Table Of ContentBLUMEA 45 (2000) 403-425
Notes on Malesian Fabaceae
(Leguminosae-Papilionoideae).
7. The genus Millettia
Frits Adema
Nationaal Herbarium Nederland,UniversiteitLeiden branch, P.O. Box9514
2300RALeiden,The Netherlands
Summary
The genus Millettia Wight& Arn.is revised for theFlora Malesiana area.Two newspecies are
described: M.borneensis and M.glabra.Pongamiavelutina (C.T.White)Verdc, is transferred to
Millettia;astheepithetonvelutina isalreadyinuseinMillettia,anewnameisproposed:M.velve-
tinaAdema.Intotal 15speciesarerecognisedfortheareaconcerned.Akeytothespecies isgiven.
An identification listand anindex ofnamesareincluded.
Keywords: Millettia,Malesia,Solomon Islands.
Introduction
Millettiawas described by Wight & Arnott(1834: 263) to accommodatetwo new
species:M.rubiginosa Wight&Am.andM.splendens Wight&Am.Sincethenmany
species havebeenaddedtothis genus.Dunn(1912)accepted 127species in15sections.
Geesink(1981 a, 1984) transferredseveralsectionstoothergeneraandunitedPongamia
Vent,withMillettiasect.Fragiliflorae(seealsoAdema,2000). InMalesiatwo sections
ofMillettiacanbefound:sect.Millettia(= sect. Typicae Dunn+sect. Macrospermae
Dunn)andsect.FragilifloraeDunn.Inthefollowingtreatment thegenus,bothsections,
andtheirspecies willbereviewed anddiscussed.Twonewspecies willbe described,
M.borneensis andM. glabra,, andonenewname, M. velvetinaAdema,is proposed.
Only thenewspecies are described infull; theotherspecies arementionedwith full
synonymy andnotes. Akey toallspecies ofMillettiainMalesiais given; anidentifi-
cationlistand anindexof namesare included.
MILLETTIA
Millettia Wight& Am., Prod.Fl.Penins. Ind.Or. 1 (1834)263, nom. cons.; Benth. in Miq., PI.
Jungh.(1852)247;Baker in Hook.f.,Fl.Brit.Ind. 2(1876) 104;Taub. in Engl.&Prantl,Nat.
Pflanzenfam.3, 3(1894)270;Prain,J.As.Soc. Beng.66, 2(1897)358;Merr.,Philipp.J.Sci.,
Bot. 5 (1910)70; Dunn,J.Linn. Soc. Bot. 41 (1912) 123;Gagnep.,Fl. G6n. Indo-Chine 2
(1916)361;Merr.,Enum.Philipp.Flow. PI.2(1923)278;Backer& Bakh.f.,Fl.Java 1(1964)
595;Verde., Man. New Guinea Leg. (1979) 332;Allen & Allen,Leguminosae(1981)435;
R.Geesinkin Polhill &Raven,Adv.Leg.Syst. 1(1981)257;Leiden Bot.Ser. 8(1984) 102.—
Millettia Wight& Arn. sect. TypicaeDunn,J.Linn. Soc. Bot.41 (1912) 134.— Typespecies:
Millettia rubiginosaWight&Arn.
404 BLUMEA Vol. 45, No. 2, 2000
PongamiaVent.,Jard. Malm. (1803) t.28, nom. cons.; Merr., Enum.Philipp. Flow. PI.2(1923)
298;Backer& Bakh.f.,Fl.Java 1 (1964)616;Verde.,Man.NewGuineaLeg.(1979)311;Allen
& Allen,Leguminosae(1981)543; R.Geesink in Polhill & Raven,Adv.Leg. Syst. 1 (1981)
259. —Typespecies: PongamiaglabraVent.[= P.pinnata (L.)Pierre],
GaledupaLam.,Encycl. 2(1876)594;Taub. in Engl.&Prantl,Nat. Pflanzenfam.3,3(1891)344.
— Typespecies: Galedupaindica Lam.
Millettia Wight& Arn.sect.FragilifloraeDunn,J.Linn.Soc. Bot. 41 (1912) 136.—Typespecies
notindicated.
Millettia Wight&Am. sect.MacrospermaeDunn, J.Linn.Soc. Bot.41 (1912)136.—Typespecies
notindicated.
Trees, shrubs, or lianas.Leaves imparipinnate, rarely pinnately trifoliolate; stipules
present, caducous; stipellae presentorabsent. Leaflets entire, lateralonesopposite.
Inflorescences axillary and/orterminal,panicles, racemes,pseudoracemes orpseudo-
panicles; insect. Fragiliflorae with apulvinus atthebase; flowersfascicled or not.
Bractspresent, usually caducous;bracteolespresent,usuallycaducous.Calyxcampanu-
lateor ± tubular, teethinconspicuous or uptoslightly longer thanthe tube. Corolla:
standardwith or withoutbasalcallosities, rarely withauricles; wings adhering to the
keel petals; keel petals slightly shorterthanthewings, usually withalateralpocket.
Stamensmono-or diadelphous, intheformercase withbasalfenestrae; anthersbasi-
fixed,insect.Fragiliflorae hairy.Discinconspicuoustoannularor 10-lobedandadnate
tothehypanthium. Ovary subsessiletostipitate, with 2(or3) toseveralovules; style
almostglabrous, or hairy inlowerhalfand glabrous upwards. Pods strap-shaped to
ellipsoidor semi-circular, compressed ornot, dehiscent,rarely indehiscentor tardily
dehiscent.Seedstransversely ellipsoid to 'quadrate' or ± oblong.
Distribution—More than 100 species in thetropics ofAfrica andAsia intothe
Pacific; in Malesiac. 15 species.
Note—Rootnoduleshavebeenreported forM.pinnataandseveralAfricanspecies
(Allen &Allen, 1981).
KEY TOTHESECTIONS OF MILLETTIA IN MALESIA
la. Axillary buds button-shaped, tightly enclosedby thebudscales. Inflorescences
withapulvinus atthebase ofthepeduncle. Standard withtwo basalcallosities.
Anthershairy sect. Fragiliflorae
b. Axillarybudsflattenedovoidto globular,loosely enclosedby thebudscales.Inflo-
rescences withoutapulvinus atthebaseofthepeduncle. Standardwithoutcallosi-
ties.Anthers glabrous sect.Millettia
SectionMillettia
SectionMillettia.—SectionTypicae■Dunn,J.Linn. Soc. Bot. 41 (1912)134,nom.illeg. —Type:
Millettia rubiginosaWight&Am.
SectionMacrospermaeDunn, J.Linn. Soc. Bot. 41 (1912) 136.—Lectotype (heredesignated):
Millettiasericea Benth.
Trees,shrubs,or lianas.Axillarybudsflattenedovoidto globular, looselyenclosedby
thebudscales. Inflorescences axillary orterminalpseudoracemes orpseudopanicles,
nopulvinus atthebaseofthepeduncle. Standardwithoutcallosities.Anthersglabrous,
veryrarelywith somehairs.
F.Adema: Notes on Malesian Fabaceae - Millettia 405
Distribution— SEAsiafromIndiaandSriLankato SChina,Indochina,Thailand
andWMalesia.
Note—- Theonly differencebetweenDunn'ssections 'Typicae’ (= Millettia) and
Macrospermae isthepresenceorabsenceofstipellae.As thisisanunimportant differ-
enceinothersections of.Millettia,thetwo sectionsshouldbeunited.
InMalesiafourspecies belongto thissection:
1.Millettiachrysamaryssa Adema—Fig. 1a
Millettia chrysamaryssa Adema,Novon 9(1999)289. —Type:Maxwell 78-107(AA, L,SINU),
Malaysia,Perak,Padang,Selim River, 12.4.1978.
Note — This new species was, rather surprisingly, found in thematerial identi-
fiedas M. sericea by earlierbotanists. Millettiachrysamaryssa differsmainly from
M.sericeainthesizeofthestipules (5 by3mmvs. 2-3.5by 2-3mm), thesizeofthe
anthers(0.7-1.1 by 0.4-0.6mmvs. 0.6-0.7by 0.4-0.6 mm), thepods (flattened, 14
by 5 by 0.4 cm vs. thickened,5-7.5 by 1.5-3.5by 1-2cm) andtheseeds(flattened,
upright-oblong, 24by 16by 2mmvs. thickened,transverse-ellipsoid, 25-45 by 10-
25by 4-13mm). SeeforafullaccountAdema, 1999.
Fig. 1. Millettia Wight& Arn. sect. Millettia. Pods. a. M.chrysamaryssa Adema;b.M. sericea
(Vent.)Wight&Am.exHassk. (a:Maxwell 78—107;b:King’scoll. 709).
2.Millettiagalliflagrans Whitmore
Millettia galliflagransWhitmore,Fed. Mus.J.13(1970) 136;TreeFl.Mai. 1 (1972)297.—Type
FRI 7551 (RF.Cockburn), Malaysia, Johore,KluangF.R.,atfootofG.Blumut(n.v.).
Note—Aratherenigmaticspecies. Itis asmalltree withunifoliolateleaves.Up to
nowonly knownfromthetype specimen, whichIhave not seenas yet. Thespecimen
FRI 17804(T. Suppiah) maybelong tothisspecies.
3. Millettiasericea (Vent.)Wight& Am.ex Hassk. —Fig. 1b
Millettia sericea(Vent.)Wight&Am.exHassk., Cat.Hort.Bogor.(1844)283;Benth.in Miq.,PL
Jungh.(1852)248;Miq., Fl.Ned. Ind. 1(1855) 153;Prain,J.As.Soc.Beng.66, 2(1897)88;
Dunn,J.Linn. Soc. Bot.41 (1912)169;Gagnep.,Fl.Gen. Indo-Chine 2(1916)383;K.Heyne,
406 BLUMEA Vol. 45, No.2, 2000
Nutt.PL 2(1916)280;Ridl.,Fl.Mai. Penins. 1 (1922)584;Craib,Fl.Siam. Enum. 1 (1928)
394;Burkill,Diet. 2(1935) 1472;Backer&Bakh.f.,Fl. Java 1(1964)596;Whitmore,TreeFl.
Mai. 1 (1972)298. —PongamiasericeaVent.,Jard. Malm. (1803)t. 28.—Type:De Lahaye
s.n.(P?, Herb. Jussieu?),Java (n.v.).
DalbergiaangustifoliaHassk., Cat. Hort.Bogor(1844)284. —Typeunknown
Millettia sericea(Vent.)Wight&Am.exHassk.var. obtusaMiq.,Fl.Ned. Ind. 1 (1855) 154.—
Type:Blumes.n.[holoL(L 908.120-305)],Java.
Millettiasericea (Vent.)Wight& Am.exHassk. forma subcordata Miq.,Fl.Ned. Ind. 1 (1855)
154.—Type: Blumes.n.[holoL(L908.120-311)],Moluccas.
Millettiasericea(Vent.)Wight&Am.exHassk. formabrachycarpa Miq.,Fl.Ned.Ind. 1 (1855)
154.—Type: Blume s.n.[holoL(L908.120-317)], Sumatra.
Millettia argenteaZipp. ex Miq., Fl. Ned. Ind. 1 (1855) 156. —Lectotype (here designated):
Zippeliuss.n., Java[holoL(L908.120-407)].
Millettia sericea(Vent.)Wight&Am.exHassk.var.majorMiq.,Fl.Ned.Ind.,Eerstebijv., Sumatra
(1861)300. —Type: Teijsmanns.n., Sumatra (K).
Millettia sericea (Vent.)Wight & Arn. ex Hassk. var.aurata Miq., Fl.Ned. Ind., Eerstebijv.,
Sumatra (1861) 300. —Type:Teijsmanns.n., Sumatra(n.v.).
Millettia turgidaMiq.,Fl.Ned.Ind.,Eerstebijv., Sumatra(1861)300. —Type:Diepenhorsts.n.,
Sumatra(n.v.).
Millettia sericea (Vent.)Wight& Arn.ex Hassk. var.malaccensis Pram,J.As. Soc. Beng.66, 2
(1897) 88.—Syntypes: Griffith 1764 (K), Maingay518 (A, K,L), Goodenough1706 (n.v.),
Malaysia.
Notes— 1.Stone 15598fromMalaysia hasrather small, thick leaflets,but is not
differentin other characters. Lorzing 6809from Sumatra is in some aspects quite
similar to M. chrysamaryssa; however, the pods ofthis specimen clearly belong to
M. sericea.
2. Several specimens from Java have ± obtuse or rounded leaflets ('var. obtusa
Miq.'). The fruitsofthese specimens appearratherwell developed (size: 50-55 by
18-19by 5-10mm). However, the seeds are flat or indentedandare probably not
viable.No differencescouldbefoundinothercharacters.Thesespecimens are included
inM. sericea.
3.ThetypeofM.sericeaformasubcordataMiq.is givenas collectedby Zippelius
in New Guineaor in theMoluccas. However, thisspecimen was probably collected
by Blumein Java.
4. Aspecimen withaKorthals label(Korthalss.n.,A)gives Borneoas thelocality.
Thespecimen wasprobably collectedinSumatraasweremostofKorthals'specimens
ofM.sericea.
5. 'Var. malaccensisPrain':Of the syntypes Griffith1764(K) and Maingay 518
(A, K,L)were seen. Maingay 518is amixedcollection:duplicates atKew belong to
M. sericea; however,onesheetalsobearstheannotation' 11846,Feb61868,Malacca'.
Anothersheetisalso numberedMaingay 1184,andathird sheetalsoMaingay 1182.
The Leidenand Harvard sheetsbelong to M. chrysamaryssa, ofwhichthe Leiden
sheet alsobears the labelMaingay 508.Nolectotype hasbeen chosen.
4.MillettiasolomonensisVerdc.
Millettia solomonensis Verde.,Kew Bull. 32 (1977)247; Man. New GuineaLeg. (1979)332. —
Type: Schodde & Craven 3805 (holoK; iso A,BRI, CANB, L,LAE), Papua New Guinea,
Bougainville.
F.Adema: Notes on Malesian Fabaceae - Millettia 407
Notes— 1. Verdcourt(1977) refrained fromincluding this species in one ofthe
sectionsofMillettia.Theabsenceofcallosities, thelarge fruitsandseeds indicatethat
M. solomonensisbelongs to sect.Millettia.In inflorescenceandflowercharacters it
is quitesimilarto M. chrysamaryssa andM.sericea.
2. The descriptions ofthefruitsandseedshavebeencopied fromVerdcourt(1977).
3.Thereare severalsmalldifferencesbetweenthespecimens. Muchmorematerial
is neededto seeifthereis morethanonetaxon involved.NGF 31614is describedas
an 8 ft high shrub,it also differs in leaves andfruits. NGF 13393is describedas a
shrubwith deep purple-red flowers (Verdcourt, 1977).
SectionFragiliflorae
SectionFragiliflorae Dunn,J. Linn. Soc. Bot. 41 (1912) 136; Merr., Enum.Philipp. Flow. PI.2
(1923)298; Backer & Bakh.f., Fl. Java 1 (1964)616;Verde.,Man. New Guinea Leg. (1979)
311;Allen & Allen,Leguminosae(1981)543; R.Geesink inPolhill & Raven,Adv.Leg. Syst.
1(1981)258;Leiden Bot.Ser. 8(1984)103.—Lectotype(heredesignated):Millettiamerrillii
Perkins.
PongamiaVent.,Jard.Malm.(1803)t. 28.—Type:PongamiaglabraVent.,nom.illeg.[=Pongamia
pinnata(L.)Pierre =Millettia pinnata(L.)Panigrahi],
Trees.Axillarybudsbutton-shaped tightlyenclosed in thebudscales. Inflorescences
axillary, pseudoracemes with apulvinus atthe baseofthepeduncle. Standardwith
2 largebasal callosities.Anthershairy (at leastsomehairsatthebase).
Distribution— TropicalAsia,throughout Malesia;PacificIslands,Australia,Mas-
carene Islands.
Note— Millettiais conservedoverPongamia (Geesink, 1981b;Brummitt, 1984);
however,Pongamia hasretaineditsstatus as conserved name (Brummitt, 1984).
ThesectionFragiliflorae can be dividedintotwo groups:A:Pinnata-group:Ovules
2 (rarely 1 or3); pods ratherthick, tardily dehiscentto indehiscent;seeds usually 1,
ratherlarge andthick. B: Merrillii/Xylocarpa-group: Ovules 3-10(rarely 2);pods
flattened, dehiscent;seeds 3or more(rarely less), usually small, flattened.
A.PINNATA-GROUP
Thisgroupconsistsof3species: thenewspecies Millettiaborneensis,andM. brachy-
carpaandM.pinnata.
5.MillettiaborneensisAdema,spec. nov. —Fig. 2a,b, 3a
Arbor.Foliaimparipinnata(5-vel)7-foliolata.Foliolaellipticaadobovatabreviteracumi-
natainfrasericea adfereglabra.Pseudo-racemi axillares pedunculobasipulvinato. Ovula
2 (vel 3).Leguminaplerumque 1-seminalis indehiscentia circumsementumescentia.
—Typus: S 27606 (Soepadmo & Smith)(holoL;iso A, K), Borneo,Sarawak,Bukit
Raya, 1.4.1969.
Treesor shrubs, 4-50 m high, dbh7-80cm.Bark grey,innerbark whiteor pale yel-
low, wood whiteto ochre. Twigs terete,3-4 mmdiam., thinly hirsute to glabrous.
Stipules obliquelyelliptic, c.4.5by 2mm,caducous,outsidesericeous,inside sericeous
atbase. Leaves (5-or) 7-foliolate.Petiole3-5 cmlong, striate, thinlyhirsuteto gla-
408 BLUMEA Vol. 45, No. 2, 2000
Fig. 2.Millettia Wight&Arn. sect.Fragiliflorae Dunn.Leaflets from below, a,b. M.borneensis
Adema;c, d. M.brachycarpaMerr.;e.M.pinnata(L.)Panigrahi; f, g.M. velvetina Adema;h,i.
M.xylocarpaMiq. (a,b: SAN25299; c, d:SMHI 1560;e: SAN25767; f, g:NGF 10747; h,i:KEP
98642).
F.Adema: Notes on Malesian Fabaceae - Millettia 409
Fig. 3.MillettiaWight&Arn.sect.FragilifloraeDunn. Pods.a.M.borneensisAdema;b.M.brachy-
carpa Merr.;c. M.pinnata(L.) Panigrahi; d.M.ahernii Merr. & Rolfe;e. M.merrillii Perkins;
f.M. xylocarpaMiq. (a:SAN60019;b: SMHI 363; c:NBFD 9455; d:PNH 11130;e: Ridsdale &
Reynoso 1917; f:FRI 32637).
brous;rachis 6.5-10.5 cmlong, striate, thinly hirsuteto glabrous; pulvinus 5-7 mm
long.Leaflets: terminalelliptic toobovate,7-11.5by 3-5cm,index2.2-2.7, ±papyra-
ceous, oftenrolledupwardswhendry,base cuneate,apex(shortly)acuminate,acumen
4-9mmlong, bothsurfaces(thinly) sericeous,rarely morehirsute, midribandnerves
flator slightly sunken above,nerves 8 or9perside,5-13 mmapart; lateral mostly
astheterminal,3.5-9by 2-3.5cm, index 1.4-3.3, acumen 1-9 mmlong; pulvinus
4-6mm long.Inflorescences 3-16cm long, thinly hirsutetoglabrous; peduncle 3-6
cm long, thinly hirsuteto glabrous; pulvinus 3-5mmlong.Bractstothebrachyblasts
narrowly triangular, c. 3 by 0.7 mm,outside hirsute, inside sericeous. Brachyblasts
(1- or) 2-flowered.Bracts tothe flowers ± ovate,0.6-0.8 by 0.4-0.5 mm, outside
410 BLUMEA Vol. 45, No. 2, 2000
sericeous, insidesericeous or glabrous. Pedicels c. 5 mmlong. Bracteoles attopof
pedicel, elliptictoovate,0.4-0.6 by 0.2-0.4mm, outsidesericeous,insidesericeous
orglabrous. Calyx c.4mmlong,outsidesericeous tohirsute, insideglabrous; lobes5,
inconspicuous. Corolla(light)purplish, standardgreenatbase. Standard:claw 2-3
mmlong;blade±orbicular, 9.5-10by 10mm,retuse,outside sericeous, glabrousto-
wards thebaseand margin,insideglabrous. Wings: clawc. 3mmlong; bladeelliptic
or 'axe-shaped', c.8 by3.5 mm,outsideandinsidesericeous atapex,auriclesinconspic-
uous.Keelpetals: claw3-3.5mm;blade± boat-shaped, 6.5-7by 3mm,outsideseri-
ceous on ventral side, inside sericeousalong ventral margin at least in upperpart,
lateralpocket c. 2 mm long,ratherwide, auricles upright, abovelateralpocket, c. 1
mm high. Stamensc. 10mm long, withbasal fenestrae,free part 1.5-3.5mmlong,
glabrous; anthers0.6-1.0by 0.5-0.6mm, withsome hairsatthebase. Disc annular
oralmostabsent,upto0.1 mmhigh. Ovary3.5-4mmlong, hirsuteorsericeous; stipe
1.5-2mmlong;ovules2(or 3);style 5-6mmlong, sericeousinlowerpart, upwards
glabrous; stigmaterminal, capitate. Pods green,flattenedto inflated,± sharp-edged,
bulging around theseed, oblong to half-moon-shaped, 5.5-8.5by 2-3.5by 0.3-1.2
cm,stipe c. 2 mm long, valves c. 1.7mmthick, outsideglabrous or withsomehairs.
Seedsdiscoidtotransversely ellipsoid,18-20by 15-16by4-8mm,hilum 1—1.5mm
long.
Distribution—Sumatra,PeninsularMalaysia, Singapore, Borneo.
Habitat&Ecology —(Disturbed)primaryforest, swampforest,along rivers. Soil:
sandorlimestone.Altitudeupto 100m.Flowering: MarchtoJune,October, November.
Fruiting: April to November.
Notes — 1. Very flat pods do not containripe seeds, the undeveloped seeds are
filledwith upto5 developing insects.
2. This new species is in several aspects rather similar to M. pinnata. It differs
especially inthe more dense indumentum,thenarrowerand usually thinnerleaflets
thatoftencurl upwards in drying,thesmallerflowers andthesharp-edged pods with
smaller, thinnerseeds. As inM. pinnata thepods are probably tardily dehiscent.In
this characterthey differfrommostother species ofsection Fragiliflorae.
3. Twospecimens fromPeninsularMalaysia arerecordedwith(pale) orangeflowers.
4.Specimens fromPeninsularMalaysia, SingaporeandSumatradifferinindumen-
tumfromtheBorneanspecimens. Inthefirst groupthelowersurfaces oftheleaflets
are(very) thinly sericeousto± hirsutewiththehairstending to becomepatent.The
Bornean specimens are sericeouson thelower surface oftheleaflets withthehairs
appressed. Otherwisethere seemtobe no differences;however, all plants from the
firstgroupseen sofarareinflowerandpods arereally neededforafinaldecision.For
thepresenttheseplants havebeen includedhere.
6.Millettiabrachycarpa Merr.—Fig. 2c, d,3b
Millettia brachycarpaMerr.,Philipp.J.Sci.,Bot. 10(1915)17;Enum. Philipp.Flow. PI.2(1923)
278. —Type:BS 9578 (A,K,L), Philippines,Palawan.
Notes— 1.Thisspecies is aPhilippineendemic.Only foundinLuzonandPalawan.
2.Redexudatehasbeen observedinseveral specimens.
3. InflorescencesofBS29185are sometimesonce branched.
F.Adema: Notes onMalesian Fabaceae - Millettia 411
7.Millettiapinnata (L.) Panigrahi —Fig. 2e,3c
Millettiapinnata(L.) PanigrahiinPanigrahi& Murti,Fl.BilaspurDistr. 1 (1989)210. —Cytisus
pinnatusL., Sp. PI. (1753)741. —Pongamiapinnata (L.)Pierre, Fl.For. Cochin. 5 (1907)
t. 384;Merr., Enum. Philipp. Flow. PI. 2 (1923)298;Craib,Fl. Siam. Enum. 1 (1928)494;
Backer &Bakh.f.,Fl. Java 1 (1964)616;Whitmore,TreeFl.Mai. 1 (1972)303;Verde.,Man.
New Guinea Leg.(1979)312. —Pongamiapinnata(L.)Merr.,Interpr.Rumph.Herb. Amboin.
(1917)271. —Type:Pluk.,Phyt. (1690) 104,f. 3.
[MalapariusRumph., Herb.Ambon. 3(1743) 183,t. 117.]
[MalapariuseNussanive Rumph., Herb. Ambon. 5(1747) 184.]
Robinia mitis L.,Sp. PI.ed. 2(1763)1044,nom.illeg. —Pongamiamitis(L.)Kurz, Prelim.Rep.
For. Veg.Pegu,App.A(1875)49.—Pongamiamitis (L.)Merr.,Philipp.J.Sci.,Bot.5(1910)
101.—Type unknown.
PongamiaglabraVent.,Jard. Malm. (1803)t.28,nom.illeg.; Descaisne,Herb. Timor.(1835)448;
Miq.,Fl.Ned.Ind.l (1855)147;K.Heyne,Nutt.PI.2(1916)306;Gagnep.,Fl.Gen. Indo-Chine
2(1916)441;Ridl.,Fl.Mai. Penins. 1 (1922)593;Burkill,Diet.2(1935)1797.— unknown.
Pongamiaxerocarpa Hassk., Hort.Bogor. descr.,ed. 2(1858) 208. — Pongamiamitis (L.) Merr.
var.xerocarpa (Hassk.) Merr.,Philipp. J.Sci.,Bot. 5 (1910) 101. —Pongamiapinnata(L.)
Merr. var.xerocarpa (Hassk.) Merr.,Enum.Philipp.Flow. PI.2(1923)298. —Typeunknown.
PongamiagrandifoliaZoll. & Moritziin Moritzi,Syst.Verz.(1845)2;Hassk., Hort.Bogor.descr.,
ed. 2(1858)210. —Type:Zollinger1245 (n.v.).
Millettia novoguineensisKaneh. & Hatus.,Bot.Mag.(Tokyo) 56 (1942)567. —Type: Kanehira
& Hatusima 12587(A), DutchNew Guinea,Sennen,40km inwardfrom Nabire,8.3.1940.
Notes— 1.This well-knownwidespread species really belongs toMillettia sect.
Fragiliflorae. SeeGeesink, 1981a,bandAdema,2000.
2. Very variable, especially in NewGuinea.The species varies greatly in shape,
size, andindumentumof leaflets; thereareoften also smalldifferencesin thesizeof
thepetals andtheindumentumofthekeelpetals. OnegroupofNewGuineanspecimens
(Kostermans & Soegeng 975, LAE50088, McDonald<6 Ismail3706, NGF39315)
hasmuchsmallerfruitswithwrinkledwalls.Furtherfieldstudiesmayshowtheexis-
tence ofmoretaxa.
3.1observed dark(red) sap instemsandpod walls.
Uses — Thehard wood is used fortemporary housing constructions. Twigsand
smallbranches are usedas toothbrush.Leaves are usedinlocalmedicine.
B.MERRILLII/XYLOCARPA-GROUP
InMalesiathis groupconsistsof7 species ofwhichM. glabra is describedas new.
8.MillettiaaherniiMerr. & Rolfe—Fig. 3d
Millettia ahernii Merr. & Rolfe,Philipp. J.Sci.,Bot. 3 (1908) 103;Merr.,Philipp. J.Sci.,Bot. 5
(1910) 71;Dunn, J.Linn. Soc. Bot. 41 (1912) 181;Merr.,Enum.Philipp. Flow. PI.2(1923)
278. — Type: BS3373 (Ahern's collector)(K),Philippines,Luzon.
Millettia canariifoliaMerr.,Philipp. J.Sci., Bot. 5(1910)71; Dunn,J.Linn. Soc. Bot. 41 (1912)
178;Merr.,Enum.Philipp.Flow. PI.2 (1923)279.—Syntypes: BS4711 (n.v.),4727(K).
Millettia cavitensis Merr.,Philipp. J.Sci., Bot. 5 (1910)72; Dunn, J.Linn. Soc.Bot. 41 (1912)
179;Merr.,Enum.Philipp. Flow.PI.2(1923)279. —Type:Merrill 4181 (L,P),Philippines,
Luzon.
Notes— 1. Rather similar toM. merrilliibut larger in sizeofleafletsand fruits
withonly aslight overlap. Alsosimilarto M. tenuipes whichdiffers inthe length of
thepedicels andtheslightly largerflowers.
412 BLUMEA Vol. 45, No. 2, 2000
Fig. 4.Millettia glabraAdema.a.Habit;b. flower;c. standardfrom inside;d.wingfromoutside;
e. keelpetalfrom outside;f.staminaltube;g. pistil (allDe Vogel4488).
