The Herbaceous Angiosperm Flora of Mahoning - Knowledge Bank

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THE HERBACEOUS ANGIOSPERM FLORA OF
MAHONING COUNTY, OHIO1' 2
CARL F. CHUEY
Department of Biology, Youngstown State University, Youngstown, Ohio 44503
ABSTRACT
A checklist of the herbaceous angiosperms of Mahoning County, Ohio, based upon
field observations and collections made during the summer of 1967 and upon the holdings
of four major Ohio herbaria are presented. The list represents 73 families, 284 genera,
and 527 species, of which 101 species, from the author's collection, are new county records.
INTRODUCTION
Mahoning County (fig. 1) is located in northeastern Ohio and adjoins Trumbull
County on the north, Portage and Stark Counties on the west, Columbiana County
on the south, and the Commonwealth of Pennsylvania on the east. It has an
area of 422 square miles. The maximum elevation of the county is 1314 feet,
found in Green Township, and the lowest elevation is 795 feet, found in Poland
Township, giving a maximum relief of 519 feet. The county was established in
1846.
Mahoning County is located in the Glaciated Allegheny Plateau of the Appa-
lachian Plateau Province of the Appalachian Highlands (Braun, 1950), and is
underlain by sandstone bedrock formed during the Mississippian and Pennsyl-
vanian Periods (Bownocker, 1947). Later, during the Pleistocene Epoch, the
county was invaded by both the Illinoian and Wisconsin glaciers (White, 1953).
The county is drained by the Mahoning River and its tributaries, with the
exception of a very small section of the northeastern corner which is drained by
Yankee Creek. Both the Mahoning River and Yankee Creek flow into the Ohio
River. The drainage pattern of the Mahoning River is unusual in that, due to
glacial blocking, the course changes from northwest to northeast and finally to
southeast, so that, because of the political boundaries of the county, it enters and
leaves the county four times. Similar glacial action also caused several other
rivers in the area to have their original directions of flow reversed (Stout, Ver Steeg,
and Lamb, 1943). Because of impoundments in the drainage system of the
Mahoning River, the county has seven major bodies of water (fig. 1).
Since the time of its early settlement, the northeastern corner of the county
has remained primarily farmland. In recent years, however, most of the county
has undergone rapid changes, owing to strip mining, to urbanization, and to in-
creased population. The population of the county has increased from 300,480
in I960, of which 212,192 were in cities (U.S. Bureau of Census, 1963), to an
estimated population of 318,048 in 1967, of which 214,566 were in cities (Ohio
Division of Economic Research, 1967). If this trend continues, the entire county
should be almost completely urbanized in the near future. The need for making
a floristic study of the area before any future changes occur, therefore, is obvious.
One such change is the plan of the U.S. Army Corps of Engineers, announced in
1968, to make a reservoir out of "Egypt Swamp".
The purpose of this study, therefore, was to make a survey of the entire
flora of Mahoning County. This county lies entirely within the Maple-Beech
Forest Region of Ohio (Braun, 1950). Although reference has been made to the
county in previous state or regional floristics works (Braun, 1961 and 1967;
Schaffner, 1914 and 1932), no intensive work has been done on the flora of Mahon-
xPresented as a thesis in partial fulfillment of the requirements for the Degree Master of
Science, Ohio University, 1969.
2Manuscript received April 20, 1971.
THE OHIO JOURNAL OF SCIENCE 72(1): 30, January, 1972.
No. 1 HERBACEOUS FLORA OF MAHONING COUNTY 31
ing County. Notes on the flora of Mill Creek Park are presented in a visitor's
guide to the various features of the Park (Galaida, 1941), but no herbarium speci-
mens are available for the species listed. Certain areas of the county were ex-
cluded from this present study: Mill Creek Park, an area of over 2000 acres,
because it is heavily planted with exotic species; Mahoning Valley Sanitary Dis-
trict, a watershed area of over 2500 acres, because this area is planted mainly with
pine trees; and all residential areas, which often contain exotic species.
A 512-acre section of the county, the northern third of what is locally known as
"Egypt Swamp", was chosen by the author for intensive study. The area is
located in Green Township (fig. 1) and consists of a broad valley with slopes
averaging 20 degrees (maximum of 45 degrees) facing east and west. The area is
bounded on the north by State Route 165 and on the south by Pine Lake Road.
Mill Creek Park Egypt Swamp
Man. Vol. Sant. Dist.
lakes freeways
•—k Mahoning River and its direction of flow
FIGURE 1. Map of Mahoning County showing location of townships and "Egypt Swamp".
Ecologically "Egypt Swamp" contains many different communities: oldfield,
beech-maple woods, and roadside communities on all topographical features;
powerline right-of-way and swamp communities on the bottomland and terraces;
and strip-mine vegetation only on the upland. Because of the large number of
habitats present, it is believed that this portion of "Egypt Swamp" is floristically
representative of the various habitats of the entire county.
METHODS OF COLLECTION AND IDENTIFICATION
The main study and collection period was from March until September, 1967.
During this time, weekly trips were made to "Egypt Swamp" and to other parts
of the county as time and weather permitted. Trips were made to all the town-
ships of the county, but only species that had not been previously taken in "Egypt
Swamp" were collected in the other townships. At least one specimen of each
species collected is deposited in the Bartley Herbarium of Ohio University (BHO),
and the remainder are in the Herbarium at Youngstown State University (YUO).
32 CARL F. CHUEY Vol. 72
Two manuals were used for identification: Gleason (1963) and Braun (1967).
Specimens which presented particular difficulties in determination were checked
against specimens in the Bartley Herbarium of Ohio University.
As a result of this study, a list of all plants observed or known in Mahoning
County was prepared, and is presented below. The order of arrangement of the
families in this check list follows Gleason (1963) for the dicotyledons and Braun
(1967) for the monocotyledons. The tribes, genera, and species in the check list
are arranged in alphabetical order under the families. In the listing of the taxa,
the following information is given at the end of the scientific name for each speci-
men collected by the author: the collection number, the place of collection by
township or topographical feature within "Egypt Swamp" (bottomland—"Bot";
terrace—"Ter"; upland—"Up"), the date of collection, the location of the speci-
men, and the degree of abundance. For those specimens collected by other col-
lectors, the order of information is as follows: the collector, the collection number
if any, the place of collection by township if known, the date of collection, and the
location of the specimen. The location of such specimens is indicated by the in-
ternational abbreviations for the following herbaria (Lanjouw and Stafleu, 1964):
BHO (Bartley Herbarium of Ohio University), KE (Herbarium of Kent State
University), OS (Herbarium of The Ohio State University), and YUO (Herbarium
of Youngstown State University). All specimens were examined personally by
the author. The author's collection resulted in 101 new county records, which
are identified in the list by an asterisk.
The abundance of those species found in Mahoning County by the author is
indicated by the use of the following terms.
Rare—few scattered locations
Sparce—infrequently found
Common—frequently found
Abundant—large populations, regularly found
Species reported by others but not seen by the author carry no abundance labels.
Few specimens were collected in the county between 1880 and 1950 as indicated
by the dates. Most have been collected since 1955 and the author's collection
of 302 species is the most intensive work done to date.
CATALOGUE OF THE
HERBACEOUS ANGIOSPERM FLORA OF MAHONING COUNTY, OHIO
MONOCOTYLEDONEAE
TYPHACEAE
Typha angustifolia L. CFC669, Bot, July 1, 1967, BHO
CFC689, Beaver Twp., July 1, 1967, YUO
Common
Typha latifolia L. CFC670, Bot, July 1, 1967, BHO
CFC795, Ter, July 18, 1967, BHO
CFC705, Springfield Twp., July 18, 1967, BHO
Common
SPARGANIACEAE
Sparganium americanum Nutt. P. A. Bricker, Poland Twp., July 2, 1955, OS
Sparganium eurycarpum Engelm. CFC520, Bot, May 31, 1967, BHO & YUO
Abundant
ZOSTERACEAE
Potamogeton crispus L. Tom S. Cooperrider, #7060, Jackson Twp., July 19,
1960, KE
Potamogeton diversifolius Raf. Tom S. Cooperrider, #7015, Milton Twp., July 18,
1960, KE
ALISMATACEAE
Alisma subcordatum Raf. CFC880, Bot, Aug. 5, 1967, BHO
CFC860, Jackson Twp., Aug. 1, 1967, YUO
Common
No. 1 HERBACEOUS FLORA OF MAHONING COUNTY 33
Sagittaria latifolia Willd. CFO866, Bot, Aug. 5, 1967, BHO Common CFC
*Sagittaria rigida Pursh CFC721, Jackson Twp., July 8, 1967, BHO Sparse
HYDROCHARITACEAE
Elodea nuttallii (Planch.) St. John Tom S. Cooperrider, #7039, Ellsworth Twp., July
19, 1960, KE
GRAMINEAE (POACEAE)
A grostideae
Agrostis alba L. CFC675, Ter, July 1, 1967, BHO Common
CFC703, Springfield Twp., July 8, 1967, YUO
Agrostis perennans (Walt.) Tuckerm. Allison W. Cusick, Beaver Twp., Aug. 12, 1962, OS
Agrostis scabra Willd. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Alopecurus aequalis Sobol. Allison W. Cusick, Springfield Twp., Aug. 12, 1962,
OS
Brachyelytrum erectum (Schreb.) Beauv. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Calamagrostis canadensis (Michx.)
Beauv. Ervin M. Herrick, Beaver Twp., June 12, 1963, OS
Cinna arundinacea L. CFC919, Ter, Aug. 17, 1967, BHO Common
Muhlenbergia frondosa (Poir.) Fern. CFC707, Springfield Twp., July 8, 1967, BHO
Common
Muhlenbergia schreberi Gmel. Ervin M. Herrick, Goshen Twp., Sept. 10, 1960, OS
* Muhlenbergia sobolifera (Muhl.) Trin. CFC835, Up, Aug. 1, 1967, BHO Sparse
Phleum pratense L. CFC621, Ter, June 22, 1967, BHO
CFC601, Jackson Twp., June 18, 1967, YUO
Abundant
Sporobolus asper (Michx.) Kunth Ervin M. Herrick, Berlin Twp., Sept. 4, 1962, OS
Sporobolus neglectus Nash Ervin M. Herrick, Canfield Twp., Oct. 3, 1961, OS
Sporobolus vaginiflorus (Torr.) Wood Ervin M. Herrick, Goshen Twp., Aug. 25, 1962, OS
Andropogoneae
Andropogon virginicus L. Ervin M. Herrick, Goshen Twp., Sept. 10, 1960, OS
Aveneae
Arrhenatherum elatius (L.) Mert. &
Koch Ervin M. Herrick, Boardman Twp., June 30, 1960,
OS
Danthonia compressa Aust. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
*Danthonia spicata (L.) Beauv. CFC578, Jackson Twp., June 18, 1967, BHO
Common
*Deschampsia flexuosa (L.) Trin. CFC554, Ter, June 8, 1967, BHO Sparse
Holcus lanatus L. Tom S. Cooperrider, #6969, Milton Twp., July 18,
1960, KE
Sphenopholis intermedia Rydb. CFC522, Bot, June 8, 1967, BHO
CFC539, Ter, June 8, 1967, BHO Common
Chlorideae
Eleusine indica (L.) Gaertn. Ervin M. Herrick, Poland Twp., Aug. 14, 1960, OS
Festuceae
Bromus inermis Leyss. CFC602, Jackson Twp., June 18, 1967, BHO & YUO
Sparse
*Bromus japonicus Thunb. CFC517, Bot. May 31, 1967, BHO
CFC547, Ter. June 8, 1967, BHO Common
Bromus purgans L. Ervin M. Herrick, Ellsworth Twp., June 13, 1963,
OS
Bromus secalinus L. Ervin M. Herrick, Coitsville Twp., June 30, 1960,
OS
Bromus tectorum L. Ervin M. Herrick, Poland Twp., June 31, 1960, OS
Cynosurus cristatus L. Ernest W. Vickers, #2378, Ellsworth Twp., June
25, 1909, KE
Dactylis glomerata L. CFC480, Bot, May 31, 1967, BHO & YUO
CFC542, Ter, June 8, 1967, BHO & YUO Common
Er agrostis hypnoides (Lam.) BSP. Allison W. Cusick, Beaver Twp., Sept. 15, 1960, OS
Eragrostis pectinacea (Michx.) Nees Allison W. Cusick, Springfield Twp., Aug. 12, 1962,
OS
Festuca obtusa Biehler CFC579, Jackson Twp., June 18, 1967, BHO
Common
34 CARL F. CHUEY Vol. 72
Glyceria canadensis (Michx.) Trin. Gareth E. Gilbert, Green Twp., July 22, 1955, OS
Glyceria striata (Lam.) Hitchc. CFC652, Bot, June 27, 1967, BHO Sparse
Poa annua L. Arthur S. Brooks, #3344, Jackson Twp., Oct. 22,
1962, OS
Poa compressa L. CFC913, Bot, Aug. 17, 1967, BHO Common
Poa cuspidata Nutt. CFC392, Jackson Twp., May 4, 1967, BHO & YUO
Sparse
*Poa palustris L. CFC481, Bot, May 31, 1967, BHO Common
CFC631, Ter, June 22, 1967, BHO
CFC696, Springfield Twp., July 8, 1967, YUO
Common
*Poa pratensis L. CGC481, Bot, May 31, 1967, BHO Common
Poa sylvestus Gray Ervin M. Herrick, Ellsworth Twp., June 12, 1963,
OS
Poa trivialis L. Ervin M. Herrick, Ellsworth Twp., June 12, 1963,
OS
Phragmites commnnis Trin. William C. Werner, Youngstown Twp., Aug.,
1887, OS
Triodia flava (L.) Smyth Ervin M. Herrick, Ellsworth Twp., Sept. 16, 1960,
OS
Hordeae
Agropyron repens (L.) Beauv. CFC632, Ter, June 22, 1967, BHO Common
*Elymus riparius Wieg. CFC848, Bot, Aug. 1, 1967, BHO Common
Elymus villosus Muhl. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Elymus virginicus L. CFC825, Bot, July 25, 1967, BHO Common
Hordeum jubatum L. CFC699, Springfield Twp., July 8, 1967, BHO &
YUO Common
Hystrix patula Moench Ervin M. Herrick, Ellsworth Twp., June 12, 1963,
OS
Loliutn perenne L. Ronald W. Hathaway, #339, Boardman Twp., May
29, 1968, YUO
Oryzeae
Leersia oryzoides (L.) Swartz Ervin M. Herrick, Coitsville Twp., Aug. 14, 1960,
OS
Leersia virginica Willd. CFC920, Bot, Aug. 17, 1967, BHO Common
Paniceae
Digitaria ischaemum (Schreb.) Muhl. Ervin M. Herrick, Milton Twp., Sept. 10, 1960, OS
Digitaria sanguinalis (L.) Scop. Ervin M. Herrick, Coitsville Twp., Aug. 14, 1960,
OS
Echinochloa crusgalli (L.) Beauv. CFC843, Bot, Aug. 1, 1967, BHO Abundant
Panicum capillare L. CFC929, Ter, Sept. 9, 1967, BHO Common
Panicum clandestinum L. CFC882, Bot, Aug. 5, 1967, BHO
CFC899, Up, Aug. 8, 1967, BHO Abundant
Panicum dichotomiflorum Michx. Arthur S. Brooks, #3347, Jackson Twp., Oct. 22,
1962, OS
Panicum dichotomum L. Ervin M. Herrick, Ellsworth Twp., June 12, 1963,
OS
Panicum lanuginosum Ell. CFC604, Jackson Twp., June 18,1967, BHO & YUO
CFC697, Springfield Twp., July 8, 1967, YUO &
BHO Common
Panicum latifolium L. CFC639, Ter, June 22, 1967, BHO Common
*Panicum linearijolium Scribn. CFC603, Jackson Twp., June 18, 1967, BHO
Sparse
Panicum stipitatum Nash Ervin M. Herrick, Berlin Twp., Sept. 4, 1962, OS
Panicum virgatum L. Ervin M. Herrick, Canfield Twp., Oct. 3, 1961, OS
Setaria glauca (L.) Beauv. CFC937, Bot. Sept. 9, 1967, BHO Common
Setaria viridis (L.) Beauv. Tom S. Cooperrider, #8432, Goshen Twp., Sept.
20, 1960, KE
Phalarideae
Anihoxanihum odoratum L. CFC535, Ter, June 8, 1967, BHO
CFC497, Up, May 31, 1967, BHO
CFC695, Springfield Twp., July 8, 1967, YUO
Common
Phalaris arundinacea L. Allison W. Cusick, Beaver Twp., Sept. 15, 1960, OS
No. 1 HERBACEOUS FLORA OF MAHONING COUNTY 35
CYPERACEAE
*Bulbostylis capillaris (L.) C. B. Clarke CFC856, Jackson Twp., Aug. 1, 1967, BHO & YUO
Abundant
* Car ex albursina Sheldon CFC822, Bot, July 25, 1967, BHO Common
Carex amphibola Steud. CFC589, Jackson Twp., June 18, 1967, BHO
Common
* Carex annectens Bickn. CFC592, Jackson Twp., June 18, 1967, BHO
Sparse
* Carex artitecta Mackenz. CFC459, Up, May 25, 1967, BHO & YUO Sparse
* Carex communis Bailey CFC394, Jackson Twp., May 4, 1967, BHO & YUO
Sparse
Carex convoluta Mackenz. E. L. Moseley, June 16, 1896, OS
Carex crinita Lam. CFC514, Bot, May 31, 1967, BHO & YUO
Abundant
Carex festucacea Schkuhr. Allison W. Cusick, Beaver Twp., June 11, 1961, OS
Carex grayii Carey CFC537, Ter, June 8, 1967, BHO Common
Carex hirsutella Mackenz. Ervin M. Herrick, Ellsworth Twp., June 12, 1963,
OS
Carex laevivaginata (Kukenth.) Mackenz Allison W. Cusick, Springfield Twp., June 11, 1961,
OS
Carex lupulina Muhl. P. A. Bricker, Poland Twp., July 2, 1955, OS
Carex lurida Wahlenberg CFC589, Jackson Twp., June 18, 1967, BHO
Common
*Carex molesta Mackenz. CFC576, Jackson Twp., June 18, 1967, BHO
Sparse
Carex normalis Mackenz. P. A. Bricker, Poland Twp., July 2, 1955, OS
Carex pensylvanica Lam. Ervin M. Herrick, Ellsworth Twp., May 8, 1963,
OS
Carex rosea Schkuhr. CFC492, Up, May 31, 1967, BHO
CFC536, Ter, June 8, 1967, BHO Common
Carex scoparia Schkuhr. William C. Werner, Youngstown Twp., July 2,
1887, OS
Carex squarrosa L. CFC590, Jackson Twp., June 18, 1967, BHO &
YUO Abundant
Carex stipata Muhl. CFC408, Bot, May 13, 1967, BHO & YUO
Common
Carex tetanica Schkuhr. Ronald W. Hathaway, #333, Boardman Twp., May
15, 1968, YUO
Carex tribuloides Wahlenb. Ervin M. Herrick, Coitsville Twp., June 30, 1960,
OS
*Cyperus esculentus L. CFC864, Bot, Aug. 4, 1967, BHO Common
Cyperus ferruginescens Boeckl. Allison W. Cusick, Beaver Twp., Aug. 27, 1961, OS
Cyperus filiculmis Vahl Tom S. Cooperrider, Jackson Twp., July 19, 1960,
KE
Cyperus strigosus L. CFC889, Ter. Aug. 8, 1967, BHO Abundant
Eleocharis acicularis (L.) R. & S. Allison W. Cusick, Springfield Twp., Sept. 15,
1960, OS
Eleocharis obtusa (Willd.) Schultes. CFC854, Bot, Aug. 1, 1967, BHO & YUO
CFC628, Ter, June 22, 1967, BHO Abundant
Scirpus atrovirens Willd. CFC639, Ter, June 22, 1967, BHO Abundant
*Scirpus cyperinus (L.) Kunth. CFC914, Bot, Aug. 17, 1967, BHO & YUO
Common
Scirpus polyphyllus Vahl CFC679, Ter, July 1, 1967, BHO Common
Scirpus validus Vahl var. creber Fern. CFC700, Springfield Twp., July 8, 1967, BHO
Abundant
Acorus calamus L. Ervin M. Herrick, Coitsville Twp., June 30, 1960,
OS
Arisaema atrorubens (Ait.) Blume. CFC461, Up, May 25, 1967, BHO & YUO
Common
Arisaema dracontium (L.) Schott. Ervin M. Herrick, Ellsworth Twp., June 12, 1963,
OS
Arisaema triphyllum (L.) Schott. Tom S. Cooperrider, #6840, Poland Twp., July 15,
1960, KE
*Peltandra virginica (L.) Schott & Endl. CFC724, Jackson Twp., July 8, 1967, BHO Sparse
Symplocarpus foetidus (L.) Nutt. CFC342, Bot, April 6, 1967, BHO Abundant
36 CARL F. CHUEY Vol. 72
LEMNACEAE
Lemna minor L. Tom S. Cooperrider, #6962, Milton Twp., July 18,
1960, KE
Wolfia columbiana Karst Tom S. Cooperrider, #7062, Jackson Twp., July 19,
1960, KE
COMMELINACEAE
* Tradescantia ohiensis Raf. CFC654, Ter, June 27, 1967, BHO Sparse
PONTEDE RIACEAE
Pontederia cor data L. Tom S. Cooperrider, #7016, Milton Twp., July 18,
1960, KE
JUNCACEAE
Juncus acuminatus Michx. CFC591, Jackson Twp., June 18, 1967, BHO
Common
Juncus bufonius L. Lindley Vickers, #2853, Youngstown Twp., Aug. 2,
1948, KE
Juncus effusus L. CFC629, Ter, June 22, 1967, BHO
CFC593, Jackson Twp., June 18, 1967, YUO
Abundant
Juncus tenuis Willd. CFC600, Jackson Twp., June 18, 1967, BHO &
YUO Common
Juncus platyphyllus (Wieg.) Fern. Allison W. Cusick, Springfield Twp., Sept. 15,
I960, OS
Luzula bulbosa (Wood) Rydb. Tom S. Cooperrider, #1834, Milton Twp., April 27,
1966, KE
Luzula carolinae S. Wats. CFC344, Jackson Twp., April 8, 1967, BHO
Abundant
Luzula echinata (Small) F. J. Hermann Allison W. Cusick, #1834, Milton Twp., April 27,
1966, KE
Luzula multiflora (Retz.) Lejeune CFC467, Up, May 25, 1967, BHO Abundant
LILIACEAE
A Ilium canadense L. CFC861, Jackson Twp., Aug. 1, 1967, BHO & YUO
Sparse
*A Ilium tricoccum Ait. CFC353, Ter, April 15, 1967, BHO Common
* Asparagus officinalis L. CFC555, Ter, June 8,1967, BHO & YUO Common
Camassis scilloid.es (Raf.) Cory Cheri K. Jones, #2, Austintown Twp., May 21,
1968, YUO
Disporum lanuginosuni (Michx.)
Nicholson Ronald W. Hathaway, #53, Boardman Twp., May
14, 1967, YUO
Erytkronium albidum Nutt. James E. Madden, #36A, Poland Twp., April 3,
1967, YUO
Erythronium americanum Ker. CFC353A, Bot, April 15, 1967, BHO
CFC359, Ter, April 15, 1967, BHO Abundant
*Hemerocallis fulva L. CFC684, Bot, July 1, 1967, BHO
CFC676, Ter, July 1, 1967, BHO Abundant
Lilium canadense L. Ervin M. Herrick, Boardman Twp., Aug. 8, 1963,
OS
Maianthemum canadense Desf. CFC450B, Sprinfield Twp., May 23, 1967, BHO &
YUO Common
YUO Common
Medeola virginiana L. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Polygonatum commutatum (Schultes f.)
A. Dietr. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Polygonatum pubescens (Willd.) Pursh. CFC464, Ter, May 25, 1967, BHO & YUO
CFC486, Up, May 31, 1967, BHO & YUO
Common
Smilacina racemosa (L.) Desf. Inga S. Worley, #33, Jackson Twp., May 13, 1965,
YUO
Smilax herbacea L. CFC610, Jackson Twp., June 18, 1967, BHO
Sparse
Trillium erectum L. CFC391, Boardman Twp., April 26, 1967, BHO
Common
Trillium grandiflorum (Michx.) Salisb. CFC474, Ter, May 25, 1967, BHO
CFC489, Up, May 31, 1967, BHO Abundant
No. 1 HERBACEOUS FLORA OF MAHONING COUNTY 37
Trillium sessile L. Ronald W. Hathaway, #181 & Mark Singer, #127,
Poland Twp., April 26, 1968, YUO
Uvularia grandifiora Sm. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Uvularia perfoliata L. CFC483, Up, May 31, 1967, BHO & YUO
Common
Uvularia sessilifolia L. James E. Madden, #53, Poland Twp., April 23,
1967, YUO
DIOSCOREACEAE
Dioscorea villosa L. Ervin M. Herrick, Ellsworth Twp., July 20, 1963,
OS
IRIDACEAE
Iris pseudacorus L. CFC612, Jackson Twp., June 18, 1967, BHO
Common
Sisyrinchium angustifolium Mill. CFC507, Bot, May 31, 1967, BHO & YUO
CFC529, Ter, June 8, 1967, BHO
CFC681, Jackson Twp., July 1, 1967, YUO
Common
OKCHIDACEAE
Cypripedium acaule Ait. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Goodyera pubescens (Willd.) R. Br. Ervin M. Herrick, Poland Twp.,'Aug. 8, 1963, OS
Habernuria lacera (Michx.) Lodd. Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Liparis lilifolia (L.) Richard Ervin M. Herrick, Poland Twp., June 30, 1960, OS
Spiranthes cernua (L.) Richard. Allison W. Cusick, Beaver Twp., Sept. 15, 1960, OS
DlCOTYLEDONEAE
URTICACEAE
Boehmeria cylindrica (L.) Sw. Tom S. Cooperrider, #6930, Milton Twp., July 18,
1960, KE
Laportea canadensis (L.) Gaud. CFC832, Up, Aug. 1, 1967, BHO Common
Pilea pumila (L.) G CFC915, Bot, Aug. 17, 1967, BHO & YUO
Common
Urlica dioica L. CFC842, Bot, Aug. 1, 1967, BHO Common
ARISTOLOCHIACEAE
Asarum canadense L. Tom S. Cooperrider, #6826, Poland Twp., July 15,
1960, KE
POLYGONACEAE
Polygonum "arifolium L. CFC916, Ter, Aug. 17, 1967, BHO Sparse
Polygonum aviculare L. CFC971, Jackson Twp., Sept. 25, 1967, BHO
Abundant
_ _mum convolvulus L. CFC834, Up, Aug. 1, 1967, BHO Abundant
Polygonum hydropiperoides Michx. Tom S. Cooperrider, #7036, Ellsworth Twp., July
19, 1960, KE
Polygonum lapathifoliuni L. CFC851, Bot, Aug. 1, 1967, BHO Abundant
^Polygonum orientale L. CFC941, Green Twp., Sept. 9, 1967, BHO & YUO
Rare
Polygonum pensylvanicum L. CFC845, Bot, Aug. 1, 1967, BHO
CFC900, Up, Aug. 8, 1967, BHO Common
Polygonum persicaria L. CFC911, Bot, Aug. 1, 1967, BHO Common
Polygonum punctatum Ell. CFC932, Bot, Sept. 9, 1967, BHO Common
Polygonum sagittatum L. CFC926, Bot, Sept. 9, 1967, BHO Sparce
Polygonum scandens L. CFC924, Bot, Sept. 9, 1967, BHO Common
Polygonum virginianum L. CFC837, Ter, Aug. 1, 1967, BHO & YUO
CFC833, Up, Aug. 1, 1967, BHO Common
Rumex acetosella L. CFC404, Bot, May 13, 1967, BHO
CFC421, Ter, May 13, 1967, BHO Common
Rumex crispus L. CFC651, Bot, June 27, 1967, BHO
CFC702, Springfield Twp., July 8, 1967, YUO
Common
Rumex obtusifolius L. CFC660, Ter, June 27, 1967, BHO Common
Rumex verlicillatus L. Tom S. Cooperrider, #6933, Milton Twp., July 18,
1960, KE
38 CARL F. CHUEY Vol. 72
CHENOPODIACEAE
*Atriplex patula L. CFC942, Ellsworth Twp., Sept. 9, 1967, BHO
Sparse
Chenopodium album L. CPC862, Bot, Aug. 5, 1967, BHO Abundant
Chenopodium glaucum L. Tom S. Cooperrider, #7043, Jackson Twp., July 19,
1960, KE
AMARANTHACEAE
Acnida altissima Riddell Allison W. Cusick, Beaver Twp., Aug. 12, 1962, OS
*Amaranthus retroflexus L. CFC885, Bot, Aug. 5, BHO
CFC927, Ter, Sept. 9, 1967, BHO Abundant
*Amaranthus albus L. CFC966, Jackson Twp., Sept. 25, 1967, BHO &
YUO Rare
PHYTOLACCACEAE
Phytolacca americana L. CFC677, Ter, July 1, 1967, BHO Common
AIZOACEAE
Mollugo verticillata L. CFC973, Jackson Twp., Sept. 25, BHO Common
PORTULACACEAE
* Claytonia virginica L. CFC346, Bot, April 10, 1967, BHO
CFC360, Ter, April 15, 1967, BHO Abundant
CARYOPHYLLACEAE
A Isinoideae
*Arenaria serpyllifolia L. CFC505, Bot, May 31, 1967, BHO Common
Cerastium nutans Raf. Allison W. Cusick, Beaver Twp., June 11, 1961, OS
*'Cerastium vulgatum L. CFC407, Bot, May 13, 1967, BHO & YUO
CFC421A, Ter, May 13, 1967, BHO Common
Stellaria graminea L. E. B. Williamson, June 1, 1900, OS
Stellaria longifolia Muhl. Clara Weishaupt, Goshen Twp., June 16, 1956, OS
Stellaria media (L.) Cyrill. CFC390, Bot, April 25, 1967, BHO
CFC438, Ter, May 13, 1967, BHO Abundant
Paronychioideae
*Paronychia canadensis (L.) Wood CFC840A, Bot, Aug. 1, 1967, BHO Common
Scleranthus annuus L. Warren Knopp, Canfield Twp., June, 1960, OS
Silenoideae
*Dianthus armeria L. CFC641, Bot, June 27, 1967, BHO & YUO
Abundant
*Dianthus barbatus L. CFC656, Ter, June 27, 1967, BHO Sparse
Lychnis alba Mill. CFC659, Ter, June 27, 1967, BHO Common
*Lychnis coronaria (L.) Desr. CFC655, Ter, Tune 27, 1967, BHO Sparse
Saponaria officinalis L. CFC850, Bot, Aug. 1, 1967, BHO & YUO
CFC881, Ter, Aug. 5, 1967, BHO Common
*Silene antirrhina L. CFC606, Jackson Twp., June 18, 1967, BHO
Common
Silene cucubalus Wibel Tom S. Cooperrider, #7052, Jackson Twp., July 19,
1960, KE
CERATOPHYLLACEAE
Ceratophyllum demersum L. Tom S. Cooperrider, #7061, Jackson Twp., July 19,
1960, KE
NYMPHAEACEAE
*Nuphar advena (Ait.) Ait. f. CFC790, Bot. July 18, 1967, BHO Sparse
Nymphaea odorata Ait. CFC711, Beaver Twp., July 8, 1967, BHO
Common
RANUNCULACEAE
Actaea alba (L.) Mill, Ervin M. Herrick, Poland Twp., June 30, 1960, OS
-dwewowe quinquefolia L. Michael C. Kniseley, #36, Canfield Twp., May 7,
1967, YUO
*Anemonella thalictroides (L.) Spach CFC425, Ter, May 13, 1967, BHO
CFC482, Up, May 13, 1967, BHO & YUO
Common
No. 1 HERBACEOUS FLORA OF MAHONING COUNTY 39
Aquilegia canadensis L. Cheri K. Jones, #16, Austintown Twp., June 1,
1968, YUO
Caltha pahcstris L. CFC343, Bot, April 6, 1967, BHO & YUO
Common
Cimicifuga racemosa (L.) Nutt. CFC805, Ter, July 18, 1967, BHO
CFC715, Beaver Twp., July 8, 1967, YUO Sparse
Clematis virginiana L. CFC829A, Bot, July 25, 1967, BHO Sparse
Delphinium ajacis L. Tom S. Cooperrider #6855, Poland Twp., July 15,
1960, KE
Delphinium exaltatum Ait. Cheri K. Jones, #1, Austintown Twp., May 21,
1968, YUO
*Delphinium tricorne Michx. CFC473, Ter, May 25, 1967, BHO Sparse
Hepatica acutiloba DC. Tom S. Cooperrider, #7889, Poland Twp., July 15,
1960, KE
Hepatica americana (DC.) Ker CFC350, Ter, April 13, 1967, BHO
CFC838, Up, Aug. 1, 1967, BHO Common
* Ranunculus abortivus L. CFC386, Bot, April 25, 1967, BHO
CFC437, Ter, May 13, 1967, BHO & YUO
CFC455, Up, May 25, 1967, BHO Common
Ranunculus acris L. CFC551, Ter, June 8, 1967, BHO & YUO
CFC496, Up, May 31, 1967, BHO Common
^Ranunculus bulbosus L. CFC453, Ter, May 25, 1967, BHO Sparse
Ranunculus pensylvanicus L. f. Allison W. Cusick, Beaver Twp., Aug. 27, 1961, OS
Ranunculus recurvatus Poir. CFC452, Ter, May 25. 1967, BHO Common
*Ranunculus repens L. CFC419, Bot, May 13, 1967, BHO & YUO
CFC451, Ter, May 25, 1967, BHO
Ranunculus septentrionalis Poir. James Madden, #43, Poland Twp., April 13, 1967,
YUO
Thalictrum dasycarpum Fisch. &
Ave'-Lall. Ervin M. Herrick, Poland Twp., June 10, 1961, OS
Thalictrum dioicum L. Tom S. Cooperrider, #6481, Poland Twp., July 15,
1960, KE
Thalictrum polygamum Muhl. CFC642, Bot, June 27, 1967, BHO
CFC619, Ter, June 22, 1967, BHO Common
Thalictrum revolutum DC. Clara Weishaupt, Goshen Twp., June 16, 1956, OS
BERBERIDACEAE
Caulophyllum thalictroides (L.) Michx. Tom S. Cooperrider, #6890, Poland Twp., July 15,
1960, KE
Podophyllum peltatum L. CFC540, Ter, June 8, 1967, BHO
CFC498, Up, May 31, 1967, BHO & YUO
Abundant
PAPAVERACEAE
* Chelidonium majus L. CFC530, Bot, June 8, 1967, BHO
CFC532, Ter, June 8, 1967, BHO & YUO Sparse
Sanguinaria canadensis L. Gayle J. Grover, #26, Poland Twp., April 25, 1967,
YUO
FUMARIACEAE
*Dicentra canadensis (Goldie) Walp. CFC389, Ter, April 25, 1967, BHO Sparse
*Dicentra cucullaria (L.) Bernh. CFC345, Bot, April 10, 1967, BHO
CFC355, Ter, April 15, 1967, BHO Sparse
CRUCIFERAE (BRASSICACEAE)
*Arabis laevigata (Muhl.) Poir. CFC469A, Ter, May 25, 1967, BHO Common
Barbarea vulgaris R. Br. CFC413, Bot, May 13, 1967, BHO
CFC435, Ter, May 13, 1967, BHO
CFC395, Jackson Twp., May 4, 1967, YUO
Common
*Capsella bursa-pastoris (L.) Medic. CFC411, Bot, May 13, 1967, BHO & YUO
Abundant
*Cardamine bulbosa (Schreb.) BSP. CFC412, Bot, May 13, 1967, BHO & YUO
CFC432, Ter, May 13, 1967, BHO Common
Cardamine douglassii (Torr.) Britt. CFC352, Up, April 15, 1967, BHO Abundant
Cardamine pensylvanica Muhl. Tom S. Cooperrider, #6854, Poland Twp., July 15,
1960, KE
*Dentaria diphylla Michx. CFC387, Ter, April 25, 1967, BHO & YUO
Common