Table Of ContentZOOSYSTEMATICA ROSSICA, 22(1): 141–149 25 JUNE 2013
Types of Scathophagidae (Diptera) at the Zoological Institute
of Russian Academy of Sciences, St Petersburg
Типы двукрылых семейства Scathophagidae (Diptera)
в Зоологическом институте РАН, С.-Петербург
A.L. OZEROV* & M.G. KRIVOSHEINA
А.Л. ОЗЕРОВ*, М.Г. КРИВОШЕИНА
A.L. Ozerov, Zoological Museum, Lomonosov Moscow State University, Bol’shaya Nikitskaya 6, Moscow 125009,
Russia. E-mail: [email protected]. *Corresponding author.
M.G. Krivosheina, A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, 119071 Moscow,
Russia. E-mail: [email protected]
The data are summarized on the type specimens (holotypes, lectotypes, paratypes, paralecto-
types) of 19 species in the family Scathophagidae (Diptera) kept at the Zoological Institute
of Russian Academy of Sciences, St Petersburg. The lectotypes are designated for three spe-
cies, Scatophaga arctica Becker, 1897, S. septentrionalis Becker, 1897, and Scopeuma longinqua
Becker, 1915. Acerocnema obscuripes Becker, 1915 is transferred to the genus Nanna Strobl,
1894. Three new synonymies are established: Scathophaga multisetosa (Holmgren, 1883) =
Scatophaga arctica Becker, 1897, syn. nov.; Nanna obscuripes (Becker, 1915) = N. indota-
tum Engelmark, 1999, syn. nov.; Allomyella frigida (Holmgren, 1883) = Microprosopa va-
ritibia Becker, 1897, syn. nov. Both syntypes of Ceratinostoma nudiseta Becker, 1907 are not
Scathophagidae but Muscidae possibly belonging to Spilogona Schnabl, 1911. A redescription
of Allomyella portenkoi (Stackelberg, 1952) is given.
Приведены данные по типовым экземплярам (голотипы, лектотипы, паратипы, пара-
лектотипы) 19 видов двукрылых из семейства Scathophagidae, хранящихся в коллекции
Зоологического института РАН (С.-Петербург). Обозначены лектотипы трёх видов:
Scatophaga arctica Becker, 1897, S. septentrionalis Becker, 1897 и Scopeuma longinqua Becker,
1915. Acerocnema obscuripes Becker, 1915 помещен в род Nanna Strobl, 1894. Установлены
3 новых синонима: Scathophaga multisetosa (Holmgren, 1883) = Scatophaga arctica Becker,
1897, syn. nov.; Nanna obscuripes (Becker, 1915) = N. indotatum Engelmark, 1999, syn. nov.;
Allomyella frigida (Holmgren, 1883) = Microprosopa varitibia Becker, 1897, syn. nov. Оба
изученных синтипа Ceratinostoma nudiseta Becker, 1907 относятся не к Scathophagidae,
а к Muscidae, возможно, к роду Spilogona Schnabl, 1911. Дано переописание Allomyella
portenkoi (Stackelberg, 1952).
Key words: dung-flies, taxonomy, type specimens, collection, Zoological Institute of Russian
Academy of Sciences, Diptera, Scathophagidae, new synonyms
Ключевые слова: мухи-скатофагиды, таксономия, типовые экземпляры, коллекция, Зо-
ологический институт РАН, Diptera, Scathophagidae, новые синонимы
INTRODUCTION present paper reviews the type specimens of
Scathophagidae kept in ZIN. We have ex-
The Zoological Institute of Russian amined the types of 19 species described by
Academy of Sciences in St Petersburg (ZIN) the Russian dipterologists J.A. Portschin-
possesses the richest Diptera collection in sky (1848–1916), A.A. Stackelberg (1897–
Russia. Scathophagidae is a small family 1975) and K.B. Gorodkov (1932–2001),
of about 350 valid species worldwide. The who worked in St Petersburg (Leningrad).
© 2013 Zoological Institute, Russian Academy of Scienсes
142 A.L. OZEROV & M.G. KRIVOSHEINA. TYPES OF SCATHOPHAGIDAE
The famous German dipterologist Theodor Lectotype. Male: “Kurlyk, Baingol vost.
Becker (1840–1928) described several new Tsaydam. RobKozlov 21.V.95” [Kurlyk, eastern
species of Scathophagidae from northern Baingol, Caidam, 21.V.1895, coll. Roborowsky
et Kozlov; in Russian] (designated by Ozerov &
Russia in 1897–1914. We examined the
Krivosheina, 2011: 105).
types of eleven Scathophagidae species
Paralectotypes. One male, labelled same as
described by Becker, which were found in
lectotype. Five females: (1) “r Orogyn Syrtyn
ZIN.
yu Nan’shan, Gobi, RobKozlov 3–20.V.95” [in
The type specimens described by Russian], (2) “oz. Iche – r. Orogyn s. Tsaydam
Portschinsky, Stackelberg and Becker were ts. Gobi RobKozlov 3 VII 95” [in Russian], (3)
labelled properly; hence we recognized “r Orogyn Syrtyn yu Nan’shan, Gobi, RobKo-
them without difficulties. A part of types zlov 3–20 VII 95” [in Russian], (4) “Kara Tyube,
of the species described by Gorodkov were na zap ot Khami, Gobi. RobKozlov 5.IX.95” [in
properly labelled as well. At the same time, Russian], (5) “r. Bomyn (Ichegan) sv. Tsaydam
Gobi RobKozlov VI 95” [in Russian].
the new species described and the lecto-
Notes. The species was described from
types designated by him in 1974 (Goro-
eight specimens collected in China; seven
dkov, 1974) were lacking the respective
of them, two males and five females, are
labels. Luckily, all of these specimens were
kept in ZIN. The eighth specimen, male, is
found in a separate drawer. The first author
in Museum für Naturkunde der Humboldt-
labelled as the holotypes of Cordilura sagit-
Universität, Berlin (Ozerov & Krivosheina,
tifera Gorodkov, 1974 and C. zaitzevi Goro-
2011: 106).
dkov, 1974 the only male specimen in each
The lectotype is pinned; one of middle
type series with the label consistent with
legs is glued to the geographical label and
that of the holotype (Gorodkov, 1974) and
lacking tarsomeres 2–4, the other middle
with the apex of abdomen cut and placed in
leg is broken, tibia and tarsi of the left hind
a vial with glycerol. Apparently, Gorodkov
leg are broken, as well as tarsomeres 2–4 of
(1974) illustrated the male genitalia from
the right hind leg; the abdomen is cut and
these specimens.
placed in a vial with glycerol mounted on
All taxa are listed in alphabetic order of
the same pin with the specimen.
their species names. The labels are separated
The male paralectotype in ZIN is
from each other by oblique stroke. The label
Scathophaga mihalyii (Šifner, 1975). The
data are put in quotation marks, with trans-
male paralectotype in Berlin (with labels:
lation and comments given in brackets.
“oz. Iche – r. Orogyn s. Tsaydam ts. Gobi
The terminology follows McAlpine
RobKozlov 3 VII 95” [in Russian] / “Gobi.
(1981) and Cumming et al. (2009). The fol-
Asien 51896. VII”/ “albidihirta Beck.”/
lowing abbreviations are used: a, anterior; d,
“ZMU Berl”/ “Scatophaga albidohirta Beck.
dorsal; p, posterior; v, ventral; combinations
SYNTYPE labelled 1954 by J.R. Vock-
of these four abbreviations, all used for leg
eroth”) is conspecific with the lectotype.
chaetotaxy; ALO, A.L. Ozerov.
This specimen was examined by ALO in
2012 and labelled as a paralectotype.
ANNOTATED LIST OF TYPES
Three female paralectotypes (labels
1–3) are conspecific with the lectotype.
Order DIPTERA
Two female paralectotypes (labels 4–5)
Family SCATHOPHAGIDAE are Scathophaga species, but surely are not
S. albidohirta.
albidohirta Becker, 1907b: 254 (Sca- Identity. A good species of the genus
tophaga). Scathophaga Meigen, 1803. It was rede-
Type locality: “Kurlyk” [Lake Hurleg scribed and illustrated by Ozerov & Krivo-
Hu, 37.28°N 96.89°E] (China). sheina (2011).
© 2013 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 22(1): 141–149
A.L. OZEROV & M.G. KRIVOSHEINA. TYPES OF SCATHOPHAGIDAE 143
amplipennis Portschinsky, 1887: 199 “41956”/ “Scatophaga arctica Beck. SYN-
(Scatophaga). TYPE labelled 1954 by J.R.Vockeroth”/
Type locality: “[high in the upper reach- “Scatophaga apicalis Curt. Det. 1962
es of Yellow [=Hawang] River in northeast- J.R. Vockeroth”) is kept in Museum für
ern Tibet]” (China). Naturkunde der Humboldt-Universität,
Lectotype. Male: “Scatophaga amplipennis” Berlin. This specimen was examined by
[handwritten by Portschinsky] (designated by ALO in 2012 and labelled as a paralecto-
Gorodkov, 1967: 446). type; it is conspecific with Scathophaga
Paralectotype. Male, labelled same as lecto- multisetosa (Holmgren, 1883).
type. Identity. Scathophaga arctica syn. nov.
Notes. The species was described from is a junior synonym of S. multisetosa (Hol-
both sexes. Only two males were found in mgren, 1883).
ZIN, both without geographical labels. The
infumatum Becker, 1907b: 256 (Conio-
lectotype (designated by Gorodkov (1967),
sternum).
but labelled by ALO) is pinned, in very good
Type locality: “Sogon-gomba” [upper
condition. The second male was labelled by
reaches of the Yangtze River, ca. 33.6°N
ALO as a paralectotype.
96.4°E] (China).
This species was transferred to the genus
Holotype. Male: “Sogon-gomba – r Ichu, ver
Scatomyza Fallén, 1810 (Ozerov & Krivo-
Goluboy, Kozlov, kon VII.[19]00” [in Russian]/
sheina, 2011).
“Tibet, Kozl. VII.00”/ “infumatum Beck.”/ “5”
Identity. Similar to Scatomyza mag-
[three labels handwritten by T. Becker].
nipennis (Portschinsky, 1887) (see below), Notes. The species was described from a
probably is a junior synonym of this species. single male. The holotype is pinned, in very
good condition; left wing is partially torn;
arctica Becker, 1897: 398 (Scatophaga).
the abdomen is cut and placed in a vial with
Type locality: Malye Karmakuly, Nova-
glycerol on the same pin with the specimen.
ya Zemlya (Russia).
Identity. A good species of Scathophaga.
Lectotype. Male: “Mal. Karmakuly, Novaya
It was redescribed and illustrated by Oz-
Zemlya. 31.VII.[18]96 Jacobson” [in Russian] /
“Scatophaga arctica Bec[ker]” [handwritten by erov & Krivosheina (2011).
T. Becker] / “Scatophaga apicalis Curtis male fulvisetis Becker, 1915: 64 (Pleuro-
Gorodkov det. [19]73” (designated here). chaeta).
Paralectotype. Female: “stoyanka u g. Cher- Type locality: “aus dem Polaren Ural”
nyshova, Nov. Zem. 4.VIII.96 Jacobson” [in Rus-
(Russia).
sian].
Holotype. Male: dark gold circle/ “Popyzrny
Notes. The species was described from
Ural, s. Tobol’s. gub. FZaitzev 6 VII [18]09” [in
the specimens of both sexes collected in No-
Russian] / “det. Becker” [printed] “Pleurochaeta
vaya Zemlya. There are two specimens in
fulvisetis B.” [handwritten by T. Becker].
ZIN, male and female. The male is in very
Notes. The species was described from a
good condition; its abdomen is dissected
single male. The holotype is pinned, in ex-
and placed in a vial with glycerol on the
cellent condition.
same pin with the specimen. This specimen Identity. A junior synonym of Pleuro-
has been labelled by ALO and is designated chaetella simplicipes (Becker, 1900).
here as a lectotype of Scatophaga arctica.
longinqua Becker, 1915: 66 (Scopeuma).
The lectotype is conspecific with Scathoph-
Type locality: “Tundra des Fl. Kara”
aga multisetosa (Holmgren, 1883).
[tundra in environs of the Kara River, the
The female has been labelled by ALO as
border of Nenetskii Autonomous District
a paralectotype.
and Tyumen Province] (Russia).
One male paralectotype (with labels:
Lectotype. Male: “Karskaya tundra s. Tobol’s.
“Novaja Semlja mons Tschernyschew G. gub. FZaitzev 12 VII [19]09” [in Russian] (des-
Jacobson 5.VIII.96”/ “arctica Becker”/ ignated here).
© 2013 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 22(1): 141–149
144 A.L. OZEROV & M.G. KRIVOSHEINA. TYPES OF SCATHOPHAGIDAE
Paralectotypes. Two females: (1) dark gold Beck.” [handwritten by T. Becker]. One
circle/ “Karskaya tundra s. Tobol’s. gub. FZait- female: “Ins. Kotelnyi A. Wolossowitch 10–
sev 12 VII 09” [in Russian]/ “det. Becker” 25 VI. 1901”/ “Polyarnaya ekspeditsiya”
[printed] “Scatophaga longinqua B” [handwrit-
[in Russian]/ “det. Becker” [printed] “Ce-
ten by T. Becker], (2) “Karskaya tundra s. To-
ratinostoma nudiseta Beck.” [handwritten
bol’s. gub. FZaitsev 12 VII 09” [in Russian].
by T. Becker].
Notes. The species was described from
Notes. This species was described from
three specimens of both sexes. All are in
three exemplars of both sexes; two syn-
ZIN, a male and two females. The male has
types, male and female, were found in ZIN.
been labelled by ALO and is designated
In the male, the left postpedicel, tarsi of
here as a lectotype. The lectotype is pinned,
right fore leg, tibia and tarsi of left mid leg,
in good condition; the abdomen is dissected
right mid leg and both hind legs are broken,
and placed in a vial with glycerol on the
otherwise in good condition. The female is
same pin. The two females were labelled by
in good condition.
ALO as paralectotypes. All three type speci-
Identity. Both syntypes are not Scatho-
mens are Scathophaga cordylurina (Hol-
phagidae but Muscidae probabl y belong-
mgren, 1883).
ing to Spilogona Schnabl, 1911 (A.C. Pont,
Identity. A junior synonym of Scathoph-
pers. comm.).
aga cordylurina (Holmgren, 1883).
obscuripes Becker, 1915: 64 (Aceroc-
magnipennis Portschinsky, 1887: 198
nema).
(Scathophaga).
Type locality: “Tundra des Fl. Kara”
Type locality: “San Kul’” [Sonkol Lake,
[tundra in environs of the Kara River, the
3160 m] (Kyrgyzstan).
border of Nenetskii Autonomous District
Lectotype. Male: “San Kul’” [in Russian]/
and Tyumen Province] (Russia).
“Scatophaga magnipennis” [handwritten by
Holotype. Male: dark gold circle/ “Karskaya
Portschinsky] (designated by Gorodkov, 1967:
tundra s. Tobol’s. gub. FZaitzev 12 VII [19]09”
445).
[in Russian]/ “det. Becker” [printed] “Aceroc-
Paralectotype. Female, labelled same as lec-
nema obscuripes B.” [handwritten by T. Becker].
totype.
Notes. The species was described from
Notes. The species was described from
a single male. The holotype is pinned, in
both sexes. Two specimens, male and female,
very good condition. It belongs to the ge-
are kept in ZIN. The male lectotype (des-
nus Nanna Strobl, 1894, not to Acerocnema
ignated by Gorodkov (1967), but labelled
Becker, 1894, and was repeatedly described
by ALO) and the female paralectotype are
by Engelmark (1999) as Nanna indotatum.
pinned, in very good condition. The female
Identity. Nanna obscuripes (Becker,
was labelled by ALO as a paralectotype.
1915), comb. nov., with a junior synonym
Identity. A good species of the genus
N. indotatum Engelmark, 1999, syn. nov.
Scatomyza.
orbitalis Becker, 1915: 65 (Scopeuma).
nudiseta Becker, 1907a: 4 (Ceratinosto-
Type locality: “Pe-mal” [hilly area be-
ma).
tween the northernmost parts of the Polar
Type locality: “Neu-Sibirische Inseln, In-
Urals and the Arctic sea coast] (Russia).
sel Kotelnyi, Südufer beim Stan-Michailow”
Holotype. Female: “Pe-Mal, s. Tobol’s. gub.
and “West-Taimyr-Halbinsel, Walter-Bai,
FZaitzev 15 VII [19]09” [in Russian]/ “det.
Ostufer am Cap Kriwoj” (Russia). Becker” [printed] “Scatophaga orbitalis B.”
Syntypes. One male: “Taimyr occ. Birulja [handwritten by T. Becker].
22 VII– 4 VIII 901”/ “mukhi poym. bl. pal- Notes. The species was described from
atki (stoyanka XI)” [in Russian]/ “Polyar- a single female. The holotype is pinned, in
naya ekspeditsiya” [in Russian]/ “det. very good condition. It is conspecific with
Becker” [printed] “Ceratinostoma nudiseta Scathophaga nigripalpis (Becker, 1907)
© 2013 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 22(1): 141–149
A.L. OZEROV & M.G. KRIVOSHEINA. TYPES OF SCATHOPHAGIDAE 145
and was correctly recognized by Gorodkov Holotype. Male: dark gold circle/ “O-v
(1986: 33). Vrangelya. Aternon Rodzhers 27.VI.939 Porten-
Identity. A junior synonym of Scathoph- ko” [in Russian]/ “Stackelberg det.” [printed]
“Microprosopa portenkoi sp.n. male Typ.” [hand-
aga nigripalpis (Becker, 1907).
written by A. Stackelberg].
paradoxopyga Stackelberg, 1952: 405 Notes. The species was described from a
(Acerocnema). single male. The holotype is minute pinned,
Type locality: Matochkin Shar (73.3877° in very good condition, only pedicel and
N, 55.2144°E), Novaya Zemlya (Russia). postpedicel of both antennae are lost; the
Holotype. Male: “Nov. Zemlya, Matochkin abdomen is cut and placed in a vial with
Shar xxx Vakulenko 11/VII.925” [in Russian; glycerol on the same pin with the specimen.
xxx = illegible]/ “Stackelberg det.” [printed] This species has been incorrectly recog-
“Acerocnema paradoxopyga sp.n. Typ. 41” [hand- nized by Hackman (1956: 24, 58, Figs 58,
written by A. Stackelberg].
128) and later by Gorodkov (1970: 455),
Paratype. Male: “O-v Vrangelya. Aternon
therefore its redescription and the figures of
Rodzhers 27.VI.939 Portenko” [in Russian].
the holotype sternites 4 and 5 and the geni-
Notes. The species was described from
talia are given below. Vockeroth (1965: 835)
three males taken from Novaya Zemlya (ho-
recognized this species correctly.
lotype and paratype) and Wrangel Island
Identity. A good species of the genus Al-
(paratype). We have examined the holotype
lomyella Malloch, 1923.
and the male paratype from Wrangel I.; the
Redescription (Figs 1–6). Male (holo-
other paratype could not be found. The ex-
type). Length of body 4.2 mm; length of
amined types are in good condition.
wing 3.2 mm.
Identity. A good species of the genus Ac-
Head. Frons black, reddish yellow in
erocnema Becker, 1894.
anterior part only, greyish microtrichose.
perfecta Becker, 1907a: 3 (Scatophaga). Ocellar triangle and parafacial black, with
Type locality: “Lena-Mündung, Charau- greyish microtrichia. Face blackish, yel-
lachgebirge, Cap Elijdep”, Arctic coast of lowish along margin of mouth only, greyish
Yakutia (Russia). microtrichose. Gena yellow. Postcranium
Holotype. Male: “Prijansky Tundra z-w. black, greyish microtrichose, with black se-
Lena und Jana Brusnew 8, 9 VI 1902” [tundra tae in upper half and whitish hairs in lower
between the rivers Lena and Yana, leg. Brusnev, part. Head with 3 orbitals, 3–4 frontals, 1
8–9.VI.1902]/ “na tsvetakh” [in Russian]/ “det. ocellar, 1 postocellar, 1 inner vertical, and 1
Becker” [printed] “Scatophaga perfecta Beck.”
outer vertical setae; 1 pair of strong vibris-
[handwritten by T. Becker].
sae and 2 pairs of short subvibrissae. Scapus
Notes. The species was described from a
and pedicel black, both postpedicels miss-
single male. The holotype is in good condi-
ing. Palpus yellow, with moderately long
tion; left fore leg glued on carton card; end
yellowish setae.
of abdomen cut and placed in a vial with
Thorax black, densely microtrichose.
glycerol on the same pin with the specimen.
Scutum with 2 postpronotal, 2 notopleural,
It is conspecific with Scathophaga apicalis
2+2 supraalar, 1+2 intraalar, 2 postalar, and
(Curtis in Ross, 1835) and was correctly
(4–5)+(3–4) dorsocentral setae; anterior
recognized by Gorodkov (1986: 30).
presutural directed to head; acrostichals
Identity. A junior synonym of Scatho-
long and thin, in two rows. Proepisternum
phaga apicalis.
covered with hair-like setulae in middle
portenkoi Stackelberg, 1952: 406 (Mi- part, with 1 seta near upper margin. Pro-
croprosopa). epimeron with 1 seta. Anepisternum cov-
Type locality: Aternon-Rogers (70.9832° ered with setulae along dorsal margin and
N, 178.4166°E), Wrangel Island (Russia). in posterior part only, with 2–3 setae near
© 2013 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 22(1): 141–149
146 A.L. OZEROV & M.G. KRIVOSHEINA. TYPES OF SCATHOPHAGIDAE
posterior margin. Katepisternum covered from the type locality, the abdomen was cut
with setulae and with 1 seta in posterodor- and placed by ALO in a vial with glycerol
sal corner. Anepimeron with several hairs at on the same pin with the specimen; in one
middle. Postmetacoxal bridge absent. Scu- of the female paratypes from the type local-
tellum with 2 basal and 2 apical setae, and 2 ity, the abdomen is broken. Two male para-
discal setulae. types (from Mongolia and from environs of
Legs greyish microtrichose. Coxae, tro- Irkutsk) are with abdominal apex cut and
chanters and femora of all legs black; tibiae glued on plastic rectangle on the same pin
and tarsi of all legs yellow. Foretibia with with the specimen.
rows of short spines ventrally, 1 p and 1 d Identity. A good species of Cordilura Fal-
at middle, and preapical p and d. Midfemur lén, 1810, correctly recognized by recent
with preapical p and pd. Midtibia with 1 ad authors.
and 1 p at middle and a ring of apical setae. septentrionalis Becker, 1897: 397 (Sca-
Hind femur with a row of ad and a row of tophaga).
av. Hind tibia with 2 ad, 2 pd, 1 av, and 1 Type locality: Mt. Chernyshev, Novaya
preapical d. Zemlya (Russia).
Wing tinged with brownish. Veins Lectotype (designated here). Male: “stoyanka
brown; crossveins not darkened. Calypters, u g. Chernysheva, Nov. Zem. 5.VIII.[18]96 Jacob-
including margins, and halteres yellowish. son” [in Russian]/ “Scatophaga septentrionalis
Abdomen black, densely microtrichose, Becker” [handwritten by T. Becker]/ “Scatophaga
covered with yellowish hairs. Male sternites varipes (Holmgren) male Gorodkov det. 1972”.
4 and 5 as in Figs 1–4. Epandrium, cerci and Paralectotypes. Two males, with same geo-
surstyli as in Figs 5, 6. graphical labels as in lectotype.
Distribution. Russia: Wrangel I.; USA: Notes. The species was described from
Alaska (Vockeroth, 1965: 835). five specimens of both sexes. Three male
syntypes were found in ZIN. The male in
sagittifera Gorodkov, 1974: 388 (Cordi-
good condition has been labelled by ALO
lura).
and is designated here as a lectotype of Sca-
Type locality: 6 km E of Kyren, 700 m
tophaga septentrionalis. Two other males
(ca. 51.6770°N, 102.2146°E), Khargun, Bu-
were labelled by ALO as paralectotypes.
ryat ia (Russia).
The other two syntypes (male and fe-
Holotype. Male: “Khargun, 6 km O Kyren, male) were kept in Museum für Naturkunde
Sayany, osok. boloto, Gorodkov 11 VII 965” der Humboldt-Universität, Berlin. The
[Khargun, 6 km E of Kyren, Sayan Mountains,
male has the labels: “Novaja Semlja mons
sedge bog, leg. Gorodkov 11.VII.1965; in Rus-
Tschernyschewi G. Jacobson 5.VIII.96”/
sian]/ “700 m”.
“41955”/ “Scatophaga septentrionalis Beck.
Paratypes. Six males and eight females, la-
belled as holotype. One male: “Okr.Irkutska. SYNTYPE labelled 1954 by J.R. Vock-
Malashedova 4.VI.911” [In Russian]/ “sagit- eroth”/ “Scopeuma varipes Holm. Det.
tif.” [handwritten by Gorodkov]. One male: “oz. 1954 J.R. Vockeroth”. The female has the la-
Tsagan, 100v. k W ot Urgi, Mong[olia]. Kozlov bels: “Novaja Semlja mons Tschernyschewi
28–29.VI.924” [in Russian]. G. Jacobson 5.VIII.96”/ “septentrionalis
Notes. The holotype (labelled by ALO) Beck.”/ “41955”/ “Scatophaga septentrio-
is pinned, in very good condition; the ab- nalis Beck. SYNTYPE labelled 1954 by J.R.
domen was cut and placed in a vial with Vockeroth”/ “Scopeuma varipes Holm. Det.
glycerol on the same pin with the specimen 1954 J.R. Vockeroth”. Both specimens were
(apparently, by Gorodkov). The 16 para- examined by ALO in 2012 and labelled as
types of this species mentioned by Goro- paralectotypes.
dkov (1974: 388) were found in ZIN, in Identity. A junior synonym of Scathoph-
good condition; they were labelled by ALO aga varipes (Holmgren, 1883), correctly
as paratypes. In one of the male paratypes recognized by recent authors.
© 2013 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 22(1): 141–149
A.L. OZEROV & M.G. KRIVOSHEINA. TYPES OF SCATHOPHAGIDAE 147
Figs 1–6. Allomyella portenkoi (Stackelberg), male (holotype): 1, sternite IV; 2, sternite V, left side;
3, sternite V, dorsal view; 4, sternite V, right side; 5, epandrium, cerci and surstyli, lateral view; 6,
epandrium, cerci and surstyli, dorsal view.
Fig. 7. Abdominal sternites of Microprosopa varitibia Becker, female (holotype).
© 2013 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 22(1): 141–149
148 A.L. OZEROV & M.G. KRIVOSHEINA. TYPES OF SCATHOPHAGIDAE
sibirica Gorodkov, 1974: 388 (Cordi- “Microprosopa varitibia B[ecker]” [handwritten
lura). by T. Becker].
Type locality: 6 km E of Kyren, 700 m Notes. The species was described from
(ca. 51.6770°N, 102.2146°E), Khargun, a single specimen, incorrectly indicated as
Buryatia (Russia). male. The holotype is pinned; left foreleg,
Holotype. Male: “Khargun, 6 km O Kyren, right mid- and hind legs are missing; the
Sayany, osok. boloto, Gorodkov 11 VII 965 “ abdomen is cut and placed in a vial with
[Khargun, 6 km E of Kyren, Sayan Mountains, glycerol on the same pin with the specimen.
sedge bog, leg. Gorodkov 11.VII.1965; in Rus-
Sternites (Fig. 7) are conspecific with those
sian]/ “700 m”.
of Allomyella frigida (Holmgren, 1883).
Notes. The species was described from a
Identity. Microprosopa varitibia syn.
single male. The holotype is pinned, in very
nov. is a junior synonym of Allomyella frigi-
good condition; the end of abdomen is cut
da (Holmgren, 1883).
and placed in a vial with glycerol on the
same pin with the specimen. zaitzevi Gorodkov, 1974: 386 (Cordi-
Identity. A good species of Cordilura. lura).
stackelbergi Gorodkov, 1967: 448 (Oke- Type locality: Songino, 1300 m, Central
niella). Aymag (Mongolia).
Holotype. Male: “Mongoliya, Tsentral-
Type locality: “[North of Mondy, Tun-
nyy Aymak, Songino, poyma i stepnye sklony,
kins kie Mountains (2500 m), East Sayan]”
Zaitzev, 3–4.IX.969” [in Russian]/ “zaitzevi sp.
(Russia).
n. Gorodkov det.”.
Holotype. Male: “okr. Mondy, Sayany,
Paratypes. Three females, with geographi-
Tunkinskie gol’tsy, Gorodkov 22 VII 965” [in
cal labels as in holotype. Three males: “Mon-
Russian]/ “r. Khulugayma, 2500 m, golets, bolo-
goliya, Tsentr. Aymak, Songino, Nartshuk,
to” [in Russian]/ “Holotypus ♂ Okeniella stack-
7.VII.970” [in Russian]. Two males: “Mon-
elbergi Gorodkov, 1967”.
goliya, Tsentr. Aymak, r. Kerulen, 40 km ZYuZ
Paratypes. Two males and two females, with
Bayan-Obo, Kerzhner, 28.VII.971” [in Rus-
geographical labels same as in holotype. Nine
sian]. Two females: “Khargun, 6 km O Kyren,
males and eight females: “okr. Mondy, Sayany,
Sayany, osok. boloto, Gorodkov 11 VII 965” [in
Tunkinskie gol’tsy, Gorodkov 28 VII 965” [in
Russian]/ “700 m”. One female: “Nikol’skaya
Russian]/ “Verkh r. Ikhe-Ukhgun’, gol’tsovyj
slobodka, Namsk. ul., Herz 26 VI [19]01” [in
poyas, boloto, 2200 m” [in Russian].
Russian]. One male: “Malokrasnoyarsk. na
Notes. The holotype is minute pinned,
Irtyshe Semip, Vereshchagin 27 VII 926” [in
in excellent condition. There are 21 more
Russian]. Two males and one female: “Yakutsk,
specimens of this species in ZIN (11 males,
Moskvin 30.VII.927” [in Russian]. One male:
10 females), all in good condition and la-
“Novopokrovskoe, pr. b. Amgi Yakut., Pripuzov
belled by Gorodkov as paratypes. In one 22.VII.928” [in Russian].
of the male paratypes collected on 28 July Notes. The holotype (labelled by ALO)
1965, the abdomen is cut and placed in a is pinned, in very good condition; the end
vial with glycerol on the same pin with the of abdomen is cut and placed in a vial with
specimen. glycerol on the same pin with the specimen.
Identity. A good species of the genus All 16 specimens of this species mentioned
Okeniella Hendel, 1907. by Gorodkov (1974) (9 males, 7 females)
were found in ZIN in good condition and
varitibia Becker, 1897: 400 (Micropro-
labelled by ALO as paratypes. In two of the
sopa).
Type locality: “Malija-Karmakuly, Nowa- male paratypes (those collected by Kerzh-
ja-Semlja” [Malye Karmakuly (72.371409°N ner and Moskvin), the end of abdomen is
52.741517°E), Novaya Zemlya] (Russia). cut and glued on plastic rectangle on the
Holotype. Female: “Mal. Karmakuly, Novaya same pin with the specimen.
Zemlya. 16.VII.[18]96 Jacobson” [in Russian] / Identity. A good species of Cordilura.
© 2013 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 22(1): 141–149
A.L. OZEROV & M.G. KRIVOSHEINA. TYPES OF SCATHOPHAGIDAE 149
ACKNOWLEDGEMENTS scription of two new species; Nanna indota-
tum and Cochliarium sibiricum. Entomologisk
We wish to thank Dr. Andrey Przhiboro (St Tidskrift, 120(4): 157–167.
Petersburg) for the assistance in getting the Gorodkov K.B. 1967. New data on high altitude
types and the information about some of them. Scatophagidae (Diptera) from the Asiatic
Our thanks to Dr. Emilia Nartshuk, Dr. Ol’ga part of the USSR. Entomologicheskoe Oboz-
Ovchinnikova and Mrs. Galina Suleymanova renie, 46(2): 445–449. (In Russian).
(all are from St Petersburg) for the help during Gorodkov K.B. 1970. Family Scathophagidae
our work in ZIN with the type material. We are (Cordyluridae, Scatomyzidae, Scopeumati-
grateful to Dr. James O’Hara (Ottawa) for the dae). In: Stackelberg A.A. & Nar tshuk E.P.
help in the identification of the specimens of A. (eds.). A key to the insects of the European
portenkoi determined by J. Vockeroth. part of the USSR, 5(2): 440–458. Leningrad:
Nauka. (In Russian).
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Received June 13, 2012 / Accepted June 5, 2013
© 2013 Zoological Institute, Russian Academy of Scienсes, Zoosystematica Rossica 22(1): 141–149
