Table Of ContentPROC. ENTOMOL. SOC. WASH.
98(2), 1996, pp. 264-266
A NEW SPECIES OF ECTENOPSIS (PARANOPSIS) (DIPTERA: TABANIDAE)
FROM NEW ZEALAND AND A KEY TO SPECIES OF THE
SUBGENUS PARANOPSIS
JOHN E BURGER
Department of Zoology, University of New Hampshire, Durham, NH 03824, U.S.A.
Abstract.(cid:8212)Ectenopsis (Paranopsis) mackerrasi, n. sp., from the North Island of New
Zealand is described and illustrated. It is compared to Ectenopsis lutulensis (Hutton), the
only other known species in this subgenus, and a key to the subgenus Paranopsis Mack-
erras is provided.
Key Words: Diptera, Tabanidae, Pangoniini, Ectenopsis (Paranopsis), Australasian Re-
gion, New Zealand, Key to Species
Mackerras (1955) described Paranopsis those of male slender, gradually tapered to
as a subgenus of Ectenopsis, based on Ec- blunt apex, about /% length of proboscis.
tenopsis lutulentus (Hutton). Only females Proboscis about equal to head height, stout,
were available for study. This paper de- with large, soft labella. Hind tibial spurs un-
scribes a second species in the subgenus usually large and powerful. Cerci as in Per-
Paranopsis, supplements the generic defi- isilvius Ferguson. Male genitalia similar to
nition provided by Mackerras, and includes those of other subgenera of Ectenopsis
a key to the species of Ectenopsis (Paran- (Mackerras 1955).
opsis). The subgenus Paranopsis 1s restrict-
ed to New Zealand. Ectenopsis (Paranopsis) mackerrasi
Burger, NEW SPECIES
Genus Ectenopsis Macquart, subgenus
(Figs. 1, 2)
Paranopsis Mackerras
Holotype male.(cid:8212)Length: 11.6 mm. Hol-
Paranopsis Mackerras, 1955: 473, fig. 16.
optic, eyes black, without color pattern, up-
Type-species: Apatolestes lutulentus Hut-
per % of eyes with facets enlarged, distinct-
ton, 1901. New Zealand.
ly separated from area of small facets be-
Eyes bare, without color pattern, those of low. Ocellar tubercle large, gray tomentose,
male with enlarged facets on upper %, dis- distinctly raised above level of eyes, ocelli
tinctly separated from area of smaller facets distinct. Frontal triangle yellowish gray to-
below. Ocellar tubercle raised and promi- mentose, bare. Frontoclypeus and genae
nent. Front wide, index 2, without frontal gray tomentose, white pilose. Beard white.
callus. Face convex. Third antennal seg- Antennae with scape and pedicel gray to-
ment with basal 4 flagellomeres widened mentose, pilosity yellowish on scape, mixed
and partially to mostly fused, apical four yellowish and black on pedicel, flagellum
flagellomeres clearly defined, apical flagel- reddish brown, basal four flagellomeres en-
lomere longer than preceding three flagel- larged, partially fused, apical four flagel-
lomeres combined. Maxillary palpi rather lomeres clearly defined, the apical one long
flattened, longer than shaft of proboscis, and slender, distinctly longer than preceding
VOLUME 98, NUMBER 2 265
:
Fig. 1. Holotype male of Ectenopsis (Paranopsis) mackerrasi, dorsal view. Scale bar = 4.0 mm.
three flagellomeres combined. Maxillary topleural lobes gray tomentose, white pilose
palpi gray tomentose, basal palpomere above, black pilose below. Pleuron gray to-
globular, apical palpomere slender, elon- mentose and white pilose. Legs with coxae
gate, gradually tapered to blunt apex, about gray tomentose, pale pilose, femora brown,
¥, length of proboscis, palpi mostly white darker black-brown subapically, tibiae
pilose, with mixed black and white hairs brown, becoming black-brown apically,
apically. Proboscis gray-brown, labellum hind tibial spurs very large, strongly curved
large, unsclerotized. apically, tarsi black-brown. Wings subhya-
Mesoscutum gray tomentose with a pair line with a grayish cast, veins dark brown,
of broad submedian and sublateral brown sc bare above and below, vein R, with spur.
stripes extending its entire length. Scutel- Abdominal tergites gray tomentose with
lum gray tomentose with broad median pale yellowish gray posterior margins and
brown marking, both mesoscutum and scu- distinct brown markings, tergite 1 with
tellum bearing long silky white hairs. No- broad submedian crescent-shaped mark-
266 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON
ler. In the collection of the California Acad-
emy of Sciences, San Francisco, U.S.A.
Etymology.(cid:8212)This species is named for
the late Ian M. Mackerras, whose studies
on the classification of Tabanidae and the
Australasian and Oceanian fauna laid the
foundation for the modern classification of
Tabanidae and the diversity of Tabanidae of
New Guinea, Australia, New Zealand and
the Pacific Islands.
Discussion.(cid:8212)Ectenopsis lutulentus 1s
known only from the South Island, al-
though Mackerras (1957) cites a question-
able record from Taranaki, in the south-
western part of the North Island. E. mack-
errasi is known only from the northern part
of the North Island. Although E. lutulensis
is known only from female specimens and
the species described here is known from a
single male specimen, I am confident that
Fig. 2. Head of Ectenopsis (Paranopsis) macker-
they represent distinct species because other
rast, holotype male, lateral view. Scale bar = 2.0 mm.
described species of Ectenopsis for which
both sexes are known show little sexual di-
morphism.
ings, tergites 2(cid:8212)6 with pair of large sub-
median subquadrate brown patches enclos- KEY TO SPECIES OF ECTENOPSIS
ing a median gray tomentose triangular (PARANOPSIS)
area, the brown markings largest on tergites
(cid:8212) Mesoscutum and abdomen both without distinct
2-3, progressively reduced on posterior
brown markings ....... lutulentus (Hutton) (2)
segments, tergites predominantly pale pi-
(cid:8212) Mesoscutum bearing distinct brown stripes and
lose. Sternites gray tomentose, white pilose. abdomen bearing large brown submedian quad-
Ninth tergite (epandrium) large, undivided, rate markings .......... mackerrasi n. sp. (6)
shield-shaped. Aedeagus gradually tapered
LITERATURE CITED
posteriorly, as in Ectenopsis (Perisilvius).
Hutton, E W. 1901. Synopsis of the Diptera Brachy-
Gonocoxites relatively slender, gradually
cera of New Zealand. Transactions of the New
tapered posteriorly. Gonostyli bifid, outer
Zealand Institute 33: 1(cid:8212)95.
lobe broadly rounded apically, inner lobe Mackerras, I. M. 1955. The classification and distri-
more slender apically and more strongly bution of Tabanidae (Diptera) II. History: mor-
phology: classification: subfamily Pangoniinae.
pointed, as in Ectenopsis (Ectenopsis).
Australian Journal of Zoology 3: 439-511.
Holotype male-(cid:8212)NEW ZEALAND:
Mackerras, I. M. 1957. Tabanidae (Diptera) of New
North Isl., Maropiu Distr., Omamari Beach Zealand. Transactions of the Royal Society of
N. of Dargaville, 5. III. 1969, James E. Tob- New Zealand 84: 581-610.
