Table Of ContentOriginals 周，，b1yJふKM市巨一物取1.H'hooT打M涜蜘雑LmMHA1-一三士心川凶∞、、，J
Systematic Studies of Asian Aconitum (Ranunculaceae) VIII.
，
Subgenus Tanguticathe Forth New Subgenus of the Genus Aconitum
and aN ew Species Belonging to the Subgenus Tangutica
Yuichi KADOTA
Departrnent of Botany，Na tional Science Museurn，
Arnakubo 4-1-1，Ts ukuba，Ib araki，30 5-0005 JAPAN
(Received on February 27，2 001)
Aconitum species which have scapose to subscapose habit，b ranched perennial root-
stocks，no n-tubular nectyblades in the distal part and navicular to conical helmets e
訂 紅
regarded to belong to as ubgenus，Ta ngutica (W.T.Wang) Kadota，w hich is reg紅白das
the forth subgenus in the genus Aconitum. Within the subgenus Tangutica three sections
erecognized; sect. Fletcherum Tamura，s ect. Tangutica and sect. Zeravschanica
紅
Kadota. Sect. Fletcherum is emended and Sect. Zeravschanica is newly described in this
paper. Aconitum qinghaiense is described from China as a member of subgenus
Tangutica sect. Tangutica. (Continued from J. Jpn. Bot. 76: 20-27，2 001)
Key words: Aconitum qinghaiense，n ew taxa，se ct. Fletcherum，se c .tZeravschanica，s ub-
genus Tangutica
Aconitum tanguticum was first published zomes have been calssified into subgenus
by Maximowicz (1899) as 'A. rotundifolium Lycoctonum (e.g.， Lauener and Tamura
Kar. & Kir. var. tanguticum Maxim.' Later 1979，W .T.Wang 1979，T amura 1995) [or a
Stapf (1905) recognized this entity to be a distinct genus， Lycoctonum (e.g.， Nakai
distinct species and published an ew name A. 1953)]. Then does A. tanguticum belong to
tanguticum (Maxim.) Stapf. 1a gree Stapf' s subgenus Lycoctonum? Most members of
佐eatmentbecause these two species，A . subgenus Lycoctonum have helmets higher
伽 guticumand A. rotundifolium，ar e differ- than broad. In the case of the species with
ent in the shape of nectyspurs (Kadota helmets broader than tall，t he helmets e
紅 訂
1998). Furthermore there is as ignificant diι provided with long anterior parts (e.g.，A .
ference in root system: A. rotundifolium has novoluridum Munz; Lauener and Tamura
biennial tubers (mother tuber plus daughter 1979，f ig. l-C). Aconitum tanguticum has
tuber) while A. tanguticum，a t least its navicular helmets which ec1early broader
紅
lectoparatype specimens，h as a perennial than tal1. Next， species of subgenus
rootstock (Przewalski 307，L E! ; Kadota Lycoctonum have tubular parts in the distal
1998，f ig. l-C). Aconitum tanguticum is a p tof necatry blades (e.g.，L auener and
紅
smal1 scapose plant and grows in rocky al・ Tamura 1979， fig. 1， except for A.
pine meadows. Owing to difficulty to dig the moschatum and A. fletcheranum; Kadota
rootstocks the specimens often do not have 1999，f ig. 3). On the other hand，n ect y
紅
subterranean organs. blades of A. tαnguticum enot provided
紅
Aconitum species having perennial rhi- with tubular p紅白. 1 conc1uded that A.
-183ー
184 植物研究雑誌第76巻第4号 平成13年8月
tanguticum does not belong to subgenus portions. It is highly doubtful that A.
Lycoctonum nor subgenus Aconitum. Here 1 moschatum belongs to subgenus Lycocto-
propose subgenus Tangutica based on ser. num. This issue will be discussed elsewhere.
Tangutica W.T.Wang (1979).
1 have already pointed out that A. Subgenus Tangutica (W.T.Wang) Kado-
zeravschanicum Steinb. from Central Asia ta，st at. nov.
and A. popovii Steinb. & Schisch. ex Sipliv. Ser. Tangutica W.T.Wang，F .1Reipub .1
from East Siberia have also perennial root- Pop. Sin. 27: 604 (1979).
stocks (Kadota 1997). Aconitum fletchera- Sect. Sinαconitum auct. non W.T.Wang:
num G. Tay .1 from East Himalaya has been Tamura，N atur .1Pflanzenfam. 17 aI V: 284
considered to be am ember of the subgenus (1995)，p. p. minore.
Lycoctonum (Tamura 1966，1 995，L auener Type species: Aconitum tanguticum
and Tamura 1979，W .T (Maxim.) Stapf.
this species bears perennial rootstock (Fig. Distribution: W China，E Siberia and
め5). However， necty blades of A. Central Asia.
紅
fletcheranum are not provided with tubular
parts (Tamura 1995，fi g. 15，9 -10，T amura Sect. 1. Fletcherum Tamura emend.
and Lauener 1979， fig. lB; Fig. 4C). Kadota.
Aconitum fletcheranum should be therefore Subgenus Paraconitum Rapaics sect.
classified into subgenus Tangutica. Aconitum Fletcherum Tamura in Sci. Rep. Osaka Univ.
spiripetalum Hand.-Mazz. from Sichuan 15: 30 (1966)-Lauener & Tamura in Notes
Province，C hina，is characterized by having Roy. Bot. Gard. Edinb. 37: 117 (1979)-
perennial rootstocks，n avicular helmets and W.T.Wang，F .1Reipub1. Pop. Sin. 27: 179
non tubular nectary blades and is also re- (1979)-Subgeus Lycocωnum (DC.) Peterm.
garded as a member of the subgenus sect. Fletcherum Tamura，N atur1. Pflanzen-
Tangitica. fam. 17 aI V: 280 (1995).
Conducting field examination in Qinghai Type species: Aconitum fletcheranum
Province，C hina，1 f ound Aconitum plants G.Tay1.
which were similar to but significantly diι Herba p vaperennis scaposa. Caulis sim-
紅
ferent from A. tanguticum. After repeated plex basi obtectus vestigiis fibris. Folia
field surveys and examination of herbarium basales rosulatae laminis membranaceis
specimens，1 c ame to ac onclusion that these ambitu reniformis profunde trilobatis，
plants belongs to an ew species. The subge- petiolis basi vaginatis. Flos solitarius
nus Tangutica is consequently consisted of terminalis. Cassis navicularis rostrio brevi
the following six species: A. fletcheranum， obtuso. Calcaria nectariorum bacilliformia
A. popovii，A. qinghaiense，A. spiripetalum， leviter incurvata. Labella nectariorum breves
A. tanguticum and A. zeravschanicum. fimbris conspicuis auriculatis. Carpella 6-8.
Aconitum moschatum (Bruhl) Stapf de- 1. Aconitum fletcheranum G.Tay1. in J.
scribed from Kashmir has been considered to Roy. Hort. Soc. London 77: 242 (1952)
ー-
be am ember of subgenus Lycoctonum，si nce Tamura in Sci. Rep. Osaka Univ. 15: 30
this species has perennial rhizomes (Stapf (1966)-Lauener & Tamura in No tes Roy.
1905，pl . 94，T amura 1966，1 995，f ig. 15， Bot. Gard. Edinb. 37: 117 (l979)-W.T.
11-12，L auener and Tamura 1979，T amura Wang，F 1. Reipub1. Pop. Sin. 27: 179 (1979)
and Lauener 1979，fi g. 1A ，C haudhary and -Gleason & Long，F .1Bhutan 1(2): 316
Rao 1998). However，n ectary blades of A. (1984)-Shma，F .1Ind. 1: 13 (1993).
訂
moschatum are not equipped with tubular TYPE: BHUTAN: Pangotang，T sampa，
August 2001 Journal of Japanese Botany Vo .l76 No. 4 185
alt. 14500 ft，1 2 September 1949，pe rianth a 11: 2 (1996)，p . p. minore (Forrest 14643
very bright violet，a b eautiful colour，w ith a from Yunnan，K) .
striking white edge to the inner segments，on Differt ab Aconito popovii labellis
cliff ledges of steep gravery open slopes，F. nectariorum reflexis cum marginibus leviter
Ludlow，G . Sherriff & J. H. Hicks 19734 auriculatis，ca ssidibus navicularis cum rostris
(holotype-E!) . longis; ab A. tangutico calcaribus nectario-
Distribution: SE Tibet， Bhutan and rum cupulatis，l abellis nectariorum brevis
Assam. sed manifestis.
Aconitum fletcheranum is characterized by A subscapose perennial，1 0-30 (-70) cm
having pedicels sp ingly strigose with tall. Rootstock branched into narrowly car-
紅
roughsurfaced curved hairs， scape base rot-shaped tubers 1-3 cm long，ca . 5m mi n
田
clothed with fibrous remains of petioles and diameter. Stem terete，1 -2 times branched to
nectary labia with auriculate flanges. Nectary simple. Basal leaves rosulate，s ubcoriaceous，
spurs are less developed and show ad egen- roundish reniform，3 -4 cm long，3 .5-5 cm
erated condition (Fig. 4C). It is possible that broad，m ediall y( to 1/2f rom the base) 5-9-
they secrete no nect This should be inves- lobed，g labrous on both sides; middle lobes
訂.
tigated in the field. broadly obovato-rhombic，1 -2 cm long and
broad，c oarsely dentate with obtuse teeth;
Sect. 2. Tangutica. petioles 5-8 cm long，g labrous，v aginate at
Sect. Sinaconitum auct. non W.T.Wang: base. Cauline leaves 2-3，pe ntagonal in out-
Tamura，N a turl. Pflanzenfam. 17 aI V: 284 line，d eeply trilobed with deeply incised
(1995)，p. p. lobes，g labrous，s hortly petiolate. Flowers in
2. Aconitum popovii Steinb. & Schisch. July to August，pa le greenish white or some-
ex Sipliv. in Nov. Sist. Vyssh. Rast. 11: 335 times bright blue violet，2- 4 in ac ompact ra-
(1974)-Kadota in Natur. Environ. Sci. Res. ceme，c a. 2 cm long and broad，s paringly
9: 7，fi g. 2E(1996). strigose with rough-surfaced .cu rved hairs.
田
TYPE: RUSSIA; Central Siberia，市tlontes Pedicels erect，5 -10 mml ong，s trigose with
S司jan Orientalis， jugum Tunkense， fl. rough-surfaced curved hairs; bracteoles
Chubuta，i n prato temperate irrigito' [East linear，5 mml ong，1 mmb road，s ituated
Sayan Mountains，th e Tunkinskyi Range]，11 around the middle of the pedicels. Helmets
July 1957，L . M. Malyshev & N. Misjura s. navicular with long attenuate beaks，1 c m
n. (holotype-LE!; isotypes-NS!). ta ，1l 2 cm broad; lateral sepals broadly
Distribution: the Tunkinskyi Range，E obovate，1 7-18 mml ong and broad，d evoid
Siberia (endemic). of long pollen-collecting hairs on the adaxial
Aconitum popovii is characterized by hav- side. Nectaries glabrous; spurs cup-shaped，
ing pedicels villose with smooth-surfaced erect，d ark purple，ca . 2m ml ong; labia ca. 1
patent hairs and cupshapednectary spurs. mm long， dark purple， clearly bilobed，
田
As stated in Kadota (1996)，A. popovii is dis- strongly reflexed， provided with slightly
criminated from A. tanguticum by having auriculate flange; claws slightly incurved，
cup-shaped nectyspurs and short but dis- 12-14 mm long. Carpels 4-5， glabrous.
紅
tinct emarginate labia. Stamens glabrous or sp inglyvillose with
紅
smooth-surfaced patent hairs，p rovided with
3. Aconitum qinghaiense Kadota，s p. staminal teeth. Mature seeds unknown; im-
nov. [Figs. 1-3，4 A] mature seeds smooth and not lamellate
Aconitum tanguticum auct. non (Maxim.) across.
Stapf: Kadota in Natur. Environ. Sci. Res. TYPE: CHINA; Prov. Qinghai，X ining，
186 植物研究雑誌第76巻第4号 平成13年8月
Fig. 1. Type specimen of Aconitum qinghaiense Kadota (CHINA; Prov. Qinghai，X ining，
Huanzhong Xian，L 司jishankou，alt. 3560 m，2 8 July 1999，Y. Kadota 994010，h olotype，T NS).
The left corner inset shows ap erennial rootstock (the same locality，Y . Kadota 994008，T NS).
August 2001 Journal of Japanese Botany Vol. 76 No. 4 187
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188 植物研究雑誌第76巻第4号 平成13年8月
Huanzhong Xian，L ajishankou，a lt. 3560 m， Aconitum qinghαiense is distinguished
28 July 1999，Y .Kadota 994010 (holotype- from A. popovii by having reflexed nectary
TNS). labia with slightly auriculate flanges and
Chinese name: Qinghai-wutou (青海烏頭). navicular helmets with long beaks，a nd dis-
Distribution: EQ inghai，S E Tibet and SW criminated from A. tαnguticum by cup-
，
Gansu (Fig. 3). shaped nectyspurs and short but distinct
紅
Specimens examined: CHINA; Prov. Qinghai， labia.
Xining，H uanzhong Xian，L 司jishankou，alt. 3560 m，2 8
July 1999， Y. Kadota 994004-994009， 994011- 4. ACω0凶姐E0it佃um叩spi廿ri恰petalumHa加ndι.
994012 (TNS); Prov. Qinghai，H ainan Tibetan Autom.
i凶n幻ActaHort. Gothob. 13: 91 (l939)-W.T.
State，G uide-Tongde，al t. 3750 m，1 3 August 1991，Y.
Wang，F l. Reipubl. Popul. Sin. 27: 184，p l.
Kaodta 18016a (TNS). SE TIBET: Sarong，o n Doke la，
Mekong-Salwin divide，la t. 28020'， alt. 13000 ft，pl ant 33，自gs.7-8 (1979).
of 2-3 inches，f lowers deep blue，o n stony pasture， TYPE: CHINA: Sikang，T aofu (Dawo)
Aug. 1917，G . Forrest 14673 (E); SE TIBET，1 917- Distr.， Haitzeshan， in declioli herbosa-
1919，G . Forrest 14643 (K). SWG ANSU: T'ao River fruticoso，c a. 4200 m，3 1 August 1934，H .
basin，a lpine meadow of Choni，al t. 9000ー10000ft， Smith 11711 (holotype-WU!; isotypes-E!，
July 1925，1. F. Rock 12582 (LE).
Fig. 3. Distribution of Aconitum qinghaiense Kadota.
August 2001 Journal of Japanese Botany Vo .l76 No. 4 189
PE!). 261， fig. 223 (1974)-W.T.Wang， Fl.
Distribution: W Sichuan (endemic). Reipubl. Popul. Sin. 27: 182，p l. 33，fi gs. 1-'-
Aconitum spiripetalum is characterized by 4 (1979); Fl. Xizang. 2: 22，f ig. 5，1 -2
pedicels strigose with rough-surfaced curved (1985)-Bot. Ins ，.t Acad. Sin.，I con. Corm.
hairs and strongly coiled necatry claws，h ow Sin. 1: 697，fi g. 1934 (1980)-C.Y.Wu &
細
ever，n ect訂yspurs are basically similar to a ，.l Ind. Fl. Yunnan. 97 (1984)-:-Kadota，
those of A. qinghaiense. Nectary labia of A. Himal. Pl. 1: 215 (1988)-W.T.Wang & a ，.l
spir伊etalum eprojecting downward and Vasc. Pl. Hengduan Mts. 1: 455 (1993)-
紅
entire while those of A. qinghaiense are Aconitum rotundifolium Kar. & Kir. var.
clearly bilobed and reflexed (Figs. 4，A ，B) . tanguticum Maxim.，F l. Tangut. 26 (1899)
Nectar secretion of A. spiripetalum was con- -Finet & Gagnep. in Bull. Soc. Bot. Fr.
firmed by me at Palanshan (alt. 4200 m)， 53: 213 (1905).
Prov. Sichuan，C hina，o n 21 August 1996. LECTOTYPE (Kadota 1998): CHINA:
Terra Tangutorum [Prov. Gansu]，B urchan
5. Aconitum tanguticum (Maxim.) Stapf Budda，al t. 13-15500 ft，1 3 August 1884，fl.
in Ann. Roy. Bot. Gard. Calc. 10: 51 coeruleoviolacei，N .M.Przewalski 391 (LE!，
(1905)一Hand.-Mazz.in Acta Hort. Gothob. Kadota 1998，fi g. lA; isotype-K!).
13: 90 (1939)-Vorosch. in Byull. Glavn. Distribution: Gansu， Qinghai， Shanxi，
Bot. Sada (Moskva) 64: 33，3 7 (1967)-Inst. Sichuan，Y unnan and ET ibet.
Bot. Bor.-Occ. Acad. Sin.，F l. Tsinling. 1: Aconitum tanguticum is characterized by
Fig. 4. Comparison in the spape of helmets and nectaries of Aconitum species. A. A. qinghaiense
Kadota (CHINA: Prov. Qinghai，Ha inan Tibetan Autom. State，Gu ide-Tongde，al t. 3750 m，13
August 1991，Y. Kadota 18016a，TN S). B. A. spiripetalum Hand.-Mazz. (CHINA: Sikang，Ta ofu
(Dawo) Distr.，Ha itzeshan，ca . 4200 m，31 August 1934，H. Smith 11711，is otypes，E) . C. A.
fletcheranum G.Tayl. (BHUTAN: Pangotang，T sampa，al t. 14500，白 12 September 1949，F.
Ludlow，G. Sherriff & 1. H. Hicks 19734，ho lotype，E) . Scale 5m m; short scale 1m mf or mag-
nified nectary blades.
190 植物研究雑誌第76巻第4号 平成13年8月
Fig. 5. Type specirnen of Aconitum fletcheranum G.Tayl. (BHUTAN: Pangotang，T sarnpa，al t.
14500 ft，1 2 Septernber 1949，F. Ludlow，G . Sherriff & J. H. Hicks 19734，h olotype，E) . The
right corner inset shows ap erennial rootstock (INDIA: Assarn，Z e La，al t. ca. 15000 ft，2 0
August 1938，F. Kingdon-Ward 14122，E) .
August 2001 Journal of Japanese Botany Vol. 76 No. 4 191
having pedicels villose with flexuous hairs …1. A. fletcheranum
and slightly incurved，ve rsatile nectary spurs la. Nectary spur saccate，cu p-shaped to ellip-
with obscure labia. Nectar secretion of A. soidal
tanguticum was also verified by me at 2a. Nectary spur ellipsoidal; labia long，
Balanshan (alt. 4400 m)，P rov. Sichuan， conspicuously bilobed (Sect.
China，o n 22 August 1996. Refer to Kaodta Zeravschanica)…6. A. zeravschanicum
(1998) for ad etailed description. 2b. Nectary spur cup-shaped， slightly
incurved or erect; labium short (Sect.
Sect. Zeravschanica Kadota，se ct. nov. Tangutica)
Herba mediocris perennis subscaposa. 3a. Nectary spur slightly incurved; la-
Caulis duplo vel triplo ramosum. Lamina bium obscure...・・...5.A. tanguticum
H
foliorum coriacea ambitu rotundatae. 3b. Nectary spur erect; labium short but
Inflorescentiae racemosae. Cassides obvious
naviculares rostris brevibus obtusis. Calcaria 4a. Nectystalk strongly coiling at
紅
nectariorum ellipsoidea labellis longis proximal part; labium entire.........
profunde bilobatis. Carpella 5. Semina ……4. A. spiripetalum
rugosa non lamellata. 4b. Nectystalk slightly incurved;
紅
Type species: Aconitum zeravschanicum labium bilobed
Steinb. 5a. Nectary labium not reflexed;
6. Aconitum zeravscahnicum Steinb.，F l. helmet short beaked...・・....・ ・...
H H
URSS 7: 230，7 33 (1937)一一Butkov，Fl. ・... ・....2. A. popovii
H
Uzbek. 2: 463 (1953)-Shpota，F l. Kirghis. 5b. Nectylabium reflexed; hel-
紅
6: 60 (1955)-Wasylywa in Nov. Sys .t met long beaked........・・.....
H
Vyssh. 1966: 243 (1966)-Kadota in Natur. ・・・..…...・ ・.3.A. qinghaiense
H H H
Environ. Sci. Res. 9: 6，fi g. 2-D (1996).
TYPE: CENTRAL ASIA: Uz bekistan;‘N. 1w ish to give my sincere thanks to Profs.
slope of the Turkestan Range，恥1:t.Kurgansk W. T. Wang and Q. E. Yang for discussion，
ne thesource of the Au-korchegai River， advice and suggestion on Aconitum; Mr. L.
訂
among junipers，at the height of 2710 m'，2 S.Wang，K unming Institute of Botany，f or
August 1934，K . Afanatsev 424 (holotype- his aangementof the field work in Sichuan
町
LE!). Province，C hina.恥1:Y thanks also go to the
Aconitum zerlαvschanicum is characterized curators of the Herbaria (BM，C AL，E ，K ，
by pedicels strigose with rough-surfaced KATH，K UN，L E，M W，N S，P E，T I，T NS，
curved hairs，w hitish flower color and ellip- W and WU). Field works were financially
soidal nect訂ゲ spurswith long bilobed labia. supported by the Grants-in-Aid from the
Nectar secretion of this species was con- Ministry of Education， Science， Sports，
firmed by me at the Alaisky Range， Culture and Technique， Japan (Nos.
Kirghistan，o n 10 August 1995. Refer to 07041151， 07041152). Herbarium works
Kadota (1996) for ad etailed description. were supported by the Delegation Programs
of the Jap an Society for the Promotion of
Key to the species Sciences (1992，1 998) and the Program of
la. Nectary spur rod-shaped，s lightly bend- Oversea Research Scholars from the
ing inwardly; nectylabium short，e ntire， Ministry of Education， Science， Sports，
訂
provided with auriculate flange; scape Culture and Technique，J apan (2000).
clothed with fibrous remains of petioles
(Sect. Fletcherum Tamura)…・.，
192 植物研究雑誌第76巻第4号 平成13年8月
References Nakai T. 1953. A new classification of Lycoctonum
Chaudhary L. B. and Rao R. R. 1998. Notes on the and Aconitum in Korea，Ja pan，a nd their surround-
genus Aconitum (Ranunculaceae) in North-West ing Areas. Bull. Natn. Sci. Mus.，T okyo (32): 1-53.
Himalaya (India). Feddes Repert. 109: 527-537. Sharma B. D. (eds.) 1993. Aconitum. Flora of India 1:
Gleason A. J. C. and Long D. G. 1984. Aconitum. 5-23. Botanical Survey of India，Ca lcutta.
Flora of Bhutan including ar ecord of plants from Stapf O. 1905. Aconites of India: am onograph. Ann.
Sikkim. 1，Pa rt 2: 314-321. Royal Botanic Garden， Roy. Bot. Gard. Calc. 10: 115-197.
Edinburgh. Tamura M. 1966. Morphology，ec ology and phylogeny
Kadota Y. 1996. Systematic studies of Asian Aconitum of the Ranunculaceae 6. Sci. Rep. Osaka Univ.
(Ranunculaceae) 1. Notes on Aconitum (15): 13-35.
rotundifolium and A. zeravschanicum from Tien 一一一一andLauener L. A. 1979. A synopsis of Aconitum
Shan High Mountains， Central Asia. Natur. subgenus Lycoctonum: 11. Notes Roy. Bot. Gard.
Environ. Res. 9: 1-8. Edinb. 37: 431-466.
一一 1998.Systematic studies of Asian Aconitum 一一一 1995.11Systematic part. Engler A. und Prantl K.，
(Ranunculaceae) 111. No tes on Aconitum Naturl. Pflanzenfam. Angiospermae: Ordung
tanguticum (Maxim.) Stapf. Natur. Environ. Res. Ranunculales，F am. Ranunculaceae，1 7 aI V: 223-
11: 1-6. 497. Dunker & Humbolt，Be rlin.
Lauener L. A. and Tamura M. 1979. A synopsis of Wang W. T. 1979. Aconitum. Flora Reipublicae
Aconitum subgenus Paraconitum 1. Notes Roy. Popularis Sinicae. 27: 113-326. Science Press，
Bot. Gard. Edinb. 37: 113-123. Beりing(inC hinese).
門田裕一:アジア産トリカブト属(キンポウゲ科)
の分類学的研究 VIII. 第 4の新E属Tangutica
(W.T . Wang) Kadotaとその1新種 つことからこれまでレイジンソウ亜属に所属する
Aconitum tanguticum (Maxim.) Stapfはチベット 種と考えられてきたが，A. tanguticumと同じよう
東部から雲南省，四川省などを経て険西省にかけ に舟形で幅が高さより明らかに大きい上等片と花
ての地域に分布する中国固有の種である.標本で 弁の舷部に円筒状の部分がないことから，レイジ
はしばしば地下部が失われているが，これはこの ンソウ亜属ではなく， Tangutica亜属に属すると
種が花茎状で，高山帯の岩混じりのやや乾いた風 考えられる.A. fletcheranumはまた，花弁唇部の
衝草原にはえるために植物体全体の採集が容易で 基部が耳状に広がる特異な花弁をもっている.そ
ないことが原因と考えられる.しかし，注意深く こで，単型的な節sect.Fletcherumを認め，レイジ
掘り出すことにより，分枝した多年生の貯蔵根を ンソウ亜属から Tangutica亜属に移し，再記載を
もつことが分かる. トリカブト亜属では貯蔵根は 行った.同様に，キルギスタンとタジキスタンに
越年性であり，毎年更新される.したがって，A. 分布する A.zeravschanicum Steinb.も花弁の距が
αtnguticum はトリカブト亜属の構成員ではないと 楕円体状であり，唇部が大きくかっ2裂すること
いえる. トリカブト属ではレイジンソウ亜属が多 から， トリカブト亜属から Tangutica亜属に移し，
年生の貯蔵根をもつことで特徴付けられるが，A. 単型的な節 sect.Zeravschanicaを新設した.四川
ωnguticumは上等片(ヘルメット)が舟形であり， 省固有のA.伊s~iかr~伊petωa，lωumη1. Ha加ndι.
幅が高さよりも明らかに大きい.レイジンソウ亜 貯蔵根をもつことを確認した.また，雲南省，青
属の種でも幅が高さよりも大きなものもあるが， 海省，甘粛省から A.qinghaiense Kadotaを新種と
この場合は花弁の舷部に円筒状の部分がある.A. して記載した.A.q的ghaienseは，東シベリア特
tanguticumでは花弁の舷部に円筒状の部分はない. 産のA.popovii Steinb. & Schisch. ex Sipliv.とは明
このように，A.ωnguticumはレイジンソウ亜属と 瞭に反曲する花弁唇部と鳴の長い上専片をもつこ
もトリカブト亜属とも異なるので， Tangutica節 とで，A.ωnguticumからは花弁の距がコップ状で
を亜属のランクに引き上げて subgenusTangutica 短いながらも明瞭な唇部をもつことで区別される.
(W.T.Wang) Kadotaとした.Tangutica亜属はトリ したがって，新亜属Tanguticaは3節6種から成
カブト属の中で4番目の亜属となる. り，中アジア，東シベリア，東ヒマラヤ，中国横
チベット東南部，アッサム，ブータンに分布す 断山脈などチベット高原の周辺部に分布する.
るA.fletcheranum G.Tayl.も多年生の貯蔵根をも
