Table Of ContentPROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON
lIl(4):795-798. 1998.
Acteonina permiana, a new species from the Permian Coyote Butte
Formation, central Oregon (Mollusca: Gastropoda: Actaeonidae)
Rex Alan Hanger and Ellen E. Strong
(RAH) Department of Geology, George Washington University, Washington, D.C. 20052, U.SA.;
(EES) Department of Biological Sciences, George Washington University,
Washington, D.C. 20052, U.S.A.
—
Abstract. Acteonina permiana, is described from the Early Permian Coy-
ote Butte Formation of Crook County, central Oregon. Occurrence of Ac-
teonina indicates affinity with the Carboniferous to Permian Boreal faunal
province, and implies that the Grindstone terrane need not have an "exotic"
origin thousands ofkilometers from the North American continent during the
Permian.
Acteonina is among the rarest of all Up- in a forearc basin (Blome & Nestell 1991).
per Paleozoic gastropod genera. Knight The exact dimensions of this basin, and
(1932, 1936, 1941) counted only two specifically the longitudinal separation of
known specimens, both from the Visean of this island arc from the North American
Belgium. In their survey ofPaleozoic opis- mainland remain controversial, and dis-
thobranch gastropods, KoUmann & Yoch- tances ofgreater than 5000 km with south-
elson (1976) point out that the genus, Ac- ern hemisphere origins have been hypoth-
teonina, "... has not yet been found in esized (Jones 1990, Miller et al. 1992).
North America in spite of careful search The Coyote Butte Formation fauna has
among large collections of Pennsylvanian been only partially described: brachiopods
&
and Permian gastropods." Acid etching of (Cooper 1957); fusulinids (Skinner
limestone samples from the Early Permian Wilde 1966), colonial rugose corals (Ste-
&
(Wolfcampian-Leonardina) Coyote Butte vens Rycerski 1983) and conodonts
Formation of central Oregon has produced (Wardlaw et al. 1982). Acid etching reveals
many new taxa (Hengstenberg et al. 1997), the presence of many undescribed gastro-
including four specimens of the new spe- pods, bivalves, chitons, rostroconchs, nau-
cies, Acteonina permiana. tiloids, bryozoans, sponges, trilobites, os-
The presence of Acteonina in Oregon tracods and annelids. A detailed paleobi-
has important implications for western ogeography, incorporating the paleoecolog-
North American terrane paleogeography. ical framework, is desirable, butmuchbasic
The Coyote Butte Formation is part of the taxonomic work needs to be completed on
Upper Paleozoic-Mesozoic Grindstone ter- these important taxa.
rane of central Oregon (Wardlaw et al. Ofthe describedfauna, the fusulinids and
1982), and occurs as chaotically inter- corals have thus far been most useful for
mixed limestone blocks within cherts and paleobiogeographic comparison. The colo-
volcaniclastics. The limestone is interpret- nial rugose corals are part of a ""Thysano-
ed as slide and slump blocks that became phyllum Belt" fauna that existed along the
detached from a carbonate shelf and incor- northern and western margins of Pangea,
porated into deeper-water basinal elastics but Stevens and Rycerski (1983) suggested
796 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON
Figs. 1—3. Acteoninapermiana, new species, from the Coyote Butte Limestone, Crook County, Oregon. 1,
Holotype USNM 498736. Bar is 1 mm; 2, Holotype USNM 498736, close-up of protoconch, showing heter-
ostrophic larval whorl. Bar is 100 ixm; 3, Paratype USNM 498737. Bar is 1 mm.
that the Coyote Butte fauna might have ex- Systematic Paleontology
isted several hundred kilometers west ofthe
margin based upon high Otsuka coefficient Family Actaeonidae d'Orbigny, 1842
similarity with known exotic terranes. Fu- Genus Acteonina d'Orbigny, 1850
type species Acteonina carbonaria
sulinids suggest a position at equatorial lat-
itudes and many hundreds of kilometers (DeKoninck, 1843)
Acteonina permiana, new species
west of the Permian North American mar-
Figs. 1-3
gin (Ross and Ross 1983).
—
Presence of the only other species, A. Diagnosis. Very small (less than 4 mm
carbonaria, in the plate-bound basins of length), shell outline like modem genus,
Belgium suggests a northern hemisphere Conus, with turret-stepped spire, elongate
position for the Coyote Butte Formation aperture, heterostrophic larval shell andcol-
and general affinity with the Carbonifer- umella with pli—ca.
ous-Permian Boreal fauna. Biogeographic Description. Small for genus; subcy-
evidence from so small a sample size is ten- lindrical outline, superficially resembling
uous, but corroborates preliminary paleo- Conus; flat whorl profile and rectangular
magnetic evidence (Harbert et al. 1995) of sutures, producing a turreted spire; broad
a northern hemisphere position for the sutural ramp; narrow aperture, shallowly
Oregon Blue Mountains region. Long dis- expanded at base; concave, plicate colu-
tance transport of the Grindstone terrane is mella; heterost—rophic larval whorl (Fig. 2).
not required for this distribution. Depository. Four silicified specimens
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VOLUME 111, NUMBER 4 797
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are housed in the collections ofthe Depart- Oregon. Smithsonian Miscellaneous Collec-
ment of Paleobiology, National Museum of tions 134(12):1-79.
Natural History (USNM), Smithsonian In- Harbert, W., J. Hillhouse, & T. Vallier. 1995. Paleo-
stitution, Washington, D.C. Type specimens magnetism of the Permian Wallowa Terrane;
include: USNM 498736 (holotype), USNM niym.p—licJaotuironnalfoorf GteerorpahnyesimciaglraRteisonearacnhd 1o0r0o(g7e):-
498737 (paratype). The two other speci- 12573-12588.
mens (USNM 498738, USNM 498739) are Hengstenberg, C. A., R. A. Hanger, E. E. Strong, & J.
fragments of turreted spires that also have M. Yarnell. 1997. Paleogeographic significance
the sutural ramp a—ndUSfiNveMwhorls. ioafnaCrothyrootpeodBsutatnedFmoorlmlautsicosn,ofcetnhteralEaOrrleygoPne.rm-
Measurements. — 498736 (holo- Geological Society ofAmerica, Abstracts With
type, Figs. 1, 2): length 4.10—mm, width Programs 29(3):23.
3.61 mm, whorl number 6. USNM Jones, D. L. 1990. Synopsis of Late Palaeozoic and
498737 (p—aratype. Fig. 3): length——2.15 Mesozoic terrane accretion wit—hin the Cordil-
nam, width 1—.66 mm, whorl number 5.5. lera of western North America. Philosophical
Transactions of the Royal Society of London,
Discussion. The other species of the series A 331:479-486.
genus, A. carbonaria, is about two times Knight, J. B. 1932. The gastropods of the St. Louis,
the length ofA. permiana. Whorl counts are Missouri, Penn—sylvanian outlier: IV. The Pseu-
the same for both species so we are not domelaniidae. JournalofPaleontology6:189-
comparing juveniles of the Permian with 202. —
adults of the Carboniferous. The acid . 1936. Notes on Paleozoic Gastropoda. Jour-
nal ofPaleontology 10:520-534.
etched residues from the Coyote Butte Fm. . 1941.PaleozoicGastropodGenotypes.—Geo-
contain species from many different phyla logical Society ofAmerica, Special Papers 32:
that have variable lengths from several mm 1-510.
to several cm, so we are not biased by a Kollmann, H. A., & E. L. Yochelson. 1976. Survey of
size-sorted sample. Acteonina carbonaria Paleozoic gastrop—ods belonging to the subclass
Opisthobranchia. Annals Naturhistorische
also has a prominent collar-like ridge at the Museum Wien 80:207-220.
base ofthe parietal wall, which is absent or DeKoninck, L. G. 1843. Descriptiondes animauxfos-
not preserv—ed in A. permiana. siles, qui se trouventdans le terrain carbonifere
Location. The fossil locality is in Crook de Belgique, Liege, 1-651.
County, Oregon, which may be found on . 1881. Faunaducalca—irecarboniferedelaBel-
the Twelvenule Reservoir, Oregon 7.5 min- gique, Gasteropodes. Annales Musee royal
d'Histoire naturel, 6:1-170.
ute U.S. Geological Survey quadrangle, Meek, F B. 1863. RemarksonthefamilyActaeonidae,
1981 edition. The UTM coordinate is with de—scriptions ofsome new genera and sub-
11TKU75826907. The geographic coordi- genera. American Journal of Science 35:89-
nates a—re 119°47'17"N, 97°44'10"W. 94.
Age. Early Permian (Wolfcampian to Miller,&E.RL..,MMa.drMi.d.Mil1l9e9r2,.C.LaHt.eSPtaelveeonzso,iJc.Ep.alWeroiggehot-,
Leonardian) ages for the Coyote Butte For- graphic and tectonic evolution of the western
mation are suggested by: brachiopods U.S. Cordillera. Pp. 57-106 in B. C. Burchfiel,
(Cooper 1957), conodonts (Wardlaw et al. P. W. Lipman, andM. L. Zoback, eds..TheCor-
1982), colonial rugose corals (Stevens & dilleran Orogen: Conterminous U.S. The geol-
Rycerski 1983) and fusulinids (Skinner & ogy ofNorth America, volume G-3. Geological
Wilde 1966). Society ofAmerica, Boulder, Colorado, 724pp.
D'Orbigny, A. 1842. Paleontologie Francaise,Terrains
cretaces 2, Gasteropodes. G. Masson, Paris, 1-
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Cooper, G. A. 1957. Permianbrachiopods fromcentral accrted terranes ofwestern North America. Pp.
798 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON
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