Table Of ContentElytra,Tokyo,36(2):349-355,November22,2008
ANewParanchodemus(Coleoptera,Carabidae)
fromCentralJapan
Seiji MORITA
Higashi-gotanda5-19-7,Shinagawa-ku,Tokyo,141-0022Japan,
Naoki ToDA
Hirako2-24-16, Minami-ku,Nagoya-shi, Aichi,457-0001Japan
and
Noboru KANIE
Nishibora-cho117,Seto-shi, Aichi,489-0837Japan
Abstract Anewplatyninecarabidbeetleisdescribedfrom theChubuDistrict,
CentralJapan,underthenameofParatlchodemusishiguroi. Itisrelatedtop.ca11eides
(BATES),butdiffersfromitmainlyinthecolorationandtheshapeofpronotum.
Paranchodemus(HABU, 1978)waserectedasasubgenusofthegenusAnchodemus
for A. ca11eldes BATES (1883, p 256). Later, LIEBHEER (1989) studied the type
specimen,mainlythefemalegenitaliaandregardedHABU'ssubgenusasan independent
genus. At thesametime, headdedonenewspecies fromChina.
In1998,MoRvANerectedthegenusKeriserzh foraTibetanspecies,K thibetanus,
and added its new subspecies, K t. sichuanensis. However, thisgenus wasa junior
synonymof thegenusParanchodemus (BoUsQUET, 2003). Up to thepresent, three
speciesandonesubspeciesofthegenusParanchodemushavebeenknownfromEastAsia
(BOUSQUET,2003).
Inthispaper,wearegoingtodescribeanewspeciesasafourthrepresentativeof
thegenus.
The abbreviations used herein are as follows: L - body length, measured from
apicalmarginofclypeustoapicesofelytra;HW-greatestwidthofhead;PW -greatest
widthofpronotum;PL- lengthofpronotum,measuredalongthemid-line;PA - width
of pronota1apex; PB-width of pronota1base; EW-greatest width of elytra; EL-
greatest length of elytra; FL- length of metafemur; ML- length of metatrochanter;
TL - length ofmetatarsus; TLI- length ofsegment 1ofmetatarsus; TLV- length of
350 SeijiMORITA,NaokiToDAandNoboruKANIE
clawsegmentofmetatarsus;M-arithmeticmean;NSMT- Nationa1Museum of Na-
tureandScience, Tokyo.
Beforegoingfurther,wewishtoexpressourdeepgratitudetothefollowingfriends
forsupplyinguswithimportantmaterial:Dr.YoshiroKURosA,Messrs.MasakiIsHI-
GUR0 and Kentarou TOYOSHIMA.
Paranchodemusishiguroi MORITA,ToDAet KANIE,sp nov
[Japanesename:Hakusan-ruri-hirata-gomimushi]
(Figs.1-17)
Diagnosis. Bodyofmoderatesize;dorsalsideusuallywithbrilliantbluelustre;sides
ofpronotumweaklyarcuateinfront,andconvergentposteriadorweaklysinuatebefore
hindangles;basalpart ofelytrawide.
Description. L:8.71-11.43mm. Bodyofmoderatesize,black;headandpronotum
withgreenish,purplishorbrilliantbluishlustre; elytrawithbrilliantbluishorgreenish
lustre, rarelywithpurplish lustre;ventralsideblack withweak bluish lustre; coxae
blackishbrown;appendagesblack.
Head moderately convex; eyes large and convex; frontal furrows deep, almost
parallel fromclypea1setatoclypea1suture,andthendivergentposteriadandreaching
thelevelofbasal5/6ofeye;microsculpturenotsharplyimpressedthoughconsistingof
isodiametricmeshes;genaeoblique;mentumtoothwideandweaklybifidatthetip;apex
of labrumweaklytomoderatelyemarginate; antennaefiliform, reachingthemiddleof
elytraoralittlebeforethat level; relativelengthsofantennalsegmentsas follows: -
I : II: III : IV: V:VI : XI ≒1: 0.63: 1.64: 1.47: 1.41:1.27: 1.08.
Pronotumnarrow,weaklyconvexandwidestataboutapical2/7;sideswidelyand
weakly arcuate in front, graduallyconvergent towardshind angles or very weakly
sinuateatbasal2/7;apicalanglesweaklyproducedandmoderatelyroundedatthetips;
apexalmoststraight;medianlinedeepandnotreachingapexnorbase;anteriormarginal
setaesituatedat about basal7/20;anterior transverseimpressionveryshallow;basal
foveae rather shallow, with several coarsepunctures and irregular wrinkles, each
extendinganteriorly;hindanglesright,andwithobtuselyroundedtips;PW/HW1.01-
1.10(M1.06) in ,1.04-1.13(M1.09) in早,PW/PL1.07-1.20(M1.15) in 1.10-
,
120(M1.17)in早,PW/PAt29-1.39(MI34)in1(1',1.30-1.40(MI35)in早,PW/PB
1.30-1.38 (MI34) in(i7',1.30-1.42(MI35) in早,PA/PB0.97-1.03(M1.00) in
,
0.96-1.03(M1.00) in早;discwithfinetransversewrinkles;microsculptureconsistingof
fine transverse meshes.
Elytraelongate,moderatelyconvex,andwithwidebasalpart;EW/PW193-2.03
(M I 99) in , 1.91-2.10 (M2.01) in早,EL/EW157-1.71 (M I 65) in 1.55-1.74
,
(M I65) in早;shouldersprominentandrounded;sidesalmoststraight infront,usually
very weaklyemarginateat about basal 7/20of elytra, and then widely and weakly
arcuate posteriad; preapica1sinuation wide and shallow; apical parts narrow and
NewParanchodemusfromCentralJapan 351
、,
f
t
4,
3
Figs. 1-4. paranchodemus lshiguroi MORITA,ToDAet KANIE,sp nov.,ShOWin9ColOu「Va「iatiOn
- 1,3,4,o'l;2,・♀.
352 SeijiMORITA,NaokiToDAandNoboruKANIE
5 6
、、、、、
9
(_f
f
一
、。一.一一。。,e「。'。。,,0C・...0,・・''.-〇一・-・) '一一一.~一一一'。
Figs 5-9. Paranchodemuslshiguroi MORITA,ToDAet KANIE, sp nov. - 5, Labrum; 6, prono-
tum;7,apicalpartoftheleftelytron;8,same,showingvariationofporesandstria7;9,sternites
V-VI in♀. Scale:1.0mm for6-8:0.5mm for5and9.
moderatelyproduced;striaedeep,almostsmoothorweaklypunctate;scutellarstricto
longandsituatedon interval I;basalporesituatedonbaseof interval I, andadjoining
stria1oronthestria, rarelyon theinterval;microsculptureratherstrongly impressed
andcomposedofpolygonalorwidemeshes; intervalsweaklyconvexandimpunctate;
marginal seriesofumbilicatepores22-28innumber; interval m usuallywith three
pores,sometimesfour, rarelyfiveortwo;thefirstporeadjoiningstria3andsituatedat
about basal t/5-l/3of elytra, rarely lacking; thesecond one adjoiningstria 2and
situatedat about middle; thethirdoneadjoiningstria2andsituatedat aboutbasal
13/20-7/10ofelytra;apicalpartof intervalVII usuallywith fourorfiveporesoneach
side(cf. LIEBHEER,1989,fig 9),rarelywiththreeorsixpores,andpositionofthepores
variable,usuallyadjoiningstria7oronthestria,sometimeson theinterval,rarelyclose
tostria4;apexofstria2sometimeswithaweakporeoneachside(cf. LIEBHEER, l989,
fig.10);stria7rarelyratherclearly impressedtowardstheanastomosisofstriae1and
2;apexofstria1withaweakporeoneachside;elytra1epipleurongraduallynarrowed
NewParanchodemusfromCentralJapan 353
し
、、、一
,?(
' ノ
一/
- / l l
/,J / ノ ノ
ノ
l 5
,
a
L l
,7
b
, '
c
Figs.10-17. Paranchodemus ishiguroi MORITA, ToDAet KANIE,sp nov. - l0, Aedeagus, left
lateralview;11,apicalpartofaedeagus,dorso-apica1view;12,rightparamere, leftlateralview;
l3, leftparamere, left lateral view; 14, genital segment, ventral view; 15, femalereproductive
tract,withfoldedapicalpartofspermatheca;16,spermathecaandspermathecalglandinanother
specimen;17,stylus. Scales:a,05mmfor i5-17;b,05mmfor te-13;c, 1.0mmfor i4.
354 SeijiMORITA,NaokiToDAandNoboruKANIE
apicad; innerplicaindistinct.
Sternites(I-V)usuallywithtwoor threeshortsetaeoneachside,andrarelywith
oneadditionalseta;analsternite(VI) trapezoidal, bisetosein , quadrisetosein早,
apicalhalfdepressed,andwithshortandirregularwrinkles.
Legslongandslender;metatrochanterveryshort,usuallywithaseta;ML/FL0.29-
0.32(M0.30) in 0.29-0.33(M0.31)in早;metafemoraeachwiththreeor foursetae
,
in ventralview, rarelytwo; twoproximalsegmentsofmeso-andmetatarsieachwith
inner andoutersulci;clawsegmentsofmeso- and metatarsi without setae on ventral
side;segment4ofmeso-andmetatarsishallowlybilobed;TLI/TLV0.95-1.13(M1.02)
in 0.98-1.20(M1.06) in早.
,
Genitalsegmentelongatedovateandwithaveryshort handle.
Aedeagusofmoderatesize,andwiththinbasalpart; basalorificelarge;viewed
laterally,aedeagusweaklyarcuate;apical lobenarrowlyproducedandsimplyrounded
at thetipindorsal view; rightparamereelongateandsmall; left paramerelargeand
wide.
Spermathecabroad, withcurvedapex;spermathecal glandelongateand curved;
apicalstyliwide,withtwonematiformsetaeatapicalpit-likedepression (LIEBHEER,
1989,p 6),withtwolongspinesatbasalpartandsixshortspinesatapicalpart.
Typeseries. Holotype:フ(1'(NSMT),14-VI-2008,S.MORITAleg. Paratypes:1早,
30-VII-2006,M.IsHIGUR0leg;2(l'l(11',1早,16-VIII-2006,M.IsHIGUR0leg;1早,26-
Vm-2006, K. ToYosHIMAleg; 4 , 1早,2-IX-2006, K. ToYosHIMAleg; 11
,
6早早, 11-VIII-2007, N. KANIEleg;14 , 7早早,12-VIII-2007, N. ToDAleg;50
279-?,14-VI-2008,S.MORITA&N.ToDAleg;14 99-9,28-VI-2008,N.
, ,
ToDAleg;9 ,17早早,26-VII-2008,N.ToDAleg.
Locality.Riv.0gamigo-gawa,Shokawa-cho,Takayama-shi,GifuPrefecture,Chubu
District,CentralJapan.
N1otes. Thestandardratiosofbodypartsshowninthedescriptivepartarethoseof
ten malesand ten females.
ThisnewspeciesiscloselyalliedtoParanchodemusca11eides(BATES,1883,p 256).
It is,however,distinguished fromthelatter bythefollowingpoints:1) colorationof
dorsalside;2)shapeofpronotum;and3)wideelytra1basalparts. [in3 and 3早早
ofP.ca11eidesfromNiigataPref.,PW/HW1.07-1.09(M1.08) incフ,', 1.08-1.11 (M
1.09) in早;PW/PL1.16-1.20(M1.18) in ,1.16-1.21(M1.19) in早;PW/PA t32-
1.38(M I36) in ,1.33-1.37(MI35) in早;PW/PB130-1.35(MI33) in 1.28-
,
1.36(M I 31)in ;PA/PB0.95-1.03(M0.98)in ,0.95-1.00(M0.98) in早;EW/PW
1.88-1.98(MI93) in ,1.93-2.00(M I96) in早;EL/EW l64-1.76(MI69) in
,
1.63-1.71(MI67) in早;TLI/TLV1.06-1.24(M1.13) in ,1.08-1.12(M1.04) in早;
ML/FL 0.32-0.33 (MO32) in , 0.31-0.32 (MO32) in早; TL/HW133-1.36 (M
1.35) in ,1.28-1.36(MI31) in早; relativelengthsofantennalsegmentsasfollows:
- I:II:III:IV := 1:0.54:1.55:1.31.]
NewParanchodem1lsfromCentral Japan 355
要 約
森田誠司・戸田尚希・蟹江 昇: 中部地方から採集されたヒラタゴミムシの1新種. - 岐阜
県から採集された標本を基に, ハクサンルリヒラタゴミムシParanchodemus ishiguro1 MORITA,
ToDAetKANIEを記載した. 本種は, 中部~東北地方から知られるオオアオグロヒラタゴミムシ
p.ca11eides(BATES)に近縁であるが,背面につよい青色ないし緑青光沢をもっことのほか,前胸
背板の形, 幅広い上翅の基部などの外部形態から, 容易に識別される.
References
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GeorgeLEWIS,madeduringhissecondvisit,fromFebruary,1880,toSeptember,1881.T「ansent・SOC・
Londot1,1883:205-290,pi t3.
BoUsQUET,Y.,2003. TribePlatyniniBoNELLI,1810. In LOBL,1., &A.SMETANA(edS・),Cat・Palaea「C・t
Coleopt.,1:449-469. ApolloBooks,Stenstrup.
HABU,A.,1978.carabidae:Platynini(Insecta:Coleoptera).FalunaJaponica.vii十i 447PP.,36PIS・Kei9aku
Publ.,Tokyo.
LIEBHEER, J.K., lg8g. Reviewof thepalaearcticgenusParanchodemusHABU(Coleoptera:Carabidae:
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Elytra,Tokyo,36(2):355-356,November22,2008
) DescrjptionoftheMaleofO taop agus(fzdacrzus c
OcHIet KoN(Coleoptera,Scarabaeidae)
TeruoOcHI1),MasahiroKoN2)andAkiraKASHIZAKI3)
1)Kohudai5-21-6,Toyono-cho,Toyono-gun,Osaka,563-0104Japan
2)schoolofEnvjronmenta1Science,TheUniversityofShigaPrefectu「e,
Hassaka-cho2500,Hikone,Shiga,522-8533Japan
3)Njshj6-2-10-1202,Kita31,Kita-ku,Sapporo,Hokkaido,001-0031Japan
Onthophagus(Indachorius)cheyiwasdescribedbyOcHIandKON(2006)basedonth「ee
femalespecjmenscollected fromGomantong, nearSandakan,Sabah, Malaysia. Recently, the
thirdauthorhascollectedthemaleofthisspeciesfromSepi1oknearthetypelocality・Web「lefty
descrjbeithereinandbasedonthisspecimenillustrateitsmale9enitalia・