Table Of ContentBLUMEA 41 (1996)407-411
Name changes in Agrostis, Arundinella, Deyeuxia,
Helictotrichon, Tripogon(Gramineae)
J.F.Veldkamp
Rijksherbarium/Hortus Botanicus,P.O.Box9514,2300RA Leiden. TheNetherlands
Summary
Buse’s(1854)treatmentofJunghuhn’sGramineaeappearedbefore morepartsofSteudel’s Synopsis
(1853-1855)than waspreviously thought.Agrostis rigidulaSteud. becomes.A. infirmaBuse with
8newvarietal combinations. Arundinella fuscataNeesexBusepreviously regardedasalaterhomo-
nym is invalid, and must becalled A. goeringii Steud. Helictotrichon junghuhnii(Buse) Henrard
supersedesH. virescensNees exSteud.,and TripogonexiguusBuse replacesT. semitruncatus Nees
exSteud.
Introduction
Quite afewspecies ofgrassesoccurring inMalesiawere first describedor namedin
two works that appeared moreorless simultaneously: Steudel,Synopsis plantamm
graminearum (in 6 installmentsbetween 1853and 1855:cf. Stafleu&Cowan, 1985)
andBuse,GramineaeinMiquel'sPlantaejunghuhnianae 3(1854) 341-394.
Steudelcorresponded withmanycontemporarybotanists andalso seems to have
hada manuscript by Nees,whoreceivedandnamedmanyspecies that were sent to
himby hismany correspondents. Steudelalsovisited manyherbaria, andwas even
allowedtostudy theholdings in L(which withBlumethen beingDirectormusthave
been quiteanachievement!), butnot thoseofU,as faras canbetold fromhis refer-
ences and identificationlabels. Buse saw mainly thecollectionsin U, whereatthat
timethe Junghuhn specimens were located(most ofthem are nowin L). Healso
citedNeesmanuscriptnamesobtainedfromsheetsidentifiedby Nees in U.Whether
BuseandSteudelwere awareofeachother's workis notclear; no cross references
seemtohavebeenpublished by either.Perhaps themanuscripts were finishedalmost
simultaneously. On theotherhand,Buse(1857)wrote thatSteudelapparently hadnot
noticedthePlantaejunghuhnianae, thusimplying thathis work musthaveappeared
beforeatleast partofSteudel's.
Except forafewMalesianbotanistsBuse's work was muchneglected, whilethat
ofSteudelhadamuchgreaterimpact. Also, it was thought thatthispartofthePlan-
taejunghuhnianae hadappeared inAugust 1854(Stafleu &Cowan, 1981), whereby
manyofthenames wereantedatedby thoseofSteudel.
Theheading ofBuse's treatment hasthenote "[Febr. 1854]", whichperhaps in-
ducedHenrardto accept thatdateonseveraloccasions (1930, 1940, 1950).Whether
Henrardwascorrect or notinhisassumptions couldnot bereconstructed by me; itis
408 BLUMEA Vol. 41, No. 2, 1996
also notmade clearfromthetwo veryrare re-paged butotherwiseidenticalreprints
ofBuse'spaperinL, whichhave thesamedateintheheading (L Library: Brochure
no.311). Oneis interleavedandBuse's own copy,butwith onlya fewannotations,
and givesnofurtherclue.
Recently, Stafleu& Mennega (1995) haveregarded thisreprint asapreprintdating
fromFebruary, whichgives theworkpriorityover alargerpartofSteudel'sSynop-
sis thanbefore.Thereforeuse'snames shouldbe citedfrom this, with thepage ofthe
laterpublicationminus 340.
Fortunately very few changes are necessitatedby this as faras I can see. These
areenumeratedbelow.
ItmustbenotedthatBuseisoftenmisquoted as 'Biise' because forsomereason
thatis the orthography employed in this paperand in Miquel (1857). Hehimself
wrotehisnamewithoutanUmlaut.
1.Agrostis infirma Buse
Agrostisinfirma Buse inMiq.,PI.Jungh.(Feb.1854)preprint:2; (Aug.1854): 342.—Type: Jung-
huhn s.n.(holoL,no.903.342-37).
Agrostis rigidulaSteud.,Syn. 1 (Apr. 1854) 171.—Type:Zollinger2589 (holoP).
Note—For further informationonthe species and its varieties, see Veldkamp
(1982).
2.Agrostis infirma Buse var. arisan-montana(Ohwi) Veldkamp, comb. nov.
Agrostis arisan-montana Ohwi, ActaPhyt, Botax.Geobot. 2(1933)161.—Type:Ohwi
3463 (holo KYO,iso K).
3.Agrostis infirma Buse var. borneensis (Stapf) Veldkamp, comb. nov.
Agrostis canina L. var.borneensis Stapf, Trans. Linn. Soc., London 4(1894) 246. Type:
Haviland1399(holoK).
4.Agrostis infirma Buse var. diffusissima(Ohwi) Veldkamp, comb. nov.
Agrostis reinwardtii Buse var. diffusissima Ohwi,Bull. TokyoSc. Mus. 18(1947) 8. —Type:
Clemens 33228 =32327(holoBO, isoBM,G,K, L,NSW,NY, US).
5. Agrostis infirma Buse var. formosana(Hack.) Veldkamp, comb. nov.
Agrostis canina L.var.formosana Hack., Bull. Hb.Boiss. 11,4(1904)528. Type: Faurie 724
(holoW,isoKYO).
6. Agrostis infirma Buse var. fukuyamae (Ohwi) Veldkamp, comb. nov.
AgrostisfukuyamaeOhwi inFedde,Repert.36 (1934)39.—Type:Ohwi4147(holo KYO, iso K).
7. Agrostis infirma Buse var. infirma, comb. nov.
(Theautonymautomatically generated by theothervarietalcombinations.)
J.F.Veldkamp: Name changes in Gramineae 409
8.Agrostis infirma Buse var. kinabaluensis(Ohwi) Veldkamp, comb. nov.
Agrostis kinabaluensisOhwi,Bull. TokyoSc.Mus. 18 (1947)8.—Type: Clemens 30273,p.p.
(holoBO, iso B, K,L,M?).
9.Agrostis infirma Buse var. remota(Buse) Veldkamp, comb. nov.
Agrostis strictaBuse (nonGmel., 1791)var. remota Busein Miq.,PI. Jungh.(1854)preprint:2;
(Aug. 1854):342,comb, incorr.—Type: Junghuhns.n.(holoL,no.908.76-441).
Note—Thecorrect epithet touse is 'remota'andnot 'stricta', becauseinthiscase
theautonym was not automatically established. Autonyms are only automatically
generatedwhenthenameofthespecies is legitimate(Art.26.1). Agrostis stricta Buse
beingalaterhomonymofA.stricta Gmel.(1791) isillegitimate.
10.Arundinellafuscata Neesex Buse
ArundinellafuscataNees exBuse in Miq.,PI. Jungh.(Feb. 1854)preprint: 19; (Aug.1854): 359.
Onepair ofcompeting namesofBuse andSteudelis Arundinellafuscata. At first
sightthesituationseems clear:Buse describedaspecies fromJavawithan unnamed
varietyß ‘angustior’ (the wholediagnosis, sonot a name)andSteudel'scombination
foraspecies fromtheNilgiris,India, is alaterhomonym. Theirdescriptions are to-
tally different.Only when Itriedtoappoint a lectotype fortheBuse combinationdid
I noticethatthe specimens cited by Busehadbeen labelledby him withnotes like
"Arundinellafuscata NeesabEs.ß angustior Nees. specim. auth. PI. J. p. 359."A
closerobservation showedthatalso fromthe typography andformatofthepublica-
tionnot aspecies, butanunnamedvariety was described, makingA.fuscata Nees ex
Buseinvalidly published. Hookerf. (1896) attributedthecombinationto Buseand
placedA.fuscata Steud.andA.purpureaHochst. ex Steud. inthesynonymy,but as
A.fuscata Busewas invalidly published, thiswas erroneous. Itwas apparently vali-
datedby Koorders (1911).
Steudel's A.fuscata is not a laterhomonym, butas A. purpureais equally old,
thelatteris thecorrect name, becauseBor(1960: 424) selectedthatone.
The correct namefor'A.fuscata Buse'is A. goeringii Steud. Thetypeofthelat-
ter is Goeringsect. II, no. 139, with 'Japonia?' as thepossible provenance.Goering
collectedboth in Japan and Javaand theoriginofhismaterialis not always clear.
Thenamewasnot takenupby Honda's(1930) enumerationofthegrassesofJapan.
Arundinella goeringii Steud.
Arundinella goeringiiSteud.,Syn. 1 (Mar. 1854)116.—Type:GoeringII, 139(holoP).
ArundinellafuscataNees ex Buse var. (3 Nees ex Buse in Miq.,PI. Jungh.3 (Feb. 1854)preprint:
19; (Aug.1854):359, nom.spec, inval. —ArundinellafuscataNees ex Koord.,Exk.Fl.Java 1
(1911) 119,nonSteud. (1854).—Lectotype: Junghuhns.n.(L,no.908.83-1332).
Distribution —Sumatra, Java,Celebes.
Note—Aformwithtuberculate-hairy glumes hasbeendistinguished as var. cele-
bicaby Jansen(1953) andwas accepted as a distincttaxon by Phipps (1967), butI
doubtits distinctiveness.
410 BLUMEA Vol. 41, No. 2, 1996
ArundinellapurpureaHochst. ex Steud.
Arundinella purpurea Hochst. exSteud.,Syn. 1 (Mar. 1854) 115.— Type:Hohenacker 928 (holo
P, isoK,L, U).
Arundinella fuscataNees ex [Wight,Cat. (1834)97,nomen]Steud.,Syn. 1 (Mar. 1854) 114. —
Type: Wight, Cat. 1667(holoP,iso K, U) (= Bas Hh.prop. 1837,lost).
Arundinellafuscataauct.nonNees exBuse: Hook,f.,Fl. Br.India7 (1896)74.
Distribution — MadrasState, Nilgiris.Hookerf. (1896) citedmaterialfromBur-
ma, theidentity ofwhichis presently unknown.
11.Deyeuxia srilankensis (Davidse) Veldkamp, comb, nov.
Calamagrostis srilankensis Davidse,Fl.Ceylon 8 (1994)107(Amerindedition). —Type:Davidse
7618 (holoUS, isoCAL, CANB, K, KLU,L,MO,PDA, TAES).
Note—Davidse (1994)notedthatthisSriLankaendemicisrelatedtotheSIndian
Calamagrostis zenkeri(Trin.)Davidse.As was pointed out byKorthof& Veldkamp
(1985) thelatteris aspecies ofDeyeuxia Clarion exBeauv.,andthe SriLankaspe-
ciesbelongs to this genusas well.
TheRevisedhandbooktotheFloraofCeylon 8 (1994) appeared intwoidentical
editions(except forquality ofpaper, cover,andlogo), oneby Amerind, theotherby
Balkema. They were printed simultaneously, buttheAmerindone was distributed
first (seealso Veldkamp, 1995).
12. Helictotrichon junghuhnii (Buse) Henrard
Helictotrichon junghuhnii(Buse) Henrard, Blumea 3 (1940) 425. —AvenajunghuhniiBuse in
Miq.,PI. Jungh.(Feb. 1854)preprint: 15;(Aug. 1854):345. —Type: Junghuhns.n.(holoL,
no.903.342-86).
Trisetum virescensNees ex Steud.,Syn. 1 (Apr. 1854)226. —Helictotrichon virescens Henrard,
Blumea 3 (1940)425. —Lectotype:Royle 137 [LIV,K neg. 18361,holo?;K(? nonumbered
collections)].
Note—For an account ofthegenusandarevision ofthe Malesianspecies, see
Sevenster&Veldkamp (1983).
13. Tripogon exiguus Buse
TripogonexiguusBuse in Miq.,PI. Jungh.3 (Feb. 1854)preprint: 13; (Aug. 1854):353. —Type:
Junghuhns.n. (holoL,no.904.84-64).
Tripogonsemitruncatus Nees exSteud.,Syn. 1 (Apr. 1854)301.—Type: 'InmonteLaeou Javae'
(holo?P, butnotfound).
Distribution—Java,Lesser SundaIslands(Bali, Lombok,Alor).
Note—Bor(1960: 521) hasequated theJavanese T.semitruncatuswith thecon-
tinental T.filiformisNeesex Steud., but Ihavethe impression thatinthis concept
a numberofclosely related taxa arepresent. IfBor turns outto have beencorrect,
T. semitruncatusis theoldestnameforthewholeas well.Theresultis thenaninter-
esting disjunct distribution:Kashmirto SWChina, 'jumping' to Java, Bali, Lom-
bok, andAlor.
J.F.Veldkamp: Name changes in Gramineae 411
Ohwi(1947) distinguished T.javanicus Ohwi formaterialtwice collectedon the
Arjuno by Van Steenis (7029, p.p., BO, L; 11881, BO). Thisdiffers by having a
laxinflorescenceandfirstlemmasapically hardlybilobedwithasingle awn. Because
vanSteenis7029was mixedwiththetypical form,I doubtthat thisis a distincttaxon.
ACKNOWLEDGEMENTS
I wantto thankDr.G.Zijlstra (U) for the fruitful discussions wehad onthe Buse publication, and
forcheckingtheU holdings.Dr. H.Scholz(B)kindly wentovertheBerlin material.
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