Table Of ContentBASTERIA,71: 77-80,2007
CorrectionsinCochlodina(Gastropoda, Pulmonata, Clausiliidae)
E. Gittenberger
National Museum ofNatural History Naturalis,P.O.Box 9517,NL2300RALeiden,TheNetherlands;
[email protected]
Asubspecies ofCochlodina laminatais described a second time.Itcanbe distinguishedonthe
basis ofmainlytheprominenceoftheaperturallamellae and shell size. Someincorrect data
regardingItalian,alpineC. laminata arecorrected.
Key words: Gastropoda,Pulmonata,Clausiliidae,Alopiinae,Cochlodina,Italy, Slovenia,Croatia.
Thewell-knownnameof a very characteristicsubspecies of Cochlodina(C.) laminata
(Montagu, 1803)should not be changed for nomenclatorialreasons. Records of C. (C.)
laminatafromthe Dolomites, N. Italy, couldnotbe verified.Inallcases where theshells
were availableforresearch, thesebelonged toC. (C.)fimbriata(Rossmassler, 1835).
ClausiliidaeJ.E. Gray, 1855
Alopiinae A.J.Wagner, 1913
CochlodininiLindholm,1925
CochlodinaA.Férussac, 1821
Cochlodina(C.)laminatainaequalis (A.Schmidt, 1868)(figs 1,2)
Clausilia melanostoma var.inaequalisSchmidt,1868: 29.
Clausilia inaequalisSchmidt,1868: 31,32.
Not: Clausilia bidensvar.inaequalisM.Gallenstein,1852.
Clausiliastra grossavar.inaequalis;Clessin,1888: 339.
Cochlodina laminatainaequalis;Gittenberger, 1967: 31, fig. 8.Nordsieck, 1969:114.
Cochlodina laminata insulana;Nordsieck, 1993: 34[part.].Not Gittenberger, 1967.
Material. — Croatia,Istra:UckaMtn,800-950malt.,UTMVL31,E.G.leg.(RMNH 106889 [fig.1]&
106890/13[fig. 2]); 'Monte Maggiore' (= Ucka Mtn), UTM VL31, W. Klemm leg. (ex colln W. H.
Neuteboom)(RMNH 106892/3);ditto,W.Klemmdon. (excolln F.E.Loosjes) (RMNH 106891/4);N-slope
nearVeprinac, 350 malt.,UTMVL42,W.J.M.Maassen leg. (YU345/1);N-slopenearPoklon, 970malt.,
UTMVL31 (YU346/3); nearPoklon, ruins near motel,950 m alt.,UTM VL31,W. J. M. Maassen leg.
(YU398/12).
Description. — Shelllarge, with a very prominent, apertural, palatal callus, thatis
interrupted conspicuously by thelowerpalatal plica. Plicaeprincipalis andbasalispromi-
nentand connectedtotheradial callusinfront;plica basalisoftenbroadenedin frontand
less lamella-like.Usually withtwo small, somewhatirregular, middlepalatal plicae, that
are ofteninterconnected, one of whichbroadenedin frontand connectedto thecallus.
Lamellasubcolumellaris protruding more conspicuously than in any other C. laminata
subspecies.
78 BASTERIA, Vol. 71, No. 1-3, 2007
Figs1-4.Cochlodina (C.)laminata subspec.1,2,C.(C.)l.inaequalis(Schmidt,1868),Croatia,Istra,Ucka Mtn,
800-950malt.,UTMVL31;E.&A.C.Gittenbergerleg.(1,20.6x4.9mm;2,18.6*4.5mm).3,4,C.(C.)l.insu-
lana Gittenberger, 1967, Croatia, Krk island, near Risika (= W. of Vrbnik), UTM VK79; W.J.M.
Maassen leg. (3,18.2x4.4mm;4,17.2 x4.3mm).Photographs:J.Goud,Leiden.
Measurements(n=37): 17.6-19.9*4.5-5.0mm. Gittenberger (1967:32),onthebasisof
alargermaterial,reported alarger size-range, viz. 18.0-21.0*4.3-5.0 mm.
Differentiation.— CochlodinalaminatainsulanaGittenberger, 1967,fromtheislandof
Krk (figs3, 4)is clearly smaller.Aseries [n=49]collectedvii.1982 byMr W.J.M. Maassen
in Slovenia, Krk, near Risika, W.of Vrbnik, UTMVK79measures 14.8-18.1x 3.8-4.3mm.
Asecondsample [n =9] measures 15.0-18.0* 4.1- 4.3mm.Thepalatalcallus andthepli-
caeand lamellaeareless prominentthanin C. l.inaequalis; the lamellasubcolumellarisin
particular is less prominently protruding. The other subspecies of C. laminata
Gittenberger: Corrections in Cochlodina 79
Figs5-7.Cochlodina fimbriata(Rossmässler, 1835).5,Austria,Kärnten,KoschutaMts, 11km SE.ofFerlach,
1650m alt.,E.Gittenbergerleg.(15.5x3.8mm);6,7,Italy, Belluno,Dolomiti,nearRifugioChiggiato,6km
NNW. of Pieve di Cadore, 1900 m alt., G. Marcuzzi leg. (ex colln Venmans) (6, 13.5x3.7
mm;7, 13.3x3.4mm).Photographs:J.Goud,Leiden.
are evenmoreclearly different,
Notes. — Nordsieck (1993: 34) showed that Clausilia bidens var. inaequalis M.
Gallenstein, 1852, and Clausiliamelanostomavar. inaequalis Schmidt, 1868, are primary
homonyms. Sincetheformernameis not"alsoin use",whereasClessin(1888:339)before
1899already classified thelattertaxon withClausiliastraL.Pfeiffer, 1856,andno cases of
secondary homonymy areknown, thejuniorprimary homonym "mustnotautomatical-
ly"bereplaced (ICZNArt.23.9.5). Thecase willbereferredtothecommission.Nordsieck
80 BASTERIA,Vol. 71, No. 1-3,2007
(1993:34)suggested tousethe nameofthe insular subspecies, i.e.Cochlodinalaminatainsu-
lana Gittenberger, 1967, "vorlaufig" [= for the time being] also for the "kaum ver-
schiedene"[=hardly different] mainlandpopulations. Iseenoreason tochange mypre-
vious view (Gittenberger, 1967)in thiscaseasfar asthe interpretation oftheconchologi-
cal differencesis concerned.Therefore,the twotaxa are consideredsubspecies and char-
acterized as suchoncemore.
Cochlodina(C.)fimbriata (Rossmässler, 1835)(figs5-7)
Notes. —Despite thefact thatCochlodina(C.)fimbiata (figs 5-7) is quitevariable,with
disjunctions in theAlps and theBalkans, thespecies has largely remainedspared from
subspecific classifications. IntheDolomitesincentralnorthernItaly itoccurs withaform
thatslightlyremindsof C.(C.)laminatabecausethecharacteristic,white, apertural,palatal
callusislessconspicuous thanusual.Theshellsareless paleincolourandrelativelysmall.
They can stillberecognized as belonging to C. (C.) fimbriatabecauseof the shortlower
palatal fold.This form (figs 6, 7)has been referred tomore than once inthe literature
(Thorson, 1930; Venmans,1954, 1959),butalways incorrectly asC1 . (C.) laminata.Maybe it
deservesasubspecific status.It mightbecomparable in itsphylogeographic historytofor
example Clausilia(Andraea) dubiaalpicola Clessin, 1878(see Bank&Gittenberger, 2000).
REFERENCES
BANK,R.E.,&E.GITTENBERGER,2000.On thepolytypic andproblematicClausilia dubia:notesonits
nomenclature and systematics(Gastropoda,Pulmonata,Clausiliidae).— Basteria 64: 15-27.
CLESSIN,S., 1888.DieMollusken-Fauna Oesterreich-Ungarnsund der Schweiz 3: 321-480. Niirnberg.
GITTENBERGER,E.,1967.Cochlodina laminata (Montagu)inihremsiidostlichen Verbreitungsgebiet.—
Archiv fiirMolluskenkunde 96:25-37.
NORDSIECK, H., 1969.ZurAnatomieundSystematik der Clausilien,V.Genitalsystem undSystematik
des GenusCochlodina. —Archiv fiirMolluskenkunde 99: 107-132.
NORDSIECK, H., 1993. Beitrage zur Nomenklatur der europaischen Binnenmollusken, I. Kritische
Anmerkungenund BerichtigungenzurNomenklatur vonArttaxa derClausiliidae. — Heldia2:33-
42.
SCHMIDT,A., 1868.System dereuropaischenClausilien undihrer nachstenVerwandten: 1-176.Cassel.
THORSON, G., 1930. Zoogeographischeund okologische Studien iiber die Landschnecken in den
Dolomiten. — ZoologischeJahrbiicher.Abteilung fiir Systematik, Okologieund Geographieder
Tiere60: 85-238.
VENMANS,L.A.W.C., 1954.Land and freshwater mollusks from the Dolomites. — Atti dell'Istituto
Veneto diScienze,LettereedArti.Classediscienze matem.enaturali 112: 59-96.
VENMANS,L.A.W.C., 1959. Land and freshwater molluscs from the Dolomites (supplement). — Atti
dell'Istituto VenetodiScienze,Lettere edArti.Classe discienze matem.enaturali 117: 357-371.
