gluarnidutignol
. . . . . . . . . . . . . . . . . . . thomsoni ekcuD
- rallegalF tnemges 1 retrohs naht ro sa gnol sa stnemges 2 dna 3 .rehtegot . . . . . . . . . . 3
3. suepylC deneergahs dna ,dellud htiw on noitacidni fo .serutcnup . . . . . . . . . . . . . . . . . 4
- suepylC ynihs ot ylkaew etallesset htiw .serutcnup . . . . . . . . . . . . . . . . . . . . . . . . . . 5
* ruoF ,seiceps hcihw ew dluoc ton ,enimaxe era dedulcxe morf eht tneserp yek sa:swollof
Andrena venerabilis , n 5e; 3knm9fao1ldrArfoJ A. melba ,ekc n6r6a9W 1 ;msoursfacuaC A.
jalalabadensis ;natsinah gmf oA3r 7f,9e1kcmaW A. antigana ,zere P5981 morf.niapS
204 .0 IHCUADAT & UXIL-NAUH
4. ;serutc nluapcipocsor cnfioomitaci denriuc shb to,iewtalles syelte sm nu1eg drleatmosateM
roiretsop lanoisserped aera fo lamosatem agret ylesned ;etallesset laedoporperusolcne
htiw ylno a wef ealugur ;yllasab lasrod ecaf fo muedoporp etallesset htiwwollahs
;serutcnup aireglA dna .occoroM . . . . . . . . . . . . . . . . . . . . . . . zereP
breviscopa
- lam omsuagtreeMt 1 htooms dna ynihs hti w,elsa nn ;reolsodileisarrsmueesttrcspnoeupdp
aera fo lamosatem agret ynihs dna htooms htiw esned ;serutcn ulpaedoporperusolcne
esogur no 31.1 fo lasab ;aera lasrod ecaf fo muedopo r;pdenehguor elddiM eporuEot
naeporuE ,aissuR elddiM ,airebiS anihC .)tsewhtron( . . . . . . . . . . (Christ)
combinata
5. dniH 1aibit apocs ,gnol yletaredom llew .degnarra . . . . . . . . . . . . . . . . . . . . . . . . . 6
- dniH 1aibit apocs yrev trohs ot ,trohs ton llew .degnarra . . . . . . . . . . . . . . . . . . . . . . . . . . 7
6. suepylC htooms dna dehsilop htiw daorb lanidutignol naidem dna a esrevsnartlacipa
etatcnup m;ise claapespylcarpu saera denehguor htiw gnort s;serutcnup sriah noxetrev
.)tsewhtro na(ni h;Cxevn oecti usquepy l;csuovl umfutucsos ensmori a;hhsiwolley
. . . . . . . . . . . . . . . . . . . .n .ps
tianshana,
- Clypeus ylkaew etallesset dna etatcnup with median longitudinal tessellated space;
llaepylcarpus aera htiw wollahs ;serutcnup sriah no xetrev elap ;hsinworb sriahno
htuo sa(er o,Kani h;Cxevn oscuepy l;chsiwoll eeyl amputucsosem & .)la rtnec
. . . . . . . . . . . . . . . . .. amihsariH
koma
7. rallegalF tnemges 1 retrohs naht stnemges 2 dna 3 .rehtegot . . . . . . . . . . . . . . . . 8
- rallegalF tnemges 1 sa gnol sa stnemges 2 dna 3 .rehtegot . . . . . . . . . . . . . . . . . . 11
8. lamosateM muget 1 ynihs htiw esne d ;l lsleaearermudus soteopcslonocrugnppuer no 311
fo lasab ;noitrop lasrod ecaf fo muedoporp etallesset htiw wollahs ,serutcnupton
;de nleahigcu aoafrevof yre v ,gdnaio yrypblurcaceo nlluf ec anpesew tteibbro dnatsop
;sulleco suepylc rehtar daorb dna etiuq ;xevnoc aireglA dna ocorroM laidigyp[etalp
tuohtiw desiar ;aera laduac airbmif .]kcalb . . . . . . . . . . . . . . . . . . . . . . . zereP
rhypara
lamosateM mugret 1 ylkaew etallesset htiw lacipocsorcim enif ;serutcnuplaedoporp
erus oelscongeur lla revo ot 312 fo las a;bnoit rloapsrod ecaf fo mu;eddeonpeohrgpuor
laicaf aevof ton daorb gniypucco l/2 ot 312 ecaps neewteb tibro dna tsop;sulleco
suepylc 9.......................................................xevnoc
9. laedoporP erusolcne egral dna yltcnitsid delknirw lla ,revo htiw esrevsnart leek ;yllacipa
mutonorp htiw lacipabus nigram eritne ro semitemos yrev ylkaew etanigrame nieht
;elddim napaJ dna aeroK htuos( & )lartnec mutonorp[ tuohtiw a naidemlanidutignol
;enil laicaf aevof ,worran gniypucco tuoba 7.0 ecaps neewteb tibro dna tsop;sulleco
xetrev htiw elap hsinworb ;sriah ,rellams htgnel 9-8.]mm
. . . . . . amihsa ritiheHcuadaT
yamato
- lac itpuaoh teitwalle s ks oaentetowit rloapc it,pudabelkn iyrlwtcnit seirdusol clnaeedoporP
esrevsnart ;leek mutonorp htiw lacipabus nigram etanigrame ni eht .elddim . . . . . . . . 10
.01 mutonorP htiw naidem lanidutignol enil ;tcnitsid laedoporp erusolcne delknirw ylraenlla
;revo sriah no , xmeuttruecvsos edmn amulletu ce sl;ahpsi wsoslelceoyrp fo murbal ton
etanigrame ni eht ;elddim lamosatem mugret 1 ylkeew etallesset htiwcipocsorcim
;l s agea inr etciudvuaytaoofpco fbunr8a c ub.ncep0 eocteaiwpbts redobn; astuslolpeco
la;, ai hrhdrtset ubhiggtamgnrnuciiwaeo flollb0ra1b-9 ;m.mnapaJ
. . . . . . . . . . . . . . . amihsariH
opacifovea
NOISIV E FRSOUNEGBUS F ONRETSAAIESA 502
SZMANDRENA
- mutonorP htiw naidem lanidutignol enil ;erucsbo laedoporp erusolcone ssel,delknirw
etallesset no 3/l fo la c;inpoait rsorpiah no ,mxuetturcesvo sde nmmaul;lseutouccssuf
ssecorp fo murbal etanigrame ni eht ;elddi mlamosatem mugret 1 yltcn,iettsaildlesset
yeltraatecnnup mrio ch;itspieowrc usltoacrinccuiap mfaevo fw ogrnriaynputcucooba
7.0 ecaps neewteb tibro dna tsop ;sulleco laduac ai r;bsm ui,ofrceslu lfhatmgsneltuoba
9-8 ;mm napaJ amihsO-imam41( .).sI . . . ,amihsariH te ihcuadaT n. .tats
austroinsularis
.11 roiretsoP lanoisserped aera fo lamosatem agret htooms dna ynihs htiw tcnitsidesned
............................serutcnup . . . . . . . . . . . . . . . . . . . . . . . . . . . . .21
- roiretsoP lanoisserped aera fo lamosatem agret yletaredom ot ylkaew etallessethtiw
.........................serutcnup . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .31
12 Clypeus smooth; propodeal enclosure rugose on basal half; facial fovea broad;
laepylcarpus aera ylgnorts ,etatcnup ton ;denehguor xetrev ton ;dnuor lamosatemriah
sdnab no agret 4-3 ;etelpmoc laidigyp etalp htiw desiar aera ton ;etallessetanihC
.) h.t r.o n.( . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .n.eauw.. sp.
- Clypeus tessellate; propodeal enclosure rugose on basal ;3/2 laicaf aevof;worran
supraclypeal area roughened; vertex round; metasomal hair bands on terga 3
detpurretni ;yllaidem laidigyp etalp htiw desiar aera ;etallesset ,dnalgnE elddiMdna
htuoS eporuE ot naeporuE .aissuR . . . . . . . . . . . . . . . . . . . . . . . kcnehcS
lepida
.31 lasroD ecaf fo muedoporp etallesset htiw wollahs ,serutcnup ton ;denehguor laicafaevof
y r ge,nv diyayloprruab celcnlou fe cnaepe swttiebbro dna t;ss ouepllldeedpciooMruE
ot naeporuE Rtrssia lamosatem[ mugret 1 ,etallesset htiw erucsbo ;serutcnuproiretsop
lanoisserped aera fo lamosatem agret ,etallesset htiw tcnitsidni.]serutcnup
. . . . . . . . . . thcenkedeimhcS
congruens
- lasroD ecaf fo muedoporp ;denehguor laicaf aevof .worran . . . . . . . . . . . . . . . . . 14
.41 lamosateM mugret 1 ,etallesset htiw enif ;serutcnup laepylcarpus aera htiwlanidutignol
;ealugur ssecorp fo murbal worran dna etanigrame ni eht ;elddi mdnalgnE dnaelddiM
eporuE ot naeporuE .aissuR . . . . . . . . . . . . . . . . . . . . . . . . . . *)ybriK(
dorsata
- lamosateM mugret 1 ynihs dna ,htooms htiw esned tcnitsid ;serutcn ulpaepylcarpusaera
tuohti wlanidutign o;lealugur ssecorp fo murbal to netanigrame ni eht .elddim . . . . 15
.51 laedoporP erusolcne ylkaew esogur no lasab .flah . . . . . . . . . . . . . . . . . . . . . . . 16
- laedoporP erusolcne yltcnitsid esogur lla .revo . . . . . . . . . . . . . . . . . . . . . . . . . 17
.61 suepylC ynihs dna ,htooms htiw tcn i;tsseird ultacmnouspate mag r,eettalless ehttiwllams
lan o g rian;osei sisrytereapureruoutpcbtsecacodonpup 3/l .na p;aeJtalles s,emtug rfeot
. . . . . ihcuad aatTmeihsariH
nippon
- suepyl Cy l,keateawlles shettiw a nlaai ndeietdmauttcing un;poemlci alpasmosateamgret
yrev ylkaew etallesset dna ynihs htiw tcnitsid esned ;serutcnup roiretsoplanoisserped
tu ogbnaiypu caceora 3/2 .)htro na(ni h;Cetalless eytlka eywr e,vmugr efto
. . . . . . . . . . . . . . . . .n.ps
platydepressa,
* Wu (1985) recorded 2 ,seiceps dna morf
Simandrena A. (S.) wilkella propinqua (= dorsbta)
g ,nn nea,ofhairsngoniem ahRutiCTuTb e h rtsegmnroolf eobt dna eht,rettal
Taeniandrena
aim e,deatcuAt iltas cnniIo g if o etoelh n choc d in etoieltmielZhwuoahlwonxtocec aciniS ni
.aisA nretsaE ni noitubirtsid sti no yduts erutuf eht ot derrefed( si ,yduts tneserp eht
206
0. TADAUCHI & XU HUAN-LI
.71 suep ytluCohtiw a ln aaenitidadetumct ni;ugepncmoai lpsssecorp fo murbal etanigrameni
eht ;elddim laduac airbmif ;hsinworb sniev dna amgits elap ;hsiwoll eayivalsoguYot
naeporuE .aissuR . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . susterai nekflA
- suepylC htiw a naidem lanidutign oeltatcnup myilesrevsna rdteneergahs ;ecaps ssecorpfo
murbal ;hs i s;na nn wmdie aognedeir iavltrblt oaba s;ngnmde iii ulegftadhrocdtanimme
,napaJ anihC dna aeroK htuos( & .)lartnec . . . . . . . . . . . . . . . . . . . . . kerriae Hirashima
K.ey to the Palaearctic males of the subgenus Simandrena
.1 lamosateM agret ;suonigurref laedoporp erusolcne deneergahs lla ;revo lasrod ecaffo
muedoporp ,etalless;et htiw wollah s,serutcnup ton ;denehg uloarmosatem agret 4-2htiw
,esned etelpmoc riah ;sdnab suepylc htiw suovluf ;sriah laepylcarpus aera ,etatcnupton
.denehguor . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . transitoria .F ztiwaroM
- lamosateM agret ;kcal blaedo peorrupsolcn eesogur ta tsael no lasab ;noitrop lasrod ecaffo
muedoporp ;denehguor lamosatem agret 4-2 tuohtiw etelpmoc riah .sdnab . . . . . . . . 2
.2 rallegalF tnemges 1 regnol naht ro sa gnol sa tnemges .2 . . . . . . . . . . . . . . . . . . . . . . 3
- rallegalF tnemges 1 retrohs naht tnemges .2 . . . . . . . . . . . . . . . . . . . . , . . . . . . . . . . . . . . 6
.3 suepylC htiw a naid elmanidu teitgantoc lnyulpsmriev sdneanretergahs ;ecaps suepylchtiw
;yllare tsadl nra ibeash nyeldetare dho tm 4ia-wg2 rleatmosa t;esmr ieath iehsw n,eydrevlis
lamosatem mugret 1 yrev ylkaew etallesset htiw ,llams ,esned tcnitsid.serutcnup
. . . . . . . . . . . . . . . . . . . . . . . . kerriae Hirashima
- suepylC tuohtiw a naidem laniduttgnol etatcnupmi ;ecaps suepylc tuohtiw esned ;sriah
lamosatem agret 4-2 htiw ,esraps ylroop depoleved riah .sdnab . . . . . . . . . . . . . . . . . . 4
.4 lamosat emMugret 1 yrev ylk aeetwallesse tht icwipocsorc iemni f;serutc nluapsrod ecaffo
. sss ur uhoi ;etcad spishe neywunt orlf euhi ucmhbttftugi aoceuwncndoaiuordppnoirp
. . . . . . . . . . . . . . . . congruens thcenkedeimhcS
- lamosateM mugret 1 htooms dna ,ynihs htiw tcnitsid .serutcnup . . . . . . . . . . . . . . . . . . . . . 5
.5 daeH htiw elap ot llud hsiwolley sriah ton dexim htiw hsikcalb ro ;hsinworbmutucsosem
ylkaew ot ylesned etallesset ;yllaidem roiretsop lanoisserped aera fo lamosatemagret
ynihs dna htooms htiw llams .serutcnup . . . . . . . . . . . . . . . . . . . . . . . combinata (Christ)
- daeH htiw elap sriah deximda htiw ynam hsinworb ot hsikcalb seno no ralucoarap,aera
.yll ah yi tdndmonieuoahmtmsusc s ;ox lasedaeetnnrmraeaegv
. . . . . . . . . . . . . . nippon ihcuadaT teamihsariH
.6 lamosateM mugret 1 ynihs ot yrev ylkaew ,etallesset htiw tcnitsid .serutcnup . . . . . . . . . . 7
- lamosateM mugret 1 ylkaew etallesset htiw etunim .serutcnup . . . . . . . . . . . . . . . . . 8
.7 laedopo rePrusol cynletcnitsid esogur no 4/3 fo lasab ;noit rloapmosatem agret tuohtiwriah
;sdnab laepylcarpus aera etallesset dna ,etatcnup tuohtiw ;ealugur suepylc htiwhsitihw
;sriah rewol ralucoarap aera etallesset htiw etagnole .serutcnup . . . . . . . . susterai Alfken
- laedopor Perusolcne esogur no lasab ;flah lamosatem agret 4-2 htiw ylroop depolevedriah
;sdnab laepylcarpus aera htiw ;ealugur suepylc htiw kcalb ;sriah rewol ralucoarapaera
weakly tessellate and shiny, with round punctures. . . . . . . . . . . . . . . rhypara Perez
.8 laedoporP erusolcne esogur no tuoba lasab ;flah suepylc ylkaew ot ylesned etallessethtiw
wollahs ;serutcnup sriah no suepylc elap ot suocsuf . . . . . . . . . . . . . . . . . dorsata (Kirby)
REVISION OF SUBGENUS SZMANDRENA OF NRETSAEAISA 207
- laedoporP erusolcne yltcnitsid esogur lla revo no 4/3 fo lasab .noitrop . . . . . . . . . . . . . . 9
.9 lacipabuS nigram fo mutonorp ;eritne lamosatem munrets 7 htiw lacipa sebol;worran
munrets 8 htiw lacipa ebol ;rednels latineg eluspac htiw lasrod sebol fo etixoconogton
;detcejorp ,rellams htgnel tuoba 8 .mm . . . . . . . . . . . . . yamato ihcuadaT teamihsariH
lacipabu Snigram fo mut o;neotrapn ilgarmaomseat emmunrets 7 htiw lacipa sebol,redaorb
;dednuor munrets 8 htiw lacipa ebol daorb dna denediw ;yllacipa latineg eluspachtiw
lsaesbr oofldo etixoconsog ; yd ,lere cethugatdrrgo aenrtldepuolomba 9.mm
. . . . . . . . . . . . . . . . . . . . . aevoficapo amihsariH
1. Andrena (Simandrena) combinata (Christ)
.giF()1
Apis combinutu.]ynam reG ;elamef[ 781 .p ,.tcesnI .grutaN ,1971 ,tsirhC
Andrenu ulbiburbis ,kcnehcS ,3581 .bJ .reV .ktaN uassaN :9 431 ;elam[ .W ;]ynamreG,ekcmaW
,7691 ,soE :34.842
Andrenu combinutu: ,thcenkedeimhcS ,4881-2881 .dipA ,.poruE ,I .p 177 elamef[ & ;]ela.mE
trehkcotS ni ,thcenkedeimhcS ,0391 .myH -droN dnu .lettiM ,saporue .p 239,]elamef[
389 ;]elam[ ,uW ,2891 stcesnI fo ,gnaziX.293
Andrenu suhlbergi F. ,ztiwaroM ,8881 .roH .toS .tne .ssoR :22 142 elamef[ & ;elamelddiM
;]airebiS ,ekcmaW ,7691 ,soE :34 ,081.603
Andrena mehelyi ,nekflA ,6391 .fforeV .hcs t.userDebU-.loK .suM ,nemerB .I :3 083;elamef[
;]yragnuH ,e:kcmaW 19 67E,o s4,3 2:0 32,8 7.
Andrenu (Simundrenu) combinutu: ,ekcnraW ,8691 .meM .tsE .suM .looZ .vinU,arbmioC
:)703( ;,8k8uyn h,c7y7s9O1 an u,aiFniarkU 21 :)5(.932
Distrjbution:;) g, nnsaouzioig mX egorRnn uao:gittyjusUnAei w X.hantnariiohrnCe(b isoruE
.airebiS elddiM ;aissuR naeporuE ot eporuE elddiM
Floral records: kuynhcysO )7791( dedrocer gniwollof srewolf morf naeporuE:aissuR
Potentillu urgentutu, Trifolium prutense, Mutricuriu inodoru, Burbureu vulgaris, Melitotus
oficinulis, Inulu britunicu, Picris hierucioides.
Flight records::el a.Ma iss unRaepor unyEil ueJt aoeltn uyJlr a,eani hneCin ud Ji:melameF
.ais snuaRepo rynu liEeu tJeoa ntlyulJrae
Specimens examined selameF( = ,F selaM = :)M ]anihC[ gnaijniX rugyUsuomonotuA
:noigeR ,F2 ukuW ,daoR nahsnaiT htron ,epols ,m0061 .11 .iv 0691 gnaW(.)gnoy-uhS We
s n:e s mnwsiao aceldeple poonosrfisumlEaaxe ,Fl , .s,r ide.an irla .bnal0crVr 0ee,5Vz.,tt7ifwS
20-28. vi. 1959 (J.E. & .R ; ).nBosneB ,Fl ,stliW .t G,nywde B.62 .iiv 9691 ;.)Kd(rahciuG ,Fl
sessoB ,seplA .42 .iv 8491 .FMP( ;)ffeohreV ,Ml ,engoduoD ,talraS ,ecnarF .02 .vi ;5591 ,Ml
,egaraV ,selliesraM ,ecnarF .22 .vi.0691
Parasite: Unknown.
Remarks: ssiehiTc espash dneede rbmo ocer elferdpdoirMu Eo t ealidrdei bMdilnSaartnec
.anihC ehT elamef fo siht seiceps si evitcnitsid ni gnivah eht suepylc ,deneergahs lluddna
derevoc htiw esned elap ,sriah eh tlamos amtuegmret 1 ,dehsi lyonpihs dna htooms htiwesned
ag rleatmos a a tefelehoramtanoisser pleadc ie,phsatht deirwut c dneyeunbtpoara pseesrutcnup
ynihs dna htoom shti w ,essenreudt ce nhlutapid ie gtdyaeplppa h esht-ta Vildwleessisa,eratera
d neaht laduac airbm.infedlog
208 IH C.U0ADAT & IL -UNXAUH
.ps
2. Andrena (Simandrena) tiunshana, nov.
.sgiF( ,2)41
LB l-5.01 2.1 ,mm LW 5.8-0.8 mm .)3=n(
Female.
Color: elbidnaM htiw lacipa driht ro ssel ;tnecsefur gniw senarbmem ,enilayhyletaredom
elap ,hsinworb sniev dna amgitsoretp ;hsinworb lamosatem agret tneculsnart hsiddernworb
;yllacipa isratitsid hsidder ;nworb 1aibit srups.suoecatset
Pubescence: sriaH no d ahe sheilw ao ; ploslltua eoiayvce lav hufoks ffrieanvdwoobradbna
elap hsiwolley ;woleb ,mutucso smeumlletucs dna mutonatem htiw trohs ,)~005-003( suovluf
;sriah laedopo raplucibroc yrev llew depoleved htiw lasrod egnirf fo ,suovluf etiuq gnol-006(
,)p0001 ,esned llew degnarra ;sriah lasrod ecaf fo muedoporp htiw ,esraps suovluf;sriah
laret n,ashuccocrotl f;llams 1aibit apocs y ld e elt gla shner;utawesoirdrvwroilamauhf lT feoe rf
sriah ,yllaidem htiw trohs ,)~053-002( tcerebus sriah ;yllaretal lT htiw etihw riahegnirf
,yllaretaltsop T2 htiw a riah dnab yldaorb detpurret n,iyllaidem T3.4 htiw ylraen etelpmocot
etelpmoc riah ;sdnab laduac airbmif ;nedlog ~,IS htiw ,trohs ,esned tcere sriah ,yllaidem 5-ZS
htiw ,trohs ,tcerebus suovluf sriah ,ylroiretsop htiw lacipa egnirf yrev.trohs
Structure: :daeH WH/LH = 30.Ok68.0 (0.83-0.88, .)3=n 1annetnA epacs = tsrif 4rallegalf
;stnemges 1LF = ,3+2hE 2LF > ,3LF 2LF dna 3LF redaorb naht .gnol seyE htiw rennisnigram
gnig ryelvtnhogci ldsr a.wsoetlbid nraamlaM ecaps .raen islsecorP fo murbal etaredo mni,ezis
tuoba 5.1 semit sa daorb sa ,lgandoilozepa rott de d,nyulolr aecciapfar u.syni hssuepylCetiuq
eetc eaad s tpnhr , csate,yeni v nnduw ws,aiipaonlihwmra adsirrx neatehimnvtdniuow tcaignol
PP ,wollahs ,llams htiw dehsilop ,htooms suepylc ;yllacipa ecaps etatcnupmi ~02@ dna ,l&I LPC
= 0.I .mm rewoL ralucoarap aera ylraen htooms htiw ,gnorts esned .PP laicaF aevof,daorb
,w oLlVlFa h=s 4. 1,m mWVF = 5 . x0.de me t tlmd.r loa nwenantaueVinneoieorrvGraf > ey e
W E= :,WeGlif onnreipes ,5.0:8.0 ra eynni hysldao rebcafr u,se yrea ednedwo rPecPtun ihmt iw
eye dna ,euqapo deneer g.ayhlsroiretsop DCO:DOP:DO0 = .3.0:5.0:7.0 Mesosoma: Pronotum
rehtar ,depoleved ylthgils erutus laretal htiw mutonorp ;PP wollahs ,esraps htiw ,etallesset ylkaew
,gnol tuohtiw laretal ,ealugur lacipabus nigram fo mutonorp ton .etanigramemutucsoseM
yletaredom ,etallesset htiw wollahs PP030~ dna SI = ;5.1-1 mulletucs ynihs ni eht.elddim
elrauesd oollp lcodenrewePnife dyb a d,eenreuhtgu usnoor lasab 312 ddnea;nyelelragcaihpsa
lasrod ecaf fo muedop o;rdpeneh gruaolrucibroc aera yleni f.eta lmluemsestestipesyelMesned
d n;ighnire pyaltthg islussrati sda nb disnhuasrati sealbd dhi tm gi .ewwPLoPll ahh tseitwallesset
aibit .trohs Metasoma: WtM:WsM:W H= .6.3:3.3:2.3 Tl y le ltyaalslk alahbeets,wisewettunim
dnuor ~51-Ol@PP de rnt h ea a ythsSlolotrPI ltroiaPy yeamwp lylsess= lrksn,aeae3cvet-iwlpa
htiw yrev w oertraatncn uypnmiihs ecaps ta .secipa T2-4 y le ltyaalslk alahbeets,wisewettunim
esned ~02-Ol@PP dna SI = y hl,tl1oa ocdmi nspyaan inhhotsii twda nlolalnleasmsset -Ol@PP
20~ dna SI = ;3-l lacipa snoisserped fo agret llew .detacidni laidigyP etalp depahs-Vhtiw
.a erraalugna ilratnre tdneis iyalretare dh ot,mix wedpeadnuor 5-ZS ,esr ahp tseitwalles syeltk aew
wollahs.PP
Unknown.
Male.
Distribution:.)noigeR suomonotuA rug yU gnaijniX :tsewhtron( anihC
Floral record: toN.elbaliava
Flight record: :elameF dim.enuJ
NOISIVER FO SUNEGBUS SIMANDRENA FO NRETSAEAISA 209
Figs. 1-8: sdaeH ni latnorf weiv rof 8 seiceps fo tsaE naisA Andrena (Simandrena), 1:
combinata; 2: tianshana, .ps ;.von 3: koma; 4:yamato; 5: opacifovea; 6: austroinsularis;
sp. ;.von
7: wuae, 8: nippon.